The correct type locality of Cyaniris ladon var. quesnellii Cockle, 1910 (Lycaenidae), with designation of a lectotype

3 Pages·2003·1.9 MB·English

by  Crispin S Guppy

150 Journalofthe Lepidopterists' Society
Literature Cited Tietz, H. M. 1952. The Lepidoptera ofPennsylvania, a manual.
Penn. StateCollegeSchoolofAgriculture,Agric. Expt. Station,
Curtis, T. 1959. VegetationofWisconsin. UniversityofWiscon-
sinPJ.ress, Madison,Wisconsin.657pp. StateCollege,Pennsylvania. 194pp.
Gleaosfonn,orHt.heAa.st&erAn.UCnriotneqduiSsttat.es19a9n1d.aMdjaancueanltoCfanvaasdcau.la2rnpdlanedt.s Andrew H. Williams, Dept. ofEntomology, Uni-
NewYorkBotanicalGarden,Bronx,NewYork.910pp. versityofWisconsin, Madison, Wisconsin53706, USA.
Mathias, M. E. & L. Constance. W1944^5. Family2. Umbellif- Email: [email protected].
erae,pp.43-295.In Rickett,H. (ed.).NorthAmericanflora.
Vol. 28B. Umbellales, Cornales. NewYork Botanical Garden,
Bronx,NewYork. Receivedforpublication29March2002;revisedandaccepted4De-
SCOTT, A. 1986. ThebutterfliesofNorthAmerica. StanfordUni- cember2002.
J.
versityPress,Stanford,California.583pp.
SCUDDER, S. H. 1889. ThebutterfliesoftheeasternUnitedStates
and Canada with special reference to New England. Vol. 2.
Publ.bytheauthor,Cambridge, Massachusetts. 1774pp.
JournaloftheLepidopterists'Society
57(2),2003,150-152
THE CORRECTTYPE LOCALITYOF CYANIRIS LADONVAR. QUESNELLIICOCKLE, 1910
(LYCAENIDAE),WITH DESIGNATION OF A LECTOTYPE
Additionalkeywords: Celastrina,nigrescens,lucia,BritishColumbia.
Theoriginal descriptionof"Cyanirisladon, Cramer, and Ahbau Lake in the summer, and trapped in the
var. Quesnellii"was based on two specimens taken "at area during the winter. Apparently Cockle misread
Bala Lake, Quesnelle, northern B.C." (Cockle 1910). "Au Baw" as "At Bala". Ahbau Creek was labeled on
Cockle also statedthathethoughtitwould"provealo- maps as Graveyard Creek until 1921 (Janet Mason
cal race which will be found abundant in the Ques- pers. com.), hence the alternative name Graveyard
nelle Valley". We recently had the opportunity to ex- Lake. Ahbau Lake is about40km (25 miles) northeast
amine the type specimens ofthese butterflies in the ofdie moderntownofQuesnel, apparentlycontradict-
Canadian National Collection of Insects and Arthro- ingthe "35 miles N.W." indicated on the specimen la-
pods (Agriculture Canada, Ottawa, Ontario, Canada). bel. However, Ahbau Lake is 35 miles northwest of
The two specimens and their data labels are shown in Quesnelle Forks, asettlement (nowhistoricsite) atthe
Fig. 1, with lectotype and paralectotype designations junction ofthe Cariboo Riverwith the Quesnel River.
providedbelow. Ahbau Lake is at elevation 2950 feet, not 2480 feet,
The two specimen datalabels are in differenthand- butsucherrorsinelevationwerecommonatthattime.
writings. "J.M. Anderson"ontheparalectotypelabelis Ahbau Lake is not in the Quesnel River valley, as
written as ifit is a signature and the date is written in impliedbyCockle, andis inwhatis nowconsideredto
full. Thelabelonthelectotypeisprinted, thedateuses be central, rather than northern, British Columbia
Roman numbers for the month, and part ofthe data ("northern" is ofcourse arelative term). Ahbau Creek
on the other label is omitted. This suggests that M. is part ofthe Cottonwood/Swift Riverwatershed, the
J.
Andersonwrote the paralectotype label, and someone watershed immediately north of the Quesnel River
elsewrote theotherwhenthe specimenswerepinned. watershed. The correcttype localityis therefore "[Ah-
Dr. Fletcheris more likelythan Cockle forthe second bau] Lake, [elevation 2950 feet], [latitude 53°14', lon-
label, because of Cockle's error in reading "Aubau" gitude 122°07',] 35 miles northwest of Quesnelle
Lake as "At Bala" Lake (below). [Forks], B.C.[, Canada]", with interpolated and cor-
The spelling of the lake name on the label of the rected datashown inbrackets andthe coordinatesbe-
paralectotype can readily be seen to be "Au Baw" ingforthe outlet at the south endofthe lake.
Lake, with the alternative name of"Graveyard Lake". Thereisasecondlocalitylabelattachedtoone spec-
"Ah"is"Mr."inChinese,hencethe lakename referred imen, specifying Kaslo as the collection site. The date
to the Chinese Mr. Bawor Bau (alternative spellings). onthis labelis inadifferenthandwritingthanthe date
Formanyyearsheprospectedandworkedgoldclaims on the other two data labels, indicating that a third
on and around what are now known as Ahbau Creek personwrote it. Celastrina ladon lucia (the true iden-
Volume 57, Number 2 151
Lectotype Paralectotype
Cat* Aa u, \Asfzc <s-U>
.t,< <•
Coll,J.W.Cockle
Databased for 4
PARATYPE
The Butterflies of
Canada Projeel
?f£
1998-1999 No.
Fig. 1. ThetypespecimensofCyanirisladon,Cramer,var.QuesneUiiCockle, 1910.
tityofthe types, see Fig. 1) does notoccurnearKaslo, quesnellii is correctly placed as a synonym of lucia
so the label must be in error. Perhaps itwas intended Kirby, 1837.
to indicate that the specimens were part of Cockle's Alsoofinterestisthephrase"F &C". This indicates
collection (Cocklelivedin Kaslo). This extralabelmay that Cockle (or the person who wrote the label) con-
have contributed to the erroneous association ofthe sidered quesnellii to have been described by two
name quesnellii with nigrescens Fletcher, 1903, which people, with the initials presumablybeing an abbrevi-
is discussedbelow. ation of"F[letcher] & Cfockle]". Cockle had submit-
The two specimens from which Cockle described ted the specimens to "the late Dr. Fletcher", who had
quesnellii had labels indicating J.M. Anderson col- providedcomments onthem, butthe originaldescrip-
lected them on 8 June 1907. One ofthe labels reads tionisclearlythatofCockle aloneandhence Cockleis
"Cyaniris Quesnellii Type F & C." The designation of the sole author. Perhaps Cocklewrote the labelswhile
"Type" on this label has no bearing on the question of Dr. Fletcherwas stillalive,withtheintentionthatthey
the type status ofthe specimen, even though Cockle would co-author the description, but then assumed
mayhavewritten the label. The InternationalCode of sole authorship after Dr. Fletchers death.
Zoological Nomenclature (International Commission Blackmore (1920) lists Lycaenopsis pseudargiolus
on Zoological Nomenclature 1999) requires that the race nigrescens form quesnelli [sic]. McDunnough
designation of a type must occur in the original de- (1938) follows Blackmoreinlisting"formquesnelii [sic]
scription, anduseoftheword"type"onaspecimenla- Cockle"under"Lycaenopsispseudargiolusnigrescens",
beldoesnotmakethat specimentheholotype (Article with "niaculata-suffusa Cockle" as an infrasubspecific
72.4.7). The two specimens are syntypes, rather than synonym. Comstock and Huntington (1963) list ques-
havingthe statusofholotypeandparatype asindicated nelliiwiththecorrectspelling, andcite McDunnoughs
on the existing specimen labels, because Cockle did taxonomic placement. Dos Passos (1964) apparently
notspecifyasingle "type" or"holotype"inthe original copied McDunnough (1938) inplacing"formquesnelii
description. Accordingly, underArticle 74ofdie Inter- [sic] (Cockle), 1910" as asynonym ofCelastrina argio-
national Code ofZoological Nomenclature we hereby lus nigrescens (Fletcher), 1903. The listings by Black-
designate one specimen (the one with the existing more, McDunnoughanddos Passoshadseveralerrors.
"type" label) as the lectotype and the other as the First, they use two incorrect spellings of the taxon
paralectotype, as shown in Fig. 1. The taxonomic pur- name. Second,quesnelliiwas clearlydescribednotas a
pose of this lectotype designation (ICZN Article form but as a geographically defined variety (=sub-
74.7.3) istoclarifythatthenamequesnellii iscorrectly species). This is indicatedbyCockle's statement"diere
associatedwith lucia, ratherthanwith nigrescens, and iseveryreasontothinkdiatifthisvarietyisfoundtobe
to provide future opportunity to determine whether (as I think) adistinctlocal race, it shouldbe entitledto
,
152 Journalofthe Lepidopterists' Society
a specific name". Hence quesnellii is an available be omitted from checklists andotherpublications.
species-group name under the International Code of
WethankJanetMason,ProvincialToponyinist, Base Mapping&
Zoological Nomenclature (1999). Third, the type spec- Geomatic Services Branch, BC Ministry ofSustainable Resource
imens, and all the numerous specimens ofCelastrina Management for information on the historical names of Ahbau
thatGuppyhascollectedinthevicinityofQuesnel, are CreekandAhbauLake,andthesuggestionthat"Quesnelle"mayre-
fertoQuesnelleForks.
clearly referable to lucia (Kirby), 1837 and not to ni-
grescens (Guppy collected the nearest nigrescens 120 Literature Cited
km south ofQuesnel atWilliams Lake in 2002). Miller Blackmore, E. H. 1920. The Lycaenidae of British Columbia.
and Brown (1981) repeated the error ofplacingques- Proceedingsofthe EntomologicalSocietyofBritishColumbia
(1919)14:5-11.
nceolrliriecatseadstyhenosnpyelmlionfgnaingrdesccoernrsecrtaltyhterretahteadntluhceian,abmuet Cockilnen,oJr.tWhe.rn19B1r0it.isNhoCtoelsumobniaa.feCwanb.utEtenrtfolmieosl.fo4u2n(d6)a:t2K0a3-s2l0o4a.nd
as an available species-group name. Guppy and Shep- Comstock,W. P. &E. I. Huntington. 1963. Anannotatedlistof
the Lycaenidae (Lepidoptera: Rhopalocera) of the Western
ard (2001) placed quesnellii as a synonym ofC. ladon Hemisphere(continued). NewYorkEntomol. Soc.21:45-57.
lucia, and abbreviated the type locality to "Quesnel, DosPassos,C.F. 1964. AsynJ.onymiclistoftheNearcticRhopalo-
B.C." because at the time Guppy had not seen the cera. Lep. Soc. Mem. l:i-v, 1-145.
specimenlabels andhencecouldnotdeterminethelo- Guppbyi,a.C.UBS.C&PJr.esHs.VSahnecpoaurvedr.,2B0.0C1..41B4uttpepr.fliesofBritishColum-
cation of"Bala Lake". International Commission on Zoological Nomenclature.
An additional name is mentioned by Cockle (1910), 1999. International code ofzoological nomenclature. 4th ed.
in the sentence "I submitted them [the specimens of xTxhiex+In3t0e6rnpapt.ionalTrustforZoologicalNomenclature, London,
quesnellii] tothe late Dr. Fletcher,whowrote methat, McDunnough, J. H. 1938. Check list of the Lepidoptera of
hadtheybeen takenin Ontario, hewouldhave named Canadaandthe United States ofAmerica. Part 1. Macrolepi-
them 'macidata-suffusa'." Clearlythis name is not be- Milldeorp,teLr.a.D.Me&mF..CMa.lifB.rAocwand.. S1ci9.811.:3A-2c7a2ta(l1o-g3ue(/ccohrercikgleinsdta)o)f.the
ingformallyappliedto the specimens in question, not butterflies of America north of Mexico. Lepid. Soc. Mem.
even by Dr. Fletcher. It is clear that Cockle used the 2:i-vii, 1-280.
namequesnellii insteadofthe namemaculata-suffusa
not in addition to that name. McDunnough (1938), Crispin S. Guppy, 4627 Quesnel-Hydraulic Road,
Dos Passos (1964) and Miller and Brown (1981) were Quesnel, BC V2J 6P8 Canada and Norbert G.
in error to list "maculatasujfusa (Cockle)" as a syn- Kondla,P.O. Box244, Genelle, BC V0G1G0, Canada
onym ofquesnellii. The name maculatasujfusa has no Receivedforpublication 5 March 2002; revised and accepted 20
standingeven as an infrasubspecific name, andshould September2002.
JournaloftheLepidopterists'Society
57(2),2003,152-153
FIBST BEPOBT OF THE PALAEABCTIC DICHRORAMPHAACUMINATANA
(LIENIG &ZELLEB) IN NOBTH AMEBICA (TOBTBICIDAE)
Additionalkeywords: immigrant,holarctic,Olethreutinae,Dichroramphapetiverella, D. vancouverana.
In the course ofan ongoing inventory ofthe moths Initial identification ofthe specimen was based on
ofSteuben,WashingtonCo., Maine, asinglespecimen figures of wings and genitalia in Bentinck and
ofthe Old World olethreutine Dichrorampha acunii- Diakonoff (1968) and Kuznetsov (1987), and con-
natana (Lienig & Zeller) was captured in 2001, evi- firmedbycomparisonwith authenticPalaearcticspec-
dentlyafirstrecordforNorthAmerica. Thespecimen, imens listed below. The species is distinguished from
afresh male (Figs. 1, 3), was takenonadoorscreen at similar Nearctic forms by the acuminate shape ofits
approximately 1600 h EDST on 15 June at forewing (signalized in its name), the continuous pale
44°30'22"N, 67°59'28"W. Nothing is known ofits ori- band in its terminal fringe, its diffuse dorsal patch, its
gins, butas areportedrootfeederonChrysanthemum broad cucullus with blunt ventral cusp, and its bifid
leucanthemum L. and C. segetum L. (Asteraceae) aedeagus terminating in a distinctive open trough
(Bentinck & Diakonoff1968, Kuznetsov 1987), it can (Figs. 1-4). It belongs in the nominate subgenus in
be presumed to have developed on naturalized food- lacking anellar lobes but possessing a male forewing
plants presentwithin 1-2 km ofthe collection site. costal fold.