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Essentials of Physical Anthropology PDF

454 Pages·2013·141.99 MB·English
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Major Fossil Hominin Sites GREENLAND 75°N (KALAALLIT NUNAAT) (DEN.) Arctic ALASKA 150°W 120°W 150°9W0°W 120°W60°W 90°W30°W 60°W0° 30°W30°E 0° 60°E 30°E90°E 60°1E20°E 90°1E50°E 120°E 150°E (U.S.) 60°N ICELAND 75°N 75°N (KALGARALELEINTL NAUNNDAAT) (KALGARALELEINTL NAUNNDAAT) ARCTIC OCEANARCTIC OCEAN 75°N 75°N (DEN.) (DEN.) CANADA IRELAND U. K. 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EAST(CERH IILSEL)ANDISLPAINTCDASI R(UN.K.) EAST(CERH IILSEL)CAHNIDLE ARGENTINA URUGUAY CHILE ARGENTINA URUGUAY (dSettaetrums itnoe db)e FTSlAaoOFuRrUniITsCHgAb ad LEBSoO(dSSrTetWdHtaetOAreumZsr Ii tLnCoAe dbNa)eDve SAOFRUITCHA LESSOWTAHZOILAND Lake Mungo 30°S 30°S Klasies River Mouth FALKLAND ISLANDS (U.K.) Kow Swamp 0 1000 miles SOUTH GEORGIA ISLAND 1. SLOVENIA 1. SLOVENIA 2. CROATIA 2. CROATIA NEW ZEALAND NEW ZEALAND 0 1000 kilometers 3. BOSNIA AND HERZEGOVINA 3. BOSNIA AND HERZEGOVINA 45°S 45°S 45°S 45°S 60°S 4. ALBANIA 4. ALBANIA 5. MACEDONIA 5. MACEDONIA Antarctic Circle 0 1000 mile0s 1000 miles FALKLAND ISLANDS (U.K.) FALKLAND ISLANDS (U.K.) 6. SERBIA AND MONTENEGRO 6. SERBIA AND MONTENEGRO SOUTH GEORGIA ISLAND SOUTH GEORGIA ISLAND 0 1000 kilometers 0 1000 kilometers 75°S 60°S 60°S 60°S 60°S Antarctic Circle Antarctic Circle 75°S 75°S 75°S 75°S 150°W 120°W 150°9W0°W 120°W60°W 90°W30°W 60°W0° 30°W30°E 0° 60°E 30°E90°E 60°1E20°E 90°1E50°E 120°E 150°E 1000 Kilometers 1000 Kilometers 1000 Kilometers Parallel scale at Parallel scale at Parallel scale at Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does n0ot m anteriaolly rafftehct th e ov e0rall leearnaing sexpterience. 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A(NAHrearmtois, ,D Bikoiukrai), INDIA SRI BLAANNGKLAADESHM(BYUARNMMAA)RLTHiuALIjALiOAaSNKnDAgMPUVCIHEMTENAMaAAbMCaHAOUNG KONTGAIWANPHILIPPINEPS ANMIOCSARLRTAIHNAIEDNRSAFN IC OTroCpicE oAfM RACERNPSaUHBnALLIcLC eIOSFrL ATN1HDE5S°N Equator COLOMBIA SURFINRAENMCEH GUIANA (FR.) LIBERIA EGTQOUUGINAOETA. CAMEROON REPUBLRICWANDA UGANDA KOEamansaotp aonid West Turkana MALDIVES MALAYSBIRUANEINiah Cave FEODFE RMAICTERDO NSETSAITAES Equator 0° Tugen Hills KENYA SINGAPORE BORNEO 0° WSEASMTEORAN OCEAN GA(LEACPUAAGDOOSR )IS. ECUADOR PERU BRAZIL S&A POCR( AATINnBOgCIMNoIPEDla EA)GABOCNONGAONGOLODAFER TMEHPOEUC CBROLANITCGICO BTUARNUZNADNIIAOlduvaiC/LOMaeORtoOlSi ISLSAENYDCSHELLES INDIAN SangiranTriniNlgaFnldoorensgINDONESIA PNAEPWU AGUINEA SOLOMON ISLANDS TUVALU 15°S AMERICAN SAMOA ZAMBIA 15°S TONGA SOCIET(YFR IS.)LANDS T(AFHRI.T)I TropFiRcEN oCHf PCOLaYNpESrIiAc (FoRr.)n BOLIVIAPARAGUAY NAMIBIA BOTKSWaAbZNIwMABeABWEMOZAMMBIAQLUAEWI MADAGASCAMRAURITIUS OCEAN NEW CALEDONIA VANUATU FIJI WALVIS BAY Sterkfontein/Swartkrans/Drimolen/Malapa AUSTRALIA Tropic of Capricorn 30°S ISLPAINTCDASI R(UN.K.) EAST(CERH IILSEL)AND CHILE ARGENTINA URUGUAY (dSettaetrums itnoe db)e FTSlAaoOFuRrUniITsCHgAb ad LEBSoOSrTWdHOAeZrI LCANaDve Lake Mungo 30°S Klasies River Mouth Kow Swamp 1. SLOVENIA 2. CROATIA NEW ZEALAND 3. BOSNIA AND HERZEGOVINA 45°S 45°S 4. ALBANIA 5. MACEDONIA 0 1000 miles FALKLAND ISLANDS (U.K.) 6. SERBIA AND MONTENEGRO SOUTH GEORGIA ISLAND 0 1000 kilometers 60°S 60°S Antarctic Circle 75°S 75°S 150°W 120°W 90°W 60°W 30°W 0° 30°E 60°E 90°E 120°E 150°E 1000 Kilometers 1000 Kilometers 1000 Kilometers Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not mPateriaallyr afafectl tlhee ovle rasll lecarnaingl eexpe rieancet. Cengage Learning reserves the right to rPemoave arddaitiolnlael colnt enst act anay tilmee if suabsetquent rights restrictions require it. Parallel scale at 0 north 0 east 30 north 0 east 60 north 0 east front end map insert.indd 1 1/11/12 12:20 PM ESSENTIALS OF Physical Anthropology NINTH EDITION Robert Jurmain Professor Emeritus, San Jose State University Lynn Kilgore University of Colorado, Boulder Wenda Trevathan Professor Emerita, New Mexico State University Australia • Brazil • Japan • Korea • Mexico • Singapore • Spain • United Kingdom • United States Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 1 1/13/12 3:28 PM This is an electronic version of the print textbook. Due to electronic rights restrictions, some third party content may be suppressed. Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. The publisher reserves the right to remove content from this title at any time if subsequent rights restrictions require it. For valuable information on pricing, previous editions, changes to current editions, and alternate formats, please visit www.cengage.com/highered to search by ISBN#, author, title, or keyword for materials in your areas of interest. Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. Essentials of Physical Anthropology, © 2013, 2011 Wadsworth, Cengage Learning Ninth Edition ALL RIGHTS RESERVED. No part of this work covered by the copyright Robert Jurmain, Lynn Kilgore, herein may be reproduced, transmitted, stored, or used in any form and Wenda Trevathan or by any means graphic, electronic, or mechanical, including but Acquiring Sponsoring Editor: Erin Mitchell not limited to photocopying, recording, scanning, digitizing, taping, Web distribution, information networks, or information storage and Developmental Editor: Lin Gaylord retrieval systems, except as permitted under Section 107 or 108 of the Assistant Editor: Mallory Ortberg 1976 United States Copyright Act, without the prior written permission of the publisher. Media Editor: John Chell Marketing Program Manager: Janay Pryor For product information and technology Content Project Manager: Cheri Palmer assistance, contact us at Art Director: Caryl Gorska Cengage Learning Customer & Sales Support, 1-800-354-9706. Manufacturing Planner: Judy Inouye For permission to use material from this text or product, Rights Acquisitions Specialist: Dean Dauphinais submit all requests online at www.cengage.com/permissions. Design, Production Services, and Composition: Further permissions questions can be e‑mailed to Hespenheide Design [email protected]. Photo Researcher: Patti Zeman, Hespenheide Design Text Researcher: Ashley Liening Library of Congress Control Number: 2011945171 Copy Editor: Cheryl Smith Student Edition: ISBN‑13: 978‑1‑111‑83718‑1 Illustrator: Hespenheide Design ISBN‑10: 1‑111‑83718‑x Cover Designer: Hespenheide Design Loose‑leaf Edition: Cover Image: Tufted Capuchin Sanctuary Icamaperou, ISBN‑13: 978‑1‑111‑83816‑4 Upper Amazon, Peru: Cyril Ruoso/Photolibrary ISBN‑10: 1‑111‑83816‑x Wadsworth 20 Davis Drive Belmont, CA 94002‑3098 USA Cengage Learning is a leading provider of customized learning solutions with office locations around the globe, including Singapore, the United Kingdom, Australia, Mexico, Brazil, and Japan. Locate your local office at www.cengage.com/global. Cengage Learning products are represented in Canada by Nelson Education, Ltd. To learn more about Wadsworth, visit www.cengage.com/wadsworth Purchase any of our products at your local college store or at our preferred online store www.CengageBrain.com. Printed in the United States of America 1 2 3 4 5 6 7 16 15 14 13 12 Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 2 1/16/12 4:20 PM Brief Contents Chapter 1 Introduction to Physical Anthropology 2 Chapter 2 The Development of Evolutionary Theory 26 Chapter 3 The Biological Basis of Life 50 Chapter 4 Heredity and Evolution 72 Chapter 5 Processes of Macroevolution 96 Chapter 6 An Overview of the Primates 120 Chapter 7 Primate Behavior 154 Chapter 8 Primate and Hominin Origins 186 Chapter 9 The First Dispersal of the Genus Homo: Homo erectus and Contemporaries 230 Chapter 10 Premodern Humans 252 Chapter 11 The Origin and Dispersal of Modern Humans 282 Chapter 12 Human Variation and Adaptation 310 Chapter 13 Legacies of Human Evolutionary History: Effects on the Individual 342 Chapter 14 The Human Disconnection 366 iii Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 3 1/16/12 4:21 PM Contents Preface xi Acknowledgments xiv Supplements xv 1 ChapTeR Introduction to Physical Anthropology 2 Introduction 3 The Human Connection 6 Biocultural Evolution 6 What Is Anthropology? 10 A AS Cultural Anthropology 11 N Archaeology 11 Linguistic Anthropology 12 Physical Anthropology 12 Applied Anthropology 18 2 Physical Anthropology and the Scientific Method 20 ChapTeR The Anthropological Perspective 21 The Development of Why It Matters 22 Evolutionary Theory 26 Summary of Main Topics 23 Critical Thinking Questions 23 A Brief History of Evolutionary Thought 28 Photo Essay Making a Difference: Forensic The Scientific Revolution 29 Anthropologists in the Contemporary Precursors to the Theory of Evolution 31 World 24–25 The Discovery of Natural Selection 34 In Darwin’s Shadow 39 Natural Selection 40 Natural Selection in Action 41 Quick Review The Mechanism of Natural Selection 43 Constraints on Nineteenth-Century Evolutionary Theory 44 Opposition to Evolution Today 45 A Brief History of Opposition to Evolution in the United States 46 Why It Matters 47 Summary of Main Topics 48 Critical Thinking Questions 49 Peter Jones iv Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 4 1/13/12 3:29 PM v Contents v 3 ChapTeR The Biological Basis of Life   50 Cells 52 DNA Structure 53 DNA Replication 54 Protein Synthesis 54 CWRQueeghilclua kDtl a RIitvseo iavrs iiyGeo wGne n enCe?6eo 1sd i5n85g9 and Noncoding DNA 60 © J. Craig Venter Institute Chromosomes 61 Mitosis 62 4 Meiosis 63 ChapTeR New Frontiers 67 Heredity and Evolution 72 Why It Matters 70 The Genetic Principles Discovered by Mendel 74 Summary of Main Topics 70 Segregation 74 Critical Thinking Questions 71 Dominance and Recessiveness 75 Independent Assortment 78 Mendelian Inheritance in Humans 78 Misconceptions about Dominance and T A Recessiveness 80 C G Polygenic Inheritance 81 T A Quick Review Mendelian vs. Polygenic Traits 83 Genetic and Environmental Factors 83 T A G Mitochondrial Inheritance 84 C Modern Evolutionary Theory 84 A T The Modern Synthesis 85 A A A C A Current Definition of Evolution 85 U C Factors That Produce and Redistribute Variation 86 T U G U Mutation 86 T G T Gene Flow 86 G mRNA Genetic Drift and Founder Effect 87 T C T Recombination 89 G Natural Selection Is Directional G G C A A C and Acts on Variation 90 Review of Genetics and Evolutionary Factors 93 A Why It Matters 94 T C G Summary of Main Topics 94 DNA template strand T A Critical Thinking Questions 95 Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 5 1/13/12 3:29 PM vi Contents 5 ChapTeR Processes of Macroevolution 96 How We Connect: Discovering the Human Place in the Organic World 98 Principles of Classification 98 Constructing Classifications and Interpreting Evolutionary Relationships 101 Comparing Evolutionary Systematics with Cladistics 101 An Example of Cladistic Analysis: The Evolution- ary History of Cars and Trucks 102 Using Cladistics to Interpret Real Organisms 103 Definition of Species 105 Interpreting Species and Other Groups in the Fossil Record 106 Recognition of Fossil Species 107 Recognition of Fossil Genera 108 What Are Fossils and How Do They Form? 109 Vertebrate Evolutionary History: A Brief Summary 111 Mammalian Evolution 113 The Emergence of Major Mammalian Groups 115 Processes of Macroevolution 115 Adaptive Radiation 115 Generalized and Specialized Characteristics 116 Working Together: Microevolution and Macroevoluation 117 SWuhmy mIt Marayt toefr Ms ai1n1 8Topics 118 Nelson Ting © Critical Thinking Questions 119 6 ChapTeR An Overview Primate Classification 131 A Survey of the Living Primates 134 of the Primates  120 Lemurs and Lorises 134 Introduction 121 Tarsiers 136 Primate Characteristics 122 Anthropoids: Monkeys, Apes, and Humans 137 Primate Adaptations 126 Hominoids: Apes and Humans 141 Evolutionary Factors 126 Endangered Primates 148 Geographical Distribution and Habitats 127 Why It Matters 152 Diet and Teeth 127 Summary of Main Topics 153 Locomotion 130 Critical Thinking Questions 153 Copyright 2012 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 37181_00_i-001_FM.indd 6 1/13/12 3:29 PM

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