AMAZONIANA XIII (3/4): 325-416 Kiel, Dezember 1995 Description of nine new centipede species from Amazonia and related matters on Neotropical geophilomorphs (Chilopoda: Geophilomorpha) by Luis Alberto Pereira, Alessandro Minelli & Francesco Barbieri Dr. Luis Alberto Pereira, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque s.n., 1900 La Plata, R. Argentina. Prof. Dr. Alessandro Minelli and Dr. Francesco Barbieri, Dipartimento di Biologia, Università degli Studi di Padova, Via Trieste 15,135121Padova, Italy. (Accepted for publication: September, 1995). Abstract Nine new species of geophilomorph centipedes, all from Amazonian sites, are described: one species belongs to Ballophilidae (ltyphilus demoraisi n.sp.), three to Geophilidae (Ribautia (Schizoribautia) dfficilis n.sp.,R. (R.)ducalis n.sp.,R. (R.)proxina n.sp.),andfivetoSchendylidae(Pectiniunguistlucalis n.sp.). The d allotype of Ityphilus craåil/i PEREIRA, MINELLI & BARBIERI, 1994 is designated and described. Descriptive data and/or taxonomic comments are given for several Neotropical species of the genus Ribautia'. R. (SchiToribautía) nrontana KRAUS, I 954 is regarded as a good species and redescribed fromtypematerial. R.(5.)seydi RIBAUT, 1923,R.(5.)peruana VERHOEFF, 1941 and R.(5.)titicacde TURK, 1955 are regarded here as distinct species. Ribau¡ia si/v¿n¿ KRAUS, 1954 is placed in the subgenus Ribautia (Schiaoribautia) and the d holotype is described. Keywords: Amazon, Neotropics, Chilopoda, Geophilomorpha. Resumo Nove espécies de cenkrpéias geofilomorfas, todas da região da Arnazônia, são descritas: unra espécie representa os Ballophilidae (ltyphilus denoraisi n.sp.), trôs os Geophilidae (Ribautia (Schiaoribautia) dfficilis n.sp., R. (R.) ducalis n.sp., R. (R.) proxina n.sp., cinco os Schendylidae (Peuiniungus ducalis n.sp., Schendylurus continuus n,sp., S. janauarius n.sp., S. marchantariae n.sp., S. oligopns n.sp.). O d alotipo de ltyphilus crabilli PEREIRA, MINELLI & BARBIERI, 1994 é designado e descrikr. Dados descritívos e/ou comentários taxonômicos são dados para várias espécies Neotropicais do gênero Ribautia: R. (SchiToribautia) montana KRAUS, 1954, é considerada uma espécie válida e redescrita baseada enr material tipo; R. (S.) seydi RIBAUT, 1923, R. (5.) peruana VERHOEFF, l94l e R. (5.) titicacae TURK, I 955 são consideradas espécies dis1. rtas. Ribautia silv¿na KRA US, I 95 4 é colocada no subgênero Rlåattla (Schizoribautia) e redescrita baseada no cl holr 77 in the nrales; 75,77 or 79 in the females. In Forcipulae: when closed, the telopodites are at the sane level of the anlerior margin of head or extend sonre specintens thc midpiecc oflabrurl'l has a robust, dark and sharply pointed tooth and hyaline filanrents slightly beyond it. Basal plate with an irregular tfansverse rt¡w of 6 large setae near the posterior margin at both sides (Fig. 97), but in other specinrens it lacks the central tooth (Fig. 122). and a few additional snraller setae dispersed on the surface of the posterior half. Telopodites: Other characters with no significant variation. trochanteropraefemur with two denticles, the distal one deeply pigmented and subtriangular, proximal In àll specimens studied the lateral pieces of labrum present hyaline filaments only on the 20olo or less denticle shorter than the distal and unpigmented (Fig. 137). Fenrur and tibia without denticles. -) -) .1, -t -)-) I Tarsungulunr basally with a well developed and deeply pignrented denticlc; cl. an<ì v. edge of the ungular present only on the anterior region 0f the body, but on Ribautia ntonÍana and Ribautia peruana the porc blade not serrulate. Calyx of poison gland subtliangular (Figs. I 38- l -19). Chaetotaxy of coxosternunr an¿ fields are present along all the body, in consequence the two last cannot be conspecific with the first. lelrpodites as in Fig. ì.1ó. Second, as explained in the following (see also KRAUS, 1954) R. montano has two significative diffèrenc- Legs (last pair excepted) with chaetotaxy (Fig. 140) unifurm throughout the body length. Claws es with R. peruana and for that reason these two species should be difft:rent and valicl 9nes. ventrobasally with one anterior and one posterior spine (Fig. I 4l ). Diagnosis. - A Ribautia (Schiv.tribautia) species with coxal organs grou¡lecl in 3+3 clusters an{ ventral Stema: pore fields present from the second to the penultinrate sternum. Fields undivided on sterna pores present on both anterior and posterior sterna. Anìong the Neotropical species of the genus which II-XIV and XLVIII, but divided in two subsynìmetrical areas on sterna XV-XLVII. Fornr 6f fiel¿s share these traits it seems more closely related to R. (5.) dilJitilis n.sp. arrd R. (5.) peruana VERHOEI.-F. changing along the trunk as in Figs. I 42- I 48. Nunrber of pores on selected stema: on stemum II, 44 pores; ft. (5.) montana can be differentiated fronr these species by nreans of the characters in table 3. on V,64; <>nlX,7'7; on XIV,68; on XV,5l+49; on XLVll,2O+26; on XLVIII, 32. Type material examined. -'Pararypoid' I with 55 (57 ?) p.Ì., b.l, 24 nrm. peru: Hacienda Tau- Last leg-bearing segment without pleurites at the sides of praetergunr. Praesternunr not divided along lis (6"50'S; 79'10'W), Quebrada Pajonal, (mountain forest at ni 2700 a.s.l.) KOEPCKE \eg.27.12.1952 the sagittal plane; form and chaetotaxy of sternunr and tergum as in Figs. I 49- 1 50. Coxopleura protrucling (SMF 2146ll). This specimen is preserved partly in one slide (the twc last leg-bearing segments and at their distal v. ends, setae snrall and numerous on distal internaì eclge. the renraining surface with few temrinal segments) partly in alcohol (forcipular segment followed by 53 leg-bearing segments, hea{ and larger setae. Coxal organs arranged in 3+3 clusters, the anterior pore opens on the membrane between nrouth parts). - KRAUS says in the original description: "Beinpaare: 4i (Typus), 53 ocler 57 coxopleuron and praesternum, covered by the latter; middle and posterior pores open on the mentbrane (Paratypoide)". Because the paratypoid studied by us has 55 pairs of legs, we do not know if this is a between coxopleuron and sternunr, covered by the latter (cf. fentale. Fig. 151). Last legs with seven typographic mistake or 2 leg-bearing segments are ntissing in the specinren. podomeres, form and chaetotaxy as in Figs. 149-150. Praetarsus unguifornr, relatively snlaller than those Type locality. - Peru: At Km 35 of the Route Olnlos-Jaen 6'10'S,79'10'W, ntountain woodland at nr of the preceding legs. 1400 a.s.l., under stones and logs. Terminal segments: intermediate tergunr with posterior nrargin convex, intermediate sternum and first Known range. - Peru: Knl 35 along the way fronl Olmos to Jaen, at 6' l0'S,79. l0'W. genital sternum with posterior margin concave. Gonopods apparently r¡niarticulate (suture between Depository of type. - Senckenberg Museunr Frankfurt. presumptive basal and apical articles not evident) with ca. I I setae (Fig. 149); penis with 3+3 apical setae Additional information based on I 'Paratypoid'SMF 214611. Wirh 55 (57 ?) pairs of legs, body dorsally. Anal organs present. lenglh 24 mm, maximum body width 0.8 mnr. Female allotype. - 49 pairs of legs, body length 17 nrnt, nraximum body width 0.6 nrm. Colour (of preserved specimen in alcohol) yellowish, forcipular segment pale ochre. All features siniilar to those in the nrale except for the shape and pilosity of the last leg-bearing Antennae ca. 2.5 tinres longer than the cephalic plate, distally attenuate, all articles longer than wi¿e. segment and terminal segntents. Cephalic plate nearly rectangular but sides curved, distinctly longer than wide (ratio 1.54: I ). Lastleg-bearingsegment:formandchaetotaxyof sternumandtergunrasinFigs. l5l-l52.Coxopleura Clypeus with 3 antero-central setae placed on a subcircular clypeal area and 3 bigger setae on the sligthly protruding at their distal v. ends, setae snrall and nunìerous on the distal intemal edge, the antero-central part of the antertor half. Surface of clypeal area very densely reticulated (Fig. I 54). rernaining surface with few bigger setae. Podomeres of terminal legs with shape and chaetotaxy as in Figs. Labrum: midpiece large, separating sidepieces and slightly overlapped by thenr, with ca. 9 hyaline 151-152. teeth, the more central ones shorter than the lateral ones. Sidepieces with ca. l3+13 hyaline filantents. Terminal segments: shape and chaetotaxy as in Figs. l5l-152. First maxillae without palps on coxosternum; telopoclites with a very snrall palp (Fig. 156). Variation. - In our small series, the nunrber of pairs of legs varies as follows: in the ntales 4'7 and,49., Coxosternum without setae; median projections of coxosternum subtriangular, well developed ancl providecl the females only show 49. with l1+Tsetae.Articlelloftelopoditewith5+4v.setaean<Ì3+3d.sensilla(Figs.155-156). Second maxillae: the two coxites with 7+8 setae near the internal ntargins and 4+6 sensilla ¡rear the lateral margins are joined centrally only by a non-areolate mentbranous isthmus (Fig. I55). Apical claw of Ribautia (Schixorihautia) limaensis KRAUS, 1957 (Fig. 153) telopodite well developed ancl without teeth. Forcipulae: basal plate with an irregular transverse row of7 large setae near the posterior nrargin antl Ribautia (Schi¿oribautia) linraensis KRAUS, 1957 - Senck. biol. 38(5/6): 376 a few additional smaller setae dispersed on the surface of the posterior half. Telopoc.lites: trochanteropraefemur with two denticles, the distal one deeply pignrented, the proxinral denticle nluch Diagnosis. - Among the known Neotropical specics of the subgenus R ibautia ( St'hiz.oribaut ia) only th is shorterthanthedistalandunpigmented(relativesizeofbothteethasinFig.158).Tarsungulumbasally species and R. (5,) silvana present only one clus¡er of coxal organs on each coxopleuron. They can be with a well developed and deeply pigmented denticle; ungular blade with dorsal ancl ventral eciges n¡t differentiated as in Table 4. serrulate (Fig. 158). Calyx of poison gland subtriangular (Fig. 159). Type locality. - Peru: Lomas de Atocongo, close to Linla. This is also the only locality known to datc. Stema: the undivided subovoidal pore fields are slightly longer than wide (Fig. 160), sternunl VII with 161 pores. Last leg-bearing segment without pleurites at the sides ofpraetergunt. Praesternunl apparently divided Ribaulia (Schizorihautia) montana KRAUS, 1954 (Figs. 154-160) along the sagittal plane. Remarks. - This description is inconrplete, because the antennal setae of this specimen are lost, the Ribautia (Scltiaoribautia) monton(t KRAUS, 1954 - Senckenbergiana 34(4-6): 315 small specialized setae are also nrissing and the terminal segnìents of thc bocly are badly collapsetl in the slide. Remarks. - This species described by KRAUS in 1 954 was subsequently regarded by the sanrc author (KRAUS 1957:318) as a synonym of Ribautia sayrli RIBAUT, 1923. KRAUS (1957: 378) als¡ regar6ccl Ribautia peruana VERHOEFF, 1941 as another synonynì t¡f Ril¡au¡ia (Schitoribnuria) seyrlr. However we believe that there are reasons for revalidating these species. First, in Ribautia seydi the pore fields are 334 335 I l Ribautia (Schizoribautia) peruana VERHOEFF, l94f (Fig. l6l) two leg-bearing segments with terminal segments; (7) last left le¡¡. - Depository of type. - Senckenberg Museum Frankfurt (SMF 2152). Schizoribautia peruana VERHOEFF, 1 94 1 - Beiträge zur Fauna Perus I (2): 7 I . Type locality. - Peru: Hacienda Monteseco,6" 50' S; 79" l0' W. This is also the only locality known Ribautia peruana, ATTEMS, 1947 - Ann. Naturhistor. Mus. Wien 55: 143. to date. Schizoribautia peruana, CHAMBERLIN, 1957 - Proc. Biol. Soc. Wash. 70: 27. St'hizoribautia peruana peruÕna, TURK, 1955 - Proc. Zool. Soc. Lond. 125(3-4): 487. Redescription Diagnosis. - This species seenls to be closely related to R. (5.) montana; R. (S.) ti¡icacae and R. (S.) Male holotype. - 49 pairs of legs, body length l4 mm, maxinrurn body width 0.7 ninr. Colour of diJJicilis n. sp. For differential characters see table 3. preserved specimen on slides apparently pale yellowish with forcipular segment a little darker. Type locality. - Peru: Sivia. Antennae ca. 3.2 times longer than the cephalic plate, distally slight.y attenuate, all articles longer than Known range. - Peru: Sivia (VERHOEFF, l94l); Quebrada de Cajuraga (TURK, 1955). wide. Setae on a.a. I to VII of different length and few in number, those,of remaining articles progressively Remarks. - The type material of this species is unfortunately lost. However, according to literature shorter and more numerous towards the tip of the appendage (Figs. lf,2-163). Terminal a.a. with ca. ll descriptions this species must be considered as a valid one, not as a synonym of R. (S.) seydi as suggested claviform sensory setae on the external border and ca. l2 on the internal borcler (Fig. 165). Distal end of by KRAUS 11957:358t. this a.a. with ca.5 very small specialized setae apparently not split apically (Fig. 165). Dorsal and v. surface of a.a. II, V, IX and XIII with very snrall specialized setae, which on the v. sicle are restricted kr Ribautia (Schizoribautia) seydi RIBAUT, 1923 an internal latero-apical area and are represented by two different types: d and å. Type d setae are very thin and not split apically. Type å setae are thicker and very similar kr those on the distaì end of the Ribautia seydl RIBAUT, 1923 - N. Caledonia, Zool.3 (Lief. l):71. terminal a.a. (a, I), Fig. ló4). A.a. II with i type á sera; a.a. v (Fig. 164), IX and XIII with I type a and Ribautia seydi, ATTEMS, 1928 - Ann. S. Afr. Mus. 26: 173. I type å seta. Ribautia seydi, ATTEMS, 1929 - Das Tierreich 52: 295. Cephalicplatenearlyrectangular,butsidescurveddistinctlylongerthanwide(ratio 1.45:l),shapeand Ribautia zeydi (sic.), VERHOEFF, l94l - Beiträge zur Fauna Perus 1(2): 72. chaetotaxy as in Fig. 166. Schizoribautia seydi, KRAUS, 1954 - Senck. 34(416):315. Clypeus with 4 antero-cenlral setae placed on a subcircular clypeal area and 2+2 setae on the Schizoribautia seydi, TURK, 1955 - Proc. Zool. Soc. London 125: 485-487. postero-central partof theanteriorhalf (Fig. 167).surfaceof clypeal areadenselyreticulated(Fig. 168). Ribautia seydi, CHAMBERLIN, 1955-1956 - Acta Univ. Lund Avd.2 N.S.51(5): 17. Labrum: midpiece well developed with 13 teeth, the more external longer than the central ones. Schizoribautia seydi, CHAMBERLIN, 1957 - Proc. Biol. Soc. Wash. 70: 28. Sidepieces with ca. 9+10 hyaline filaments (Fig. 1ó9). Ribautia (Schizt¡ribautia) seydi, KRAUS, 195'7 - Senck. biol. 38(5/6): 378. Mandible: the mandibles are not dissected from the head capsule in the original slide of KRAUS, their Remarks. - Contrary to the opinion of KRAUS ( 1 957: 378) that Schizoribautia peruana VERHOEFF position on this slide does not allow us to observe the number of teeth of the pectinate lanrellae. and Ribautia (Sthircribautia) mr¡ntana KRAUS are synonyms of this species, we regard the characters First maxillae without palps on both coxosternum and telopodites. Coxosternum without setae; nredian given in table 3 as sÍrong enough k) support the validity of these three species. projections of coxosternum subtriangular, well developed and proviclef titicaca¿, which TURK considered as a with ca. 34 claviform sensory setae on the extemal border and only I on the inrernal border (Fig. 194). subspecies of peruana, it is reasonable to suspect, even though TURK did not specify it, that in titicacae Distal end of this a.a. with ca. 8 very small specialized setae apparently not divided apically (Fig. 195). there are 3+3 clusters of coxal organs. For differences between R. (5,). montona and R. (5.) peruana see Dorsal and v. surfaces of a.a. ll, V, IX and XIII with very snrall specialized setae which on the v. side are table 3. restricted to an intemal latero-apical area and are very similar to those of the apex of the terminal a.a. but Type locality. - Peru: Puno (m. 3900 a.s.l.). Only confidently known from this locality. a little thicker and shorter. Each ofa.a. II (Fig. 190), V, IX and XIII (Fig. 191) with I seta. Specialized setae on d. side are restricted to an extemal latero-apical area and are represented by two types: a and b. Type a setae are very similar to the specialized setae on the v. side; type å setae are equal in shape to Family Schendylidae thcrse ofthe apex ofthe terminal a.a. but darker (pale ochreous) in colour (a, b,Fig.193). A.a. II (Fig. 192)andVwithltype¿seta;a.a.lX(Fig.l93)andXlllwithltypeaandltypeåseta. Genus Pectiniungøis BOLLMAN, 1889 Cephalic plate longer than wide (ratio 1 3:1), shape and chaetotaxy as in Fig. 196. Clypeus with 1+1 post antennal setae, l0+8 median setae and l+l praelabral setae (Fig. 197). Diagnosis. - Pleurites of second nraxillae not fused with coxostemum; apical claw of second maxillae Labrum with l5 robust, dark and round-tipped teeth, sidepieces with l4+15 teeth very different in pectinate on both d. and v. edges. Stema with pore fields. Last pair of legs with seven podomeres; shape, size and colour from those of the midpiece, looking like long hyaline filaments (Fig. I98). praetarsus in form of a small. pilose tubercle or replaced by a small spine or altogether absent; coxopleura Mandible: dentate lamella subdivided into three distinct blocks, L mandible with 3,3,7 teeth, r. ofthe last leg-bearing segment each with two intemal coxal organs ofcomposite structure ("heterogeneous nandible with 3,3,5 teeth (Figs. 199-200); pectinare lamella with ca.26 hyaline teeth. coxal glancls" sensu BRÖLEMANN & RIBAUT, l9l2). First nraxillae with well developed palps on both coxosternunr and telopodites (Fig. 202). Coxostemum with 7 setae arranged as in Fig. 201; median projections ofcoxostemum subtriangular, well developed ancl provided with 3+3 setae. Article II of the telopodite with 5+5 v. sctae and 9+12 d. sensilla (Figs. 201-202). Pectiniunguis ducalis n.sp. (Figs. 188-225) Second maxillae (Figs.201,203-204) with l4+17 setae on the coxosternum, arranged as in Fig.201. Diagnosis. - A Pectiniungu ls species with pore fields present on anterior and posterior sterna. Among Apical claw of telopodite well developed, bipectinate, v. edge with ca. l0 teeth (Fig. 20a), d. edge with ca. 12 teeth. the Neotropical species currently included in the genus Pectiniunguis, it seems more closely related to P. galgei (CHAMBERLIN, l92l). P. ducalis n. sp. can be differentiated fronr this species by means of the plateF woricthip ualna eir:r ewghuelanr ctlroasnesdv,e rtshee mteelodpiaond irteosw doof 6no lta ergxete snedt abee ayonndd a tfheew a andtedriitoiorn narla srgnilna lolefr tsheet ahee asccla; tbtearseadl following characters (the corresponding ones in P. gaigei are given in parentheses): teeth of lateral pieces on the posterior half. Trochanteropraefemur of telopodite with a snrall round-tipped tooth on the apical of labrunr very different in shape, size and colour from those of the rnidpiece and resembling long hyaline medial edge; remaining articles without teeth. Calyx of poison gland cylindrical (Fig.206). Chaetotaxy of filanrents, Fig. I 98 (teeth of the lateral pieces snrall, subtriangular with a sharp medial extention); I with coxosternunì and telopodites as in Fig. 205. 65,67 or 69 pairs of legs, d with 65 or ó7 pairs of legs (9 with 55 or 57 pairs of legs, ct with 53 pairs Legs (last pair excepted) with chaetotaxy (Fig. 207) unifornr throughout the body length; claws of legs); ventral pores prescnt liom second to penultimate sternum (frorn second to antepenultimate); last ventrobasally with one anterior spine and two posterior spines of different size (Fig. 208). Ieg-bearing segnìent without pleurites at the sides of praetergum (with pleurites). Stema: pore fields present from the second tr¡ penultimate sternunì. All pore fields undivided and HoloTtyyppee ndt,a6te7ri apl.. |.-, Abl.ll . s5p2e cnim¡men,s 1 f0r.o1m1. 1B9r8a2z. ilA: Allontlayzpoen a9s,: 6R9e spe.1rv.,a b F.ll.. 5A2. Dnrumck, e6, .9J..1 W98. 3d.e P MarOaRryApeIS , 4l e(g9i)t,. sstuebmciarc: uolna r.s tFeronrunmr o lfl ,f iSe6ld pso crehsa;n ognin gV aI,l olnl9g ; fhoen tXruInVk, a1s4 2in; oFnig Xs.X 2I0I,9 -125155; . oNnu LnrIbIIe,r5 ol;fo pnor eLsX oVn, slel3le;c toend 69 p.1., b.l. 44 mm, 8.9. 1982. Paratype B (è), 67 p.|., b.1. 32 nrm, 1 3.3. 1 983. Paratype C (c¡), 65 p.1., b.l. LXVI,9I. 36 mm, 6.9. 1983. - Depository of typcs. - INPA (holotype, allotype); MLP (pararype,,l); AM (paratype B); Last leg-bearing segment without pleurites at the sides of praetergunt. Praesternunt not divided along JA (paratype C). the sagittal plane; shape and chaekrtaxy ofsternum and tergunr as in Figs.216-217. Coxopleura slightly Other nraterial examined. - Sanre locality ancl collector as the type series. I fragnrentary specimen, protruding at their distal v. ends, setae snraìl and nunrerous on the distal v. half, the renraining surface with 338 339 I l few larger setae. T\ryo compound ("heterogeneous") coxal organs on each coxopleuron (Figs. 216,218). mm (32 mnt); length of specialized setae on the apex of the terminal a. a. conspicuously Ionger than the coxal organs open on the membrane between coxopleuron and stemum, covered by the latter (Fig.2ló). clavifornr setae, Fig. 235 (distinctly shorter than the claviform isetae); pa¡ps of coxosternum of first Last legs moderately inflated, with seven podomeres, shape and chaetotaxy as in Figs. 216-217. Praetarsus nraxillae very small (well developed); all sterna with pore fields lacking additional pores at the sides ofthe as a very small tubercle with ca.9-13 small apical spines (Fig.2l9). nrain pore area (some stema of anterior region of the body with a few additional pores at the anterior sides Terminal segments: internrediate tergum with posterior margin convex, intermediate sternum with of the main pore area); last leg-bearing segnìent with pleurites at the sides of praetergum (without posterior margin ntedially slightly convex; first genital stemum with posterior margin medially slightly pleurites); anterior coxal organs internally with a single area of sper:ialized epithelium, posterior coxal convex, laterally slightly concave (Fig. 2ló). Gonopods biarticulate, basal article wilh ca. l8 setae and organs with ca. 2-3 areas of specialized epithelium (anterior and post:rior coxal organs internally with a apical with ca. I I setae (Fig. 220), penis dorsally with 1+1 apical setae (Fig.22l). single specialized area). Female allotype. - 69 pairs of legs, body length 52 mm, maximum body width 1.60 mm. Among the remaining Neotropical species of the genus, S. con¡inuus n.sp. seenìs to be also closely All features similar to those in the male except for the shape and chaetotaxy of the last leg-bearing related t() S. lesnei BRÖLEMANN & RIBAUT, l9l 1 , but it can be difièrentiatetype presents all the characters of an aclult specimen, including the Number of pores on selected sterna: on sternum II, 2l pores; on III, 29; on IV, 34; on V, 40; on vlll, 46; presence of spermatccae full of spermatozoa and also the presencc of nrature eggs. The two spermatecae on X, 54; on XI, 45; on XV, 16. are very clearly visible by transparence at level of segments XXXIX and XL. Last leg-bearing segnrent without pleurites at the sides of praetergunr. Praesternum not divided along Our only nrale specinten,is an immatr¡re one, so it seems to us inappropriate to describe or illustrate the sagittal plane; shape and chaetotaxy of stemunr and tergum as in Figs. 288-289. Coxopleura slightly here the nxrrphology of the last leg-bearing segment and terminal segnìents. protruding at their distal v. ends, setae snrall and nunlerous on the distal meclial eclge, the remaining surface with few and larger setae. Two single ("honrogeneous") coxal organs on each coxopleuron (Fig. 290). Coxal organs open on the membrane between coxopleuron and sternum, covered by the latter (Figs. Schendylurus janauarius n. sp. (Figs. 266-293) 288,290)' Last legs with seven podomeres, shape and chaetotaxy as in Figs. 288-2gg. praetarsus as a very snrall tubercle with 3-4 small apical spines (Fig. 291). Diagntrsis. - A Sthendylunrs species lacking pore field on the first sternum and with pore fields Terminal segments: intermediate tergum with posterior nrargin convex, internre¿iate sternunl with present only on s()nle anterior sterna. Among the Neotropical species of the genus which share these traits posterior margin slightly concave; first genital sternum with posterior nrargin medially convex, laterally the ptesent s¡recies seems 1() bc very closely related t<¡ S. andesicola CHAMBERLIN which is distributecì slightly concave (Fig.292). Gonopods biarticulate, basal article with ca. 7 serae and apical with ca. 4 setae in the high Andes of Equator at about rn 4000 a.s.l. Differential characlers are given in table 5. (Fig. 29-l), penis dorsally with 0+l apical serae. Type nraterial. - Holotype d,43 p.1., b.l.2l nrrn; paratype d,43 p.1., b.l. 17 nlm. Brazil: Amazonas: Female. - Unknown. Lago Janauarí, ntixedwater inundatioÍr forest, K, 14.09.1987, J. ADIS legit. - Depository of types: Il,lpA. Etymology. - The name of this species refers to the type krcality. f)escription Male holotype. - 43 pairs of legs, body length 21 nrnr, maximunt bocly width 0.6 mm. Colour (of Schendylurus iguøpensís VERHOEFI', l93B (Figs, 294-299 Table 6) prescrved specimen in alcohol) yellowish. Antennae ca.4.7 times longer than the cephalic plate, distally slightly attenuate; all articles, the first Schendylurus iguapensis VERHOEFF, 1938 - Zool. Jahrb., Sysr. 7l:37t. cxcepted, longer than wide. Sctae on a.a. I-lll ofdifferent lengths and few in number, those ofrenraining Sthendylurus iguapensis, BÙCHERL, l94l-42a - Mem. Inst. Butanran l5: 203. articles progressively shorter and nrore numerous towards the tip of the appendage (Figs. 266-267). schendylurus (schendylurus) iguapensis, BÜcHERL, l94l-42b -Mem. Insr. Butanran l5: 349. Terntinal a.a. with ca. l7 clavifornì sensory setae on the extemal border and ca. 3 on the internal border. schendylurus (schendylotyn) iguapensis, ATTEMS, l94j - A,nn. Naturhisr. Mus. wien 55: 87. Distal end of this a.a. with ca. 5 very snrall specialized setae apparently not divided apically. Dorsal and Schendylurus iguapensis, CRABILL, 1972 - Proc. ent. Soc. Wash. 74( I ): 20. v. surfaces ofa.a. II, V, IX and XIII with very snrall specialized setae which on the v. side are restricted schendylurus iguapensis, PEREIRA & MINELLI, 1993 - Tropical zo

Description of nine new centipede species from Amazonia and related matters on

Similar Description of nine new centipede species from Amazonia and related matters on