New genera and combinations in the Senecioneae of the Greater Antilles

2006·7.2 MB·

by  Bertil Nordenstam

5 Comp. Newsl. 44,2006
New genera and combinations in the Senecioneae
ofthe GreaterAntilles
BertilNordenstam
DepartmentofPhanerogamicBotany,SwedishMuseumofNaturalHistory
P.O.Box50007,S-10405Stockholm,Sweden
bertil.nordenstam(a)nrm.se
Abstract
GenericlimitsinthetribeSenecioneaeoftheGreaterAntillesarerevised.Sevennew
genera are described: Antillanthus B. Nord., Herreranthus B. Nord., Lundinia
B.Nord.,ElekmaniaB.Nord.,NesampelosB.Nord.,LeonisB.Nord.,andZemisia
B.Nord.Altogether33 new combinations are made.Asaconsequencethereare no
indigenous species ofSenecio L. and Pentacalia Cass, in Cuba, Hispaniola and
Jamaica.
Introduction
Thepresentstudyispartofanongoingrevisionofgeneric limits inthe Senecioneae
ofthe GreaterAntilles, particularly Cuba and Hispaniola. It has been necessary to
distinguishanumberofnewgenera,someofwhicharestillinsufficientlyknowninall
details and deserving a closer study. The treatment ofthe tribe Senecioneae in the
GreaterAntilleshasuptonoworrecentlybeenveryconservative,withmostspecies
remaininginanover-expandedconceptofSenecioL.,plusanumberoftransferstothe
poorly understoodand ill-defmedgenusPentacaliaCass.
The Jamaican Senecioneae were sorted out in a preliminary way by Nordenstam
(1978), who recognized two new endemic genera, viz. Odontocline B. Nord. and
Jacmaia B. NoRD.These were accepted by H. Robinson in Proctor (1982), who
howeveraddedthegenusPentacaliawith threeJamaican species (two describedas
newandonetransferredfromSenecio).
IntheFloradeCubavol.5(Alain 1963)thetribeSenecioneaewasrepresentedbyfour
genera.Three ofthese arequite small, namelyErechtites with one species (plusone
variety), Emilia with two species, and the monotypic and endemic Shafera. In
Senecio21 specieswererecognized, includingthetypespecies,S. vulgaris,whichis
awidespreadannualweed,although uncommoninCuba.Theother20specieslisted
underSenecioareall indigenousshrubsorvines,mostofthemendemictoCuba,but
twoofthemalsooccurringonHispaniola(S.plumbeusandS. trineums). Oneofthe
Comp. Newsl. 44, 2006 51
CubanSenecios,viz.S.almironcillo,wastransferredtothegenusPentacaliain 1982
byProctor,andanotherwasrecentlydescribedasthemonotypicOldfeltiaB.Nord.
&LuNDiN(NoRDENSTAM&LuNDiN2002).Theremainderarenowdisposedinfournew
genera, andonlytheoccasionalweedS. vulgarisremains inthegenus inthe Cuban
flora.
InHispaniola(Haiti andtheDominicanRepublic)threesmall endemicgenerahave
beenrecognized,viz.HerodotiaUrb.,MaltfeldiaUrb. andEkmaniopappusBorhidi.
Twelvespeciesofshrubs, smalltreesorvineshavebeentraditionallybeenplacedin
Senecio(Liogier 1996).Oneparticularlydeviatingtaxonisdescribedelsewhereasthe
new genus Igmirbia B. Nord. (Nordenstam 2006). Further genera ofthe tribe in
HispaniolaarethemoreorlessweedyErechtites,Emiliaand Crassocephalum.
Some ten years after Robinson and Proctor, Borhidi followed their track and
transferred anothereleven Cuban species and one from the Dominican Republic to
Pentacalia. He also described a new species from Cuba as Pentacalia acunae
Borhidi,relatedtoP.almironcillo(Borhidi 1992). Noneofthesetransferredspeciesis
wellplacedinPentacalia,whichinthebroadsenseofRobinson,Cuatrecasasa.o.,is
a largeAndean genus with a northern limit in Costa Rica, where the northernmost
outposto^paramovegetationisfound.WithinthebroadconceptoiPentacaliathere
are(atleast)twodistinctgroups,treatedassubgenerabyCuatrecasas,Robinsonand
others, but better treated as genera, as by Jeffrey, Bremer and myself Pentacalia
sensustrictocomprisesvinesorepiphytes,whereasMonticaliaC.Jeffreyisagenus
oferect shrubs with closely set leaves and terminal synflorescences. These two ge-
neraboth haveanAndean distributionandarenotrepresented intheCaribbean.
Thepurposeofthepresentpaperistobrieflydescribethenewgeneraandmakethe
necessarynewcombinations.Aflillertreatmentwithkeysanddiscussionswillfollow,
andaglobaloverviewoftheSenecioneaewithagenerickeywillbepublishedinthe
FamiliesandGeneraofVascularPlants (Nordenstam, inpress).
Newgeneraandcombinations
AntillanthusB.Nord.,gen.nov.
Fruticeserectivelscandentes.Foliacaulinasessiliavelbreviterpetiolatalineariavel
lanceolataadoblongo-ovatasupraglabrasubtustomentosa.Capitulapaucavelplura
corymbosa-cymoso-paniculata discoidea vel interdum radiata. Involucri bracteae
uniseriatae.Receptaculumalveolatumglabrum.Flosculiradiifemineivelnulli.Flosculi
discihermaphroditi;corollaresinifera,lobisovatis.Antheraebasisagittataevelbrevi-
tercaudatae.Styliramilinearesintusareisstigmaticisconfluentibusapiceobtusipilis
brevibus sparsis. Cypselae elliptico-oblongae 8-10-costatae pubescentes. Pappi
setae numerosae basi connatae albae persistentes.
52 Comp. Newsl.44, 2006
Typus: Antillanthiis ekmanii(Alain) B. N(jri).
Erectorscandentshrubs0.5-2m;branchesmostlywhite-tomentose.Leavescauhne,
sessileorshortlypetiolate,(linearor)lanceolatetooblong-ovate,3-13 cmlong, 1-4
cm wide, entire or sometimes shallowly sinuate-dentate, glabrous (or at least
glabrescent) above, densely white- or grey-tomentose beneath; margins usually
revolute. Capitula few to several in terminal corymbose or cymose-paniculate
synflorescences, discoid or rarely radiate, yellow-flowered or rarely white-cream-
coloured; peduncles bracteate. Involucre cylindrical to narrowly campanulate,
calyculate; involucral bracts uniseriate 5-13, oblong-lanceolate, obtuse-acute.
Receptacle glabrous, alveolate. Disc-florets 5-13; corolla tubular below, narrowly
campanulate or funnel-shaped above; lobes deltoid to narrowly ovate, midlined,
apically subcucullate and papillate. Resin canals often present in corolla and style.
Anthers basally sagittate or shortly caudate; apical appendage ovate, obtuse;
endothecial tissue radial with elongated cells; filament collar balusterform. Style
branchesapicallyobtusewithratherfewandshortlateralsweeping-hairs;insidewith
stigmatic areas continuous. Cypselas elliptic-oblong, 8-10-ribbed, shortly hirsute.
Pappus bristles numerous, slender, usually shorterthanthecorolla, basallyconnate,
white, persistent.
This isthelargestgenusofthetribe inCuba,with 17 speciespresentlyrecognized.
X.Antillanthusacunae(Borhidi)B.Nord.,comb.nov.
Basionym: Pentacalia acunae Borhidi, Acta Bot. Hung. 37: 88 ("1992" = 1994).
Type: J. Acuna 13412 SV, Cuba, Prov. Las Villas, Sierra de Escambray, in saxosis
calcareismontisPicoPotrerillo,900—930m,V.1939(HACholo.!).
2.Antillanthiisalmironcillo(MazaGomez)B.Nord.,comb.nov.
Basionym:SenecioalmironcilloMazaGomez,AnalesHist.Nat. 19:277(1890).
G
Syn.:Pentacaliaalmironcillo(MazaGomez) R.Proctor,J.ArnoldArbor.63(3):
312(1982).
Furthersyn.: CacaliadiscolorGriseb.,non DC;Senecio brittoniiGreenm., Field
Mus.Nat.Hist. 164,Bot.Sen2(8):323(1912).
Type:C.Wright2870,Cuba.
"i.Antillanthiisazulensis(Alain)B.Nord.,comb.nov.
Basionym:SenecioazulensisAlain,Contr.Ocas. Mus. Hist.Nat.Colegio"DeLa
Salle"no. 18: 10(1960).
Type:Alain&M. LopezFigueiras7352,Cuba,Oriente,SierraAzul,Barracoa,
4.L1960 (LSholo.,SV,"UO"iso.).
Comp. Newsl. 44, 2006 53
4.Antillanthiisbiseriatus(Alain)B.Nord.,comb.nov.
Basionym:SeneciobiseriatusAlain,Contr.Ocas.Mus. Hist.Nat.Colegio"DeLa
Salle"no. 18:11(1960).
Type:Alain3429,Cuba,Oriente,SierradeMoa,25-26.VII.1953(LShole,NYiso.).
5.Antillanthuscarinatus(Greenm.)B.Nord.,comb.nov.
Basionym:SeneciocarinatusGreenm.,FieldMus.Hist.Nat. 164,Bot.Ser.2(8):323
(1912).
Syn.:Pentacaliacarinata(Greenm.)Borhidi,ActaBot.Hung.37:88('i992"=
1994).
Type:J.A.Shafer4079,Cuba,Oriente,SierradeMoa,CampSanBenito,900m,
24.n.l9I0(F,NY).
6.Antillanthuscubensis(Greenm.)B.Nord.,comb.nov.
Basionym:SeneciocubensisGreenm.,FieldMus.Hist.Nat. 164,Bot.Ser.2(8):324
(1912).
Syn.:Pe«toca//<7a//)eA75/.9(Greenm.)Borhidi,ActaBot.Hung.37:88("1992"=
1994).
Type:J.A.Shafer4084,Cuba,Oriente,SierradeMoa,CampSanBenito,900m,
24.11.1910(F,NY).
7.Antillanthusekmanii(Alain)B.Nord.,comb.nov.
Basionym:SenecioekmaniiAlaw,Contr.Ocas.Mus.Hist.Nat.Colegio"DeLa
Salle"no. 18:11(1960).
Type:Alain,Acuna&LopezFigueiras5606,Cuba,Oriente,SierradeCristal,en
charrascos,2-7.1V.1956(LShoio.,NY,SV,"UO"iso).
8.Antillanthuseriocarphus(Greenm.)B.Nord.,comb.nov.
Basionym:SenecioeriocarphusGreenm.,Torreya 13:257(1913).
Syn.:Pentacaliaeriocarpha(Greenm.)Borhidi,ActaBot.Hung.37: 88("1992"=
1994).
Syntypes:J.A.Shafer4149,Cuba,Oriente,fromCampToatoCampLaBarga,22-
26.11.191 (F,NY,MOfragment);J.A.Shafer8257,CampLaGloria,SofSierrade
MO
Moa,24-30.XII. 191 (F,NY, fragment).
54 Comp. Newsl. 44, 2006
9.Antillanthusleucolepis(Greenm.)B.Nord.,comb.nov.
Basionym:SenecioleucolepisGreenm.,FieldMus.Hist.Nat. 164,Bot.Sen2(8):324
(1912).
Syn.:Pentacalialeucolepis(Greenm.)Borhidi,ActaBot.Hung.37:89("1992"=
1994).
Type:J.A.Shafer4146,Cuba,Oriente,CampToatoCampLaBarga,400-450m,
22-26.n.l910(F,NY).
10.Antillanthusmoaensis(Alain)B.Nord.,comb.nov.
Basionym:SeneciomoaensisAlain,Contrib.Ocas.Mus.Hist.Nat.LaSalle 18:11
(1960).
Syn.:/'e/7/flca//awoa6'm75(Alain)Borhidi,ActaBot.Hung.37: 89("1992"= 1994).
Type:Leon&Clemente23203,SierradeMoa,MinaJohnson,20.VII.1947(LSholo.,
NYiso.).
\\.Antillanthusmoldenkei(Greenm.exAlaev)B.Nord.,comb.nov.
Basionym:SeneciomoldenkeiGreenm.&Alain,Contrib.Ocas.Mus. Hist.Nat.
Colegio"DeLaSalle",Habana,no. 18: 12(1960).
Type:Acuna12775,Cuba,Oriente,GranTierra,Moa, 16.IV.1945(HACholo.!)
12.Antillanthuspachylepis(Greenm.)B.Nord.,comb.nov.
Basionym:SeneciopachylepisGreenm.,FieldMus.Hist.Nat. 164,Bot.Ser.2(8):325
(1912).
Type:J.A.Shafer4008,Cuba,Oriente,RioYamanigueytoCampToa,22-26.11.1910
(F,NY).
13.Antillanthuspachypodus(Greenm.)B.Nord.,comb.nov.
Basionym:SeneciopachypodusGreenm.,Publ.FieldMus.Nat.Hist.Chicago,Bot.
2:325(1912).
Syn.:Pentacaliapachypoda(Greenm.)Borhidi,ActaBot.Hung.37:89("1992"=
1994).
Type:J.A.Shafer8186,Cuba,Oriente,SofSierradeMoa,CampLaGloria24-
30.XII.191 (NYholotype,Ffragment).
14.Antillanthussaugetii(Alain)B.Nord.,comb.nov.
Basionym:SeneciosaugetiiAlayh,Contrib.Ocas.Mus.Hist.Nat.LaSalle 18: 12
Comp.Newsl. 44, 2006 55
(1960).
Syn.:Pentacaliasaugetii(Alain)BoR}mi,ActaBotHung.37:89("1992"= 1994).
Type:Leon 12196,SierradeImias,PuntondelMate, 1924(LShole,NYiso.).
15.Antillanthuss/iaferi(Greenm.)B.NoRD.,coinb.nov.
Basionym:SenecioshafenGreenm.,FieldMus.Hist.Nat. 164,Bot.Sen2(8):326
(1912).
Syn.:Pentacaliashafen(Greenm.)Borhidi,ActaBot.Hung.37:89("1992"= 1994).
Type:J.A. Shafer3107,Cuba,Oriente,SierradeNipe,betweenPiedraGordaand
Woodfred,400-500m,8.X11.1909(F,NY).
\6.Antillanthussubsquarrosus(Greenm.)B.Nord.,comb.nov.
Basionym: SeneciosubsquarrosusGreenm.,Ann. Missouri Bot. Gard. 8: 97(1922).
Type:J.Shafer3722,Cuba,Oriente,RioGuayabo,450-550m,21-30.1.1910(GH,NY,
MO
fragment).
\1.Antillanthustrkhotomus(Greenm.)B.Nord.,comb.nov.
Basionym:SeneciotrichotomusGreenm.,FieldMus.Hist.Nat. 164,Bot.Ser.2(8):
326(1912).
Syn.:fe/7racfl//a/r/c/70/owa(Greenm.)BoRHiDi,ActaBot. Hung.37: 89("1992"=
1994).
Type:J.A.Shafer3821,Cuba,Oriente,LomaMenquara,680m,l-3.II.1910(F,NY).
LeonisB.Nord.,gen.nov.
Fruticulus scandens glaber praeter pedunculos et petiolos rufo-asperulos. Folia
altema petiolata ovata ad eiliptica integra vel interdum trilobata trinervia.
Synfiorescentiae axillares breves coiymbosae. Capitula heterogama radiata vel
disciformia Involucrum calyculatum; involucri bracteae uniseriatae lineis resinosis
.
instructae. Flosculi radii feminei flavi; lamina brevi. Flosculi disci numerosi
hermaphroditi,corollaflavaquinquelobata, lobisdeltoideisvelovatisductomediano
resinifero notatis. Antherae basi sagittatae ecaudatae. Styli rami apice obtusi pilis
sparsis brevibus lateralibus. Cypselae elliptico-oblongae glabrae (setis sparsis
apicalibus repertis) decemcostatae stramineae. Pappi setae numerosae albae basi
connatae.
Type:Leonistrineura(Griseb.)B.Nord.
Monotypic:
56 Comp. Newsl. 44, 2006
Leonistrineura(Griseb.)B.Nord.,comb.nov.
Basionym:SeneciotrineumsGriseb.,Mem.Am.Acad.N.S.8:514(1862).
Syn.:Pentacaliatrineura(Griseb.)Borhidi,ActaBot.Hung.37(1-4):89("1992",
prob. 1994).
Type:C.Wright327,Cuba,Monteverde,Guantanamo(S=isotype!)
Furthersyn.:Senecioleonis{""Leonis''')Britton&Wilson,Bull.TorreyBot.Club50:
50(1923).
SeneciodomingensisUrb.,Symb.Antill.7:431 (1912).
Syntypes:H.v.Turckheim3347,Sto.DomingopropeConstanza.
lllustr.:Fig. 1.
Scandentshrub, glabrous inmostparts,butyoungbranches, pedunclesandpetioles
rufous-asperulous, and peduncles sometimes finely glandular-papillate below the
capitulum;stemsstriate.Leavesalternateonshortaxillaryfloweringshoots,ovateto
elliptic, 1.5^cmlong,(0.5-)1-2.5cmwide,entireorshallowlytrilobateorgrossly 1-
4-dentate, petiolate,herbaceous, three-nerved fromnearthebase; leaf-blade basally
cuneate to truncate or occasionally hastate; petiole slender, 1-3 cm long.
Synflorescences corymbose, terminatingthe lateral axillary leafy shoots. Peduncles
slender, 1^cmlong,withafewlinearbracts,asperulouswithrufousshorttrichomes
and sometimes apically papillate-glandular. Capinda campanulate, heterogamous,
radiate or disciform, yellow-flowered. Involucral bracts uniseriate, 8-12(-13),
mm mm
narrowlyoblongtolanceolate,(1.5-)2-7 long, 1-2 wide,distinctly3-5-veined
withblackish(often interruptedly)resiniferousveins,apicallyacuteandpuberulous,
otherwiseglabrousorrarelysomewhatpuberulousthroughout.Calyculusbracts2-5,
mm mm
generallyshorterthanthephyllariesbutvaryinginsize,5-8 long,0.5-1 wide,
linear and acuminate, sparsely pubescent with whitish or short rufous trichomes or
subglabrous,sometimeswithamedianresinvein.Receptacleslightlyconvex,shortly
fimbrilliferous,butnotdenticulateordistinctlyalveolate.Ray-floretsfemale, l-5(-8),
mm
glabrous;tubecylindrical,2-5 long; laminaoblongornarrowlyelliptic-oblong,
mm
1-5 long, 3-5-veined. Style branches linear, glabrous, obtuse, with separated
stigmaticlines.Disc-florets 16-24,hermaphroditic.Corollabasallytubular,widening
mm mm
above, 4.5-6 long, glabrous, shortly 5-lobed; lobes deltoid-ovate, 0.5-0.7
long,withamedian subapicaldarkresincanal,apicallysubcucuUateand±papillate
mm
outside. Anthers 1.5-2.5 long incl. appendage, basally sagittate, but ecaudate;
apical appendage narrowly ovate-lanceolate; filament collar 'balusterform" (with
enlarged cells basally). Style with two dark resin canals, branches linear with
continuous stigmatic area inside; base indurated; stylopodium distinct; style
mm
branches linear-oblong, ca. 1-1.5 long, apically obtusewith few andvery short
lateralsweeping-hairsorsubglabrous.Cypselaselliptic-oblong,glabrous(sometimes
Comp. Newsl. 44, 2006 57
7 SX
Fig.l.Leonistrineura(Griseb.)B.Nord.
APortionofplant,x 1.BRay-floret,x6.CDisc-florets,x6.DCorollaofdisc-floret,laid
out,X6.EStamens,x 12.FStylebranchesofdisc-floret,x 12.GCypselas,x6.(A-FL.
B.Smith&al.3238inS;GEkman 14504inS).Del.B.Nordenstam.
58 Comp. Newsl.44,2006
afewapicalsetae),with 1 distinctstraw-colouredribs,otherwiselightbrown.Pappus
bristlesnumerous,pluriseriate,minutelybarbellate,white,persistent,basallyconnate.
Leonis thnewa is a vine which occurs in Cuba and Hispaniola (both Haiti and
Dominican Republic). The generic name is simply adopted from the epithet ofthe
synonym Senecio leonis Britton & Wilson. Leonis agrees with Antillanthus and
some other genera in having continuous stigmatic areas on disc-floret styles.The
synflorescences are axillary along the rather slender scandent stems.The plant is
stronglyresiniferouswithdistinct,oftenblackishresinductsinphyllaries,disc-floret
corollaandstyle.
When Urban described Senecio domingensis he remarked that it has no related
Antilleantaxa("Nullialiiexantillanisaffmis").Thereisnothingtodistinguishitfrom
S. trineura, however, and it is strange that Urban overlooked this widespread
species.
The species is quite variable, which is perhaps not surprising in view ofits wide
distribution on the two largest islands ofthe GreaterAntilles. The leaf-shape varies
from entire and ovate to more or less distinctly trilobate or grossly dentate. The
calyculusbractsshowmuchvariation inlength, fromveryshorttoahnostthelength
oftheinvolucralbracts.Thenumberandsizeofray-floretsisalsovariable,fromoneor
twotoatleastfiveorrarelyeight, andsomecapitulaarebestdescribedas disciform
becauseofthereducedlaminaoftheray-florets.
Leonis trineuraisrathersimilarinvegetative morphologytoMattfeldiatriplinervia
Urb.,amonotypicgenusendemictoHaiti.Thisisalsoascandentshrubwithalternate
petiolate leaves,whichareovate,entireandthree-veined. Mattfeldiaisdistinctfrom
Leonisbythebilabiateray-floretsandthenarrowandfew-floweredcapitulawithonly
fivephyllaries.
NesampelosB.Nord.,gen.nov.
Fruticesscandentesramistomentosisglabrescentes. Foliaaltemapefiolataelliptico-
ovata ad obovata Integra margine dentata vel denticulata coriacea costata basi
rotundata vel obtusa interdum cuneata apice obtusa mucronata supra glabrescentia
nitidasubtusdensetomentosa. Capitulaparvaheterogamaradiatainsynflorescentiis
axillaribus corymbose disposita. Involucrum calyculatum; involucri bracteae
subuniseriataevelferebiseriataelineari-lanceolataevelangusteoblongaeglabraevel
pubescentes. Flosculi radii pauci lamina brevi flava vel cremea-alba. Flosculi disci
hermaphrodita; corolla supeme campanulata quinquelobata, lobis ovatis vel
lanceolatis apice papillatis. Antherae basi obtusae vel sagittatae. Styli rami apice
subtruncati pilis eferrentibus numerosis, areis stigmaticis separatis. Cypselae
striatae-costatae apicem versus sparse setosae aliter glabrae; carpopodium
Comp. Newsl. 44, 2006 59
distinctum. Pappi setaecopiosaealbaebarbellataebasi connatae.
Lianas or scandent shrublets with tomentose glabrescent stems; trichomes soft and
flexuous, branching. Leaves alternate, petiolate, elliptic-ovate to obovate, 3-7 cm
long,2-Acmwide,entirewithdentatetodenticulatemargins,midribbedandwithlate-
ral veins, coriaceous, base rounded-obtuse or cuneate; apex obtuse mucronate;
upper side initially araneose or loosely tomentose, glabrescent and glossy with a
reticulatevenationpattern,lowersidebrownishorgreyishdenselytomentose;petiole
short,3-10mm.Capitulainterminaloraxillarylateralcorymbs,small,heterogamous,
radiate. Receptacle flat, shortly denticulate with acuminate scales. Involucre
calyculatewithfewsmallsubulatecalyculusbracts;involucralbractssubuniseriateto
almost biseriate, 5-8, linear-lanceolate to narrowly oblong, glabrous ortomentose;
innerbractswithscariousormembranousmargins.Ray-floretsfew(2-5),laminashort,
mm
to 5 long, oblong, yellow, creamy or white, apically 2-3-fid. Discflorets
mm
hermaphrodite, 5-13; corolla 4-5 long, tubular below, campanulate above,
5-lobed; lobes ovate to lanceolate, apically callous-incrassate orpapillate. Anthers
basally obtuse or sagittate, ecaudate; apical appendage narrowly ovate-lanceolate.
Style branches apically subtruncate with numerous short sweeping-hairs; stigmatic
areas separated. Cypselas striateorribbed, ciliate inupperhalf, otherwiseglabrous;
carpopodium distinct. Pappus bristles numerous-copious, pluriseriate, barbellate,
white, basallyconnate,persistent. Three spp., Hispaniola.
Some characteristics ofthe genus are the scandenthabit, the petiolate entire leaves
withdentateordenticulatemargins,theshinyreticulateuppersurfaceandtomentose
lowerside,thesmallandshort-rayedfew-floweredcapitula,pluscharactersofstyles
and anthers.
X.Nesampelosliicens(Poir.inLam.)B.Nord.,comb.nov.
Basionym:ConyzalucensPoir.inLam.,Encycl.Suppl.2:341(1811).
Syn.:Seneciolucens(Poir.)Urban,Symb.Antill.3:413(1903).
Type:Nectouxs.n.,Haiti(P).
Furthersyn.:ConyzadomingensisSpr.,Syst.3:508(1826);SenecioconyzoidesDC,
Prodr.6:412(1838).
2.Nesampeloshotteana(Urb.&Ekman)B.Nord.,comb.nov.
Basionym: Senecio hotteanus Urb. & Ekman, Arkiv f Bot. 23A(11): 93 (1931).
Type: E. L. Ekman H 7430, Hispaniola, Haiti: Massifde la Hotte, western group,
Torbec, Mome Formon, limestone ridge, in thickets, 1400-1500 m, 27.XII.1926
(Sholo.!,iso.!)./
Illustr.:Fig.2.