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Atalanta (August 2000) 31 (1/2):293-326, Würzburg, ISSN 0171-0079
"Fauna lepidopterologica Volgo-Uralensis" 150 years later:
changes and additions. Part 3. Geometridae
(Insecto, Lepidoptera)
by
Vasily V. Anikin, Sergey A. Sachkov, Vadim V. Zolotuhin &
Elena M. Antonova
received 24.II.2000
Summary: 370 species of Geometridae are listed for the modern Volgo-Ural fauna. 7 species
(Euchloris volgaria Guenee, Nemoria melinaria Herrich-Schäffer, Rhodostrophia calabraria
Hübner, Ennomos effractaria Freyer, Ourapteryxpérsica Menetries, Yezognophos dilucidaría
Denis & Schiffermüller and Y. serotinaria Denis & Schiffermüller) are deleted from the list.
Supposedly they were either erroneously determinated since Eversmann's work or they are
considered now of subspecific or infrasubspecific rank. 161 species are recorded from the re
gion in addition to Eversmann's list.
This paper is the fourth in a series of publications1 and deals with the composition of the pres
ent day fauna of geometrid moths in the Middle Volga and the south-western Cisurals. This re
gion comprises the administrative divisions of the Astrakhan, Volgograd, Saratov, Samara,
Uljanovsk, Orenburg, Uralsk and Atyraus (= Gurjev) Regions, together with Tataria and Bash
kiria. As was accepted in the first part of the cycle, only material reliably labelled, and dating
from the last 20 years was used for this study. The main collections are those of the authors:
V. Anikin (Saratov and Volgograd Districts), S. Sachkov (Samara District) and V. Zolotuhin
(Uljanovsk district and southern Tataria). For the same districts we also made use of literature
data, i. e. Astrakhan district (Lvovsky, 1971), Bashkiria (Antonova, 1985; Grosser, 1983, 1987)
and Uralsk district (Ajbasov, 1974; Kuznetsov & Martynova, 1954) as well as distribution maps
of some geometrid species published in "Provisional Atlases of the Insects of the European
Part of the U.S.S.R." (Antonova, 1980-1982). All the data from the 19th and early 20th centu
ries were taken into account but only as a reference (Becker, 1854-1866; Christoph, 1867,
1868; Gross, 1925; Tauscher, 1806, 1809; Jakovlev, 1861; Krulikovsky, 1910; Zhuravlev,
1910). Whilst compiling this list we also took advantage of the information from recent papers
on this region (Kumakov & Korshunov, 1979; Novoderezhkin, 1983; Antonova, 1985; Anton
ova et al., 1990; Lastukhin, 1990a, 1990b; Sachkov et al., 1996; Anikin, 1997; Antonova &
Zolotuhin, 1998), which were in part critically reviewed and revised, and from recent mono
graphs and articles dealing especially with the Geometridae (Viidalepp, 1976-1979, 1988,
1996; Mironov, 1990, 1991).
The collection material from the Zoological Institute of the Russian Academy of Sciences at St.
Petersburg and especially of the Moscow State University (with the original Eversmannos ma
terial-curator is E. M. Antonova) was also examined for our study. Also the private collections
1 Part 1: Atalanta (1993) 24 (1/4): 89-120; part 2: Atalanta (2000) 31 (1/2): 265-292; part 4: Atalanta
(1999) 29 (1/4): 295-336.
293
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of A. & V. ISAJEv(Uljanovsk) and V. Kupayev (Samara) were studied, to whom we express our sin
cere thanks. Special thanks also due to Dr. V. G. Mironov (ZISP, St. Petersburg) for his help in
determination of Eupithecia-species and taxonomic advices during our work. We also owe spe
cial thanks to the curator of the Lepidopteran collection at the Kiev State University, I. Yu.
Kostyuk for his help to our work with the museum collections and determination of some spe
cies, and Dr. Ulf Eitschberger (Marktleuthen, Germany) for his valuable help in obtaining
some literature sources concerning the Geometridae.
In the article, the system of Viidalepp (1996) with some additions according to Wolf (1988)
and Mironov (1990, 1991) was accepted. We have to note, the system of the Geometridae is
not stable at present, thus the variant that has been used by us can differ from the usual ones.
In any case we consider our article not a taxonomic but a faunistic work and hope it would be
useful for investigators of the geometrids not only in Russia but in European states too.
For the ease of use, information is given in the form of a table, with the principal data on all
species mentioned for the Volgo-Ural region. Many localities have been renamed during the
last 150 years, the most important ones being listed below:
Orenburg - later Chkalov - now Orenburg
Samara - later Kuybyshev - now Samara
Simbirsk - now Uljanovsk
Sarepta - now Krasnoarmeisk of the Volgograd District
Waskuntschatskoi - usually noted as Baskunchak (Astrakhan District)
Zarizyn or Tsarizyn - later Stalingrad - now Volgograd.
Note: Spassk, usually interpreted as Eversmann's estate not far from Orenburg, really might
be also a town being flooded by the Volga's water during the erection of hydro-electric power
stations and following increasing waters levels. Before that Spassk had been situated at about
82 km ESE of Kasan on the left bank of the Volga.
Notes on the table and maps
Column 1: Species number
species is deleted from the list
Column 2: Species name
Column 3: Species listed by Eversmann (1844) within the regional limits of that paper
Column 4-10: Administrative units
4 Astrakhan District (centre is Astrakhan)
5 Volgograd District (Volgograd)
6 Saratov District (Saratov)
7 Samara District (Samara)
8 Uljanovsk District (Uljanovsk)
9 Bashkiria (Ufa)
10 Uralsk District (Uralsk)
+ species is present
species not found during period of this study
? species is known from old or doubtful data
o type locality
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Column 11: Flight periods
lll-XI - months
b, m, e - beginning, middle, end of month
1 (2) G - species develops 1 (2) generation(s)
Column 12: Comments and larval foodplants
L larval foodplants, * indicating original local data
TL type locality
E E. Eversmann
On the maps, the filled circles indicate the localities in which the species concerned is to be
found. Hollow circles indicate those localities in which the species concerned have not been
found during the last 15-50 years, possibly having disappeared due to environmental
changes. A plot size we accepted is 30x30 km.
E A V S S U B U
V S 0 A A L A R
E T L R M J S A
R R G A A A H L
N Species S A O T R N K S Flight Comments
M K G 0 A 0 I K period
A H R V V R
N A A S I
N N D K A
1 2 3 4 5 6 7 8 9 10 11 12
Geometridae
Archiearinae
1. Archiearis parthenias mlV-mV Local in light birch- and mixed for
Linnaeus, 1761 in 1 G est and parks. L: Betula pendula*.
2. Archiearis notha IV Rare and local in light deciduous
Hübner, [1803] in 1 G forests. L: Populus.
Alsophilinae
3. Alsophila aescularia elV-bV Local and rare in mixed and decidu
Denis & Schiffermüller, in 1 G ous forests. L: Quercus, Acer.
1775
Larentiinae
4. Lythria purpuraría + + + + + V-VI; Not rare in dry and stepped places.
Linnaeus, 1758 VII-VIII L: Rumex confertus*.
in 2 G
5. Lythria cruentaría + + + + + V; mVI- Common everywhere.
Hufnagel, 1767 mVIII
(= purpurata auct.) in 2 G
6. Cataclysme riguata + mV-VI I; Rare in deciduous forests.
Hübner, [1813] 1796 VIII in 2 G
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1 2 3 4 5 6 7 8 9 10 11 12
7 Phibalapteryx virgata + + + + + + V-VI; Was cited by E. as Aspilates
Hufnagel, 1767 bVIII Lineolata. Not rare in steppes,
in 2 G
8. Scotopteryx coa retaría + + + eV-mVI Rare and local in steppes and
Denis & Schiffermüller, in 1 G stepped biotopes.
1775
9. Scotopteryx mucronata + + + + + + mV-mVIll Common in steppes and dry mead-
Seo poli, 1763 in 1 G ows. L: Cythisus ruthenicus*.
10. Scotopteryx luridata ? Was noted by E. as Aspilates Pal
Hufnagel, 1767 umbaria. In the region ssp. plum
bada Fabricius, 1775 is native.
11. Scotopteryx chenopodiata + + + + + - mV-mVIll Was cited by E. as Larentia
Linnaeus, 1758 in 1 G Mensuraría. Common in forests
and meadows.
12. Scotopteryx moeniata + + + + - Vll-mVIll Was listed by E. as Cidaria
Scopoli, 1763 in 1 G Moeniaria. Not common in
steppes. L: Cythisus ruthenicus*.
13. Scotopteryx burgaria - ? Known only from old collection ma
Eversmann, 1843 terial.
14. Scotopteryx bipunctaria - bVII- Not common in steppes and
Denis & Schiffermüller, mVIII stepped biotopes.
1775 (map 1) in 1 G
15. Costaconvexa + + VII-VIII Rare and local in sandy steppes,
polygrammata in 1 G
Borkhausen, 1794
16. Catarhoe cuculata + + + + + mVI-bVI 11 Was cited by E. as Zerene Sinuata.
Hufnagel, 1767 Rare in old mixed forests.
17 Catarhoe rubidata Denis + + + + + mV-eVI; Was cited by E. as Cidaria Fumata.
& Schiffermüller, 1775 VIII Rare and local in old mixed forests,
in 2 G In the region, ssp. fumata Evers
mann, 1844 is native.
18. Camptogramma mVI-VIII Not rare in meadows, glades, clear-
bilineatum Linnaeus, 1758 in 1 G ings and deciduous forests.
19. Ochyría quadrifasiata mVI-mVIl Not rare but local in mixed forests,
Clerck, 1759 in 1 G
20. Orthonama vittata + eVI-mVIll Was cited by E. as Acidalia Lignata.
Borkhausen, 1794 in ?1 G
21. Orthonama obstipata - mVIII Was listed from Astrakhan Distr. by
Fabricius, 1794 Lvovsky (1971).
22. Xanthorhoe fluctuata bV-eVIII Everywhere common,
Linnaeus, 1758 in 2 G
23. Xanthorhoe montanata + + + + - bVI-mVIl Not rare in forests of all types,
Denis & Schiffermüller, in 1 G
1775
24. Xanthorhoe spadicaeria + + + + - VI; eVI11 Not common in dry clearings in old
Denis & Schiffermüller, in 2 G forests.
1775
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1 2 3 4 5 6 7 8 9 10 11 12
25. Xanthorhoe ferrugata + + + + + eV-bVIll Was cited by E. as Cidaria
Clerck, 1759 in 2 G Ferrugaria. Common in mixed and
deciduous forests and parks.
26. Xanthorhoe biriviata + - V-Vll Was noted by E. as Cidaria
Borkhausen, 1794 in 2 G Pomoeriaria Evm. Rare on humid
(= pomoeharia glades near the water. L: Impatiens
Eversmann, 1844). noli-tangere.
27 Xanthorhoe designate + + + - mVI- Was listed by E. as Cidaria
Hufnagel, 1767 mVIII Propugnaría. Rare in humid decidu
in 1 G ous and coniferous forests.
28. Euphyia scripturata - VII Rare in deciduous forests.
Hübner, 1799 in 1 G
29. Euphyia biangulata + + + - eVI-VII Was listed by E. as Cidaria Picata.
Haworth, 1809 in 1 G Rare in old humid forests.
30. Euphyia unangulata mVI Rare in old mixed forests,
Haworth, 1810 in TG
31. Epirrhoe tristata + + + + V-VI Not rare in mixed forests, stepped
Linnaeus, 1758 in 1 G biotopes and on dry meadows.
32. Epirrhoe hastülata VI Not common in sparse forests,
Hübner, 1790 in 1 G L: Galium, Asperula.
33. Epirrhoe pupillata bVI-mVIll Not common in steppes and
Thunberg, 1788 in 1 G stepped biotopes.
34. Epirrhoe galiata Denis & bVI-eVII In the region ssp. eophanata
Schiffermüller, 1775 Krulikovsky, 1906 is native. Rare in
forest-steppes.
35. Epirrhoe rivata eV-mVI Rare and local in mixed and decidu-
Hübner, [1813] 1796 in 1 G ous forests.
36. Epirrhoe alternata + + + + + mV-bVIl; Was noted by E. as Cidaria
Müller, 1764 (= sociata VIII Alchemillata. Common in clearings
Borkhausen, 1794) in 2 G and glades of mixed forests.
37 Earophlia badiata Denis + + - elV-bV Not common in sparse deciduous
& Schiffermüller, 1775 in 1 G forests on bogs.
38. Anticlea derivata Denis + - elV-mV Not common in sparse deciduous
& Schiffermüller, 1775 in 1 G forests.
(map 2)
39. Mesoleuca albicillata + + + + Vl-eVIII Not rare in forests, parks and or-
Linnaeus, 1758 in 1 G chords. L: Rubus idaeus*.
40. Pelurga comitata + + + + + + mV-IX Was cited by E. as Larentia Cheno-
Linnaeus, 1758 in 2 G podiata. Common everywhere but
more typical for anthropogenic
biotopes. L: Chenopodium albium*
41. Larentia clavaria + - eVIII-IX Was listed by E. as Larentia
Haworth, 1809 (= cer- in 1 G Cervinaria. Local in mixed forests.
vinata Denis & Schif-
FERMÜLLER, 1775)
42. Spargania luctuata Denis VI Rare and very local in coniferous
& Schiffermüller, 1775 in 1 G and mixed forests.
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1 2 3 4 5 6 7 8 9 10 11 12
43. Hydriomena furcata + + + + - VII Was listed by E. asAcidalia Elutata.
Thunberg, 1784 in 1 G Rare and local in humid old forests.
44. Hydriomena impluviata VII Very rare in deciduous forests.
Denis & Schiffermüller, in 1 G
1775 (= coerulata
Fabricius, 1775)
45. Colostygia aptata VII Rare on stoned slopes and forests
Hübner, 1813 in 1 G glades. L: Galium.
46. Colostygia olivata Denis ? Vll-Vlll Was noted from Urals by Spuler
& Schiffermüller, 1775 in 1 G (1910).
47. Colostygia pectinataria + + + - Vl-eVII Rare in old coniferous forests.
Knoch, 1781 (= viridata in 1 G
Fabricius, 1775)
48. Chloroclysta siterata - IX Was noted by E. as Larentia
Hufnagel, 1767 in 1 G Psittacata. Very rare in deciduous
forests.
49. Chloroclysta miata - ? Was listed by E. as Cidaria Miaria.
Linnaeus, 1758 No fresh material at our disposal.
50. Electrophaes corylata + + + + - eV-VI Was cited by E. as Cidaria Ruptata.
Thunberg, 1792 in 1 G Common in sparse forests and for
est glades.
51. Dysstroma truncata Vll-eVIII Was cited by E. as Cidaria Russata.
Hufnagel, 1767 in 1 G Rare in forests of different types.
52. Dysstroma citrata eVIll Very rare in humid deciduous for
Linnaeus, 1758 in 1 G ests.
53. Plemyria rubiginata Denis + ? + Vl-eVII Rare in forest steppes on chalk
& Schiffermüller, 1775 in 1 G hills.
54. Thera variata Denis & - VIII From Saratov Distr. was cited by
Schiffermüller, 1775 in 1 G Kumakov & Korshunov (1979).
55. Thera obeliscata m-eVI Not rare but local in old humid
Hübner, 1787 in 1 G mixed forests.
56. Eustroma reticulatum 22.VII. A single specimen is known, col
Denis & Schiffermüller, 1973 lected from humid meadows near
1775 Chumora. L: Impatiens
noli-tangere*.
57. Eulithis prunata ? - ? The species is known only from old
Linnaeus, 1758 material. L: Ribes*.
58. Eulithis testata - eVI Was cited by E. as Cidaria
Linnaeus, 1761 in 1 G Achatinata. Rare in deciduous for
ests. L: Salix, Populus.
59. Eulithis populata VI; VIII Common in light deciduous and
Linnaeus, 1758 in 2 G mixed forests. L: Populus, Rubus.
60. Eulithis mellinata + + + + - Vl-mVIl; Was listed by E. as Cidaria
Fabricius, 1787 VIII Marmorata. Rare in parks and or
in 2 G chards. L: Ribes nigrum*.
61. Eulithis pyropata + + - VI-VII Local in forests of different types.
Hübner, 1809 in 1 G
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62, Eulithis pyraliata Denis & + + + + + mVI-mVIl Rare in deciduous and mixed for-
Schiffermüller, 1775 in 1 G ests.
63. Ecliptopera capitata V; VII-VIII Rare and local in humid forests and
Herrich-Schaffer, 1839 in 2 G on humid meadows. L: Impatiens
noli-tangere*.
64. Ecliptopera silaceata - mVI Rare and local in old mixed forests,
Denis & Schiffermüller, in 1 G
1775
65. Cosmorhoe ocellata + - eVI-mVIl Rare in sparse forests,
Linnaeus, 1758 in 1 G
66. Lampropteryx suffumata + + + - elV-mV Rare in light deciduous forests.
Denis & Schiffermüller, in 1 G
1775
67. Operophtera brumata + + + + bX-XI Not rare in forests, parks and or
Linnaeus, 1758 ín 1 G chords. L: Malus domestica*,
Betula péndula*.
68. Epirrita dilutata Denis & ? ? From Samara noted by
Schiffermüller, 1775 Stsherbinovsky(1919).
69. Epirrita autumnaria mIX-bX Rare in forests and parks,
Borkhausen, 1794 in 1 G
70. Minoa murinata + + + + mV-eVI; Was cited by E. as Minoa
Scopoli, 1763 mVII Enphorbiata [sic!]. Rare and local
in 2 G on dry warm clearings.
71. Asthena albulata mV-mVIl Very common in humid deciduous
Hufnagel, 1767 in 1 G forests, sitting on the leaves of
Corylus avellana.
72. Euchoeca nebulata + ? m-eVI Was cited by E. as Fidonia
Scopoli, 1763 in 1 G Hepararia. Rare and local in old
humid mixed forests.
73. Venusia cámbrica 18.VII Very rare and local in mountain
Curtis, 1839 biotopes.
74. Venusia blomeri + - eVI-bVIl Was listed by E. as Cidaria
Curtis, 1839 in 1 G Pulchraria. Not common in edges
of deciduous humid forests.
L: Ulmus glabra*.
75. Hydrelia sylvata Denis & + - VII Rare in deciduous and mixed for
Schiffermüller, 1775 in 1 G ests. L: Alnus.
(= testaceata Donovan,
1810)
76. Hydrelia flammeolaria + + + + bVI-mVIl Was noted by E. as Acidalia
Hufnagel, 1767 in 1 G Luteata. Rare and local in humid
and shady forests.
77. Triphosa dubitata ? VII Rare in forest steppes,
Linnaeus, 1758 in 1 G
78. Philereme vetulata Denis + + + +.+ mV-mVIl Was cited by E. as Acidalia
«.Schiffermüller, 1775 in 1 G Affectata Evm. Not rare in mixed
and deciduous forests. L: Rhamnus
cathartica*.
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2 3 4 5 6 7 8 9 10 11 12
79. Philereme transversata + + + + + + mVI-mVIl Was noted by E. as Acidalia
Hufnagel, 1767 in 1 G Rhamnata. Rare and local in
steppe biotopes.
80. Rheumaptera hastata + + - eVI Rare in deciduous forests,
Linnaeus, 1758 in 1 G
81. Hydria undulata rnVI-mVIl Rare in mixed and deciduous for-
Linnaeus, 1758 in 1 G ests.
82. Baptria tibiale + + VI-VII Was cited by E. as Psodos Tibialata.
Esper, 1791 in 1 G From Samara Region listed by
Novoderezhkin (1983).
83. Horisme aquata ? m-eVI Local on warm glades and near
Hübner, 1813 in 1 G bushes, typical for forest-steppe.
L: Pulsatilla, Clematis, Anemone.
84. Horisme vitalbata Denis - mVI-bVIl Rare and local in chalk forest-
& Schiffermüller, 1775 in 1 G steppe. L: Pulsatilla*.
85. Horisme corticata - VIII Rare in dry sandy steppes.
Treitschke, 1835 in 1 G
86. Horisme tersata Denis & - mVI-VIII Local in steppes.
Schiffermüller, 1775 in 1 G
87. Horisme aemulata 11.VI. The only specimen was found in a
Hübner, 1813 1997 humid light mixed forest at a bog.
88. Horisme calligraphata + - VII Rare in forest-steppes.
Herrich-Schäffer, 1839 in 1 G
89. Anticollix sparsata + - eVI Rare and local in old humid mixed
Treitschke, 1828 in 1 G forests.
90. Mesotype parallelolineata - - VI Very rare in deciduous forests.
Retzius, 1783 in 1 G
91. Perizoma taeniatum + - ? No material at our disposal.
Stephens, 1831
92. Perizoma alchemillatum + + + + eVI-eVII Was noted by E. as Acidalia
Linnaeus, 1758 in 1 G Rivulata. Common in forests,
glades and meadows.
93. Perizoma hydratum + - eV-VI Local in forests,
Treitschke, 1828 in 1 G
94. Perizoma lugdunarium - 27.VII. Known only from Uljanovsk town.
Herrich-Schaffer, 1856 1987
95. Perizoma bifaciatum + eVII-VIII Rare and local in pine or fir forests,
Haworth, 1809 (= uni- in 1 G
fasciata Haworth, 1809)
96. Perizoma blandiatum + + - m-elV Was noted by E. as Zerene Albidata
Denis & Schiffermuller, in 1 G Evm. Rare in old humid forests.
1775
97. Perizoma albulatum Denis - bVI-eVII Not rare in meadows and forest
& Schiffermuller, 1775 in 1 G glades.
98. Perizoma flavofasciatum + + + + - bVI-bVIll Was listed by E. as Acidalia
Thunberg, 1792 in 1 G Decolorata. Local in humid mixed
forests near the water.
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1 2 3 4 5 6 7 8 9 10 11 12
99. Perízoma sagittatum + ? No material at our disposal.
Fabricius, 1792
100. Perízoma didymatum + - Vil Was cited by E. as Acidalia
Linnaeus, 1758 ¡n 1 G Scabraria. Rare in forests.
101. Gymnoscelis rufifasáata + elV-VII Common in forest steppes.
Haworth, 1809 ¡n 1 G
102. Chloroclystís v-ata + + + + + - mVI-bVIl Was noted by E. as Larentia
Haworth, 1809 (= coro- ¡n 1 G Coronata. Rare in parks and sparse
nata Hübner, 1813) forests.
103. Rhinoprora rectangulata - mVI-VII Rare in parks and orchards.
Linnaeus, 1758 ¡n 1 G L: Malus* Pyrus*.
104. Rhinoprora chloerata m-eVI Rare in parks.
Mabille, 1870 ¡n 1 G
105. Rhinoprora debiliata b-mVI Rare and local in sparse light birch-
Hübner, 1817 ¡n 1 G forests and on sphagnum bogs.
106. Eupithecia abietaría ? ? The nearest territory this species is
Goeze, 1781 known from is Chuvashia
(Lastukhin, 1990a); it should be
found in the northern part of the
region under study.
107. Eupithecia analoga From Uljanovsk Distr. cited by E. as
Djakonov, 1926 Larentia Residuata. The species is
(= bilunulata auct., also known from the nearby terri
= europaea Lempke, 1969) tory of Chuvashia (Lastukhin,
1990a) as Eupithecia bilunulata
Zetterstedt.
108. Eupithecia linaríata Denis + + + + + - mVI-eVIll Common everywhere in dry places,
& Schiffermüller, 1775 in 2 G L: Linaria vulgaris*, L. genistifolia*.
109. Eupithecia pyreneata + ? Was cited from S. Ural by Mironov
Mabille, 1871 (1990).
110. Eupithecia venosata + VII Not rare in forest steppes.
Fabricius, 1787 in 1 G
111. Eupithecia alliaria bVII Very rare and local in
Staudinger, 1870 in 1 G grass-steppes. L: Allium.
112. Eupithecia minusculata + eVII-VIII Rare in flooded forests.
Alpheraky, 1882 in 1 G
113. Eupithecia pusillata Denis + + + V Was cited by E. as Larentia
& Schiffermüller, 1775 (= in 1 G Sobrinata. From Ural noted by
sobrinata Hübner, 1817) Mironov (1990).
114. Eupithecia virgaureata + + - Vl-mVIll Rare in deciduous forests.
Doubleday, 1861 in 2 G
115. Eupithecia barteli Dietze, - VIII Very rare in flooded forests of
1908 in 1 G steppe zone.
116. Eupithecia lariciata ? VI Supposedely was cited by E. as
Freyer, 1842 in 1 G Larentia Residuata. Common but
local in parks with foodplants;
probably was introduced with Larix.
L: Larix sibirica*
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1 2 3 4 5 6 7 8 9 10 11 12
117 Eupithecia tantWaria + - ? No material at our disposal.
Boisduval, 1840
118. Eupithecia lanceata The species is known from the
Hübner, 1825 nearby territory of Chuvashia
(Lastukhin, 1990a); it should be
found in the northern part of the
region under study.
119. Eupithecia sinuosaria + + + + + mVI-eVII Local and rare, more typical for
Eversmann, 1848 in 1 G stepped biotopes.
120. Eupithecia selinata VII Very rare in forest-steppes.
Herrich-Schäffer, 1861 in 1 G
121. Eupithecia egenaria + + + + + eV-bVIl Not common in deciduous forests
Herrich-Schäffer, 1848 in 1 G and parks. L: Tilia.
122. Eupithecia pimpinellata + + - m-eVII Rare in old humid deciduous for
Hübner, 1813 in 1 G ests.
123. Eupithecia denotata + + + VI-VII Local and rare on the glades and
Hübner, 1813 in 1 G clearings of mixed forests.
124. Eupithecia plumbeolata VI Not common in old humid decidu
Haworth, 1809 in 1 G ous forests.
125. Eupithecia innotata + + + + + bV-mVIl; Not rare in sparse forests and dry
Hufnagel, 1767 VIII-bIX clearings. 1: Artemisia*
in 2 G (lerchiana*).
126. Eupithecia nanata b-mV Rare and local in steppes and
Hübner, 1813 in 1 G stepped biotopes.
127. Eupithecia ochridata + VI-VII I Local in steppes and stepped
Pinker & Schütze, 1968 in 1 G biotopes.
128. Eupithecia simpliciata + + + + + mVI- Common everywhere.
Haworth, 1809 (= sub- mVIII
notata Hübner, 1813)
129. Eupithecia centaureata + + eVI-bVIll Everywhere not rare,
Denis & Schiffermüller, in 2 G
1775 (= oblongata
Thunberg, 1784)
130. Eupithecia extraversaria + + + VII Local in steppes, mainly the grassy
Herrich-Schäffer, 1852 in 1 G ones.
131. Eupithecia gueneata + + + eVI-bVIl Extremely rare and local in dry
Milliere, 1862 in 1 G steppes and deciduous forests on
the stepped places.
132. Eupithecia gratiosata Was cited from S. Ural by Mironov
Herrich-Schäffer, 1861 (1991).
133. Eupithecia vulgata + mV-VIII Common everywhere but more typ-
Haworth, 1809 in 2 G ical for forest steppe.
134. Eupithecia assimilata + + + - eV-bVI Rare and local in parks and forests,
Doubleday, 1856 in 1 G
135. Eupithecia satyrata + + + + eV-eVI Was noted by E. as Larentia
Hübner, 1813 in 1 G Asteraria. Rare and local in old hu
mid forests.
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