E NCYCLOPEDIA OF FOREST SCIENCES E NCYCLOPEDIA OF FOREST SCIENCES Editor-in-Chief JEFFERY BURLEY Editors JULIAN EVANS JOHN A YOUNGQUIST Amsterdam Boston Heidelberg London New York Oxford Paris San Diego San Francisco Singapore Sydney Tokyo Elsevier Ltd., The Boulevard, Langford Lane, Kidlington, Oxford, OX5 1GB, UK Elsevier Inc., 525 B Street, Suite 1900, San Diego, CA 92101-4495, USA & 2004 Elsevier Ltd. The following articles are US Government works in the public domain and not subject to copyright: EXPERIMENTAL METHODS AND ANALYSIS/Biometric Research; Statistical Methods (Mathematics and Computers). INVENTORY/Modeling. PULPING/Fiber Resources; New Technology in Pulping and Bleaching. WOOD FORMATION AND PROPERTIES/Wood Quality. ENTOMOLOGY/Defoliators, Crown Copyright 2004 HEALTH AND PROTECTION/Integrated Pest Management Practices, Crown Copyright 2004 MENSURATION/Growth and Yield, Crown Copyright 2004 PLANTATIONSILVICULTURE/Multiple-useSilvicultureinTemperatePlantationForestry,CrownCopyright2004 PLANTATION SILVICULTURE/Tending, Crown Copyright 2004 SILVICULTURE/Coppice Silviculture Practiced in Temperate Regions, Crown Copyright 2004 SOCIALANDCOLLABORATIVEFORESTRY/CanadianModelForestExperience,CanadianCrownCopyright 2004 AFFORESTATION/SpeciesChoice,&CommonwealthScientificandInternationalResearchOrganization(CSIRO) TREE PHYSIOLOGY/Nutritional Physiology of Trees, & Commonwealth Scientific and International Research Organization (CSIRO) TREE PHYSIOLOGY/Physiology and Silviculture, & Commonwealth Scientific and International Research Organization (CSIRO) ThearticleSUSTAINABLEFORESTMANAGEMENT/CausesofDeforestationandForestFragmentationisadapted withpermission of Academic Pressfrom Ghazoul JandEvansJ (2001) Deforestationandlandclearing. In: Levin SA (Ed.) Encyclopedia ofBiodiversityISBN: 0-12-226865-2 SOLIDWOODPRODUCTS/Construction; Logs, Poles, Piles,Sleepers (Crossties) andGluedStructuralMembers areadaptedwithpermissionfromtheForestProductsLaboratory(1999).WoodHandbook–Woodasanengineering material. GeneralTechnicalReport FPL-GTR-113. Madison, WI: U.S. Department of Agriculture,ForestService TROPICAL ECOSYSTEMS/MangrovesisadaptedwithpermissionofElsevierLtd. from Spalding MD (2001) Mangroves.In:SteeleJH,TurekianKKandThorpeSA(Eds).EncyclopediaofOceanSciences.ISBN:0-12-227430-X PLANTATION SILVICULTURE/Forest Plantations is adapted with permission of Eolss Publishers from Evans (2003) Forest Plantations, Encyclopedia of Life Support Systems. http://www.eolss.net PACKAGING, RECYCLING AND PRINTING/Paper Recycling Science and Technology is adapted with permission of Elsevier Ltd. from Doshi MR and Dyer JM (2001) Paper recycling and recycled materials. In: BuschowKHJ,CahnRW,FlemmingsMC,IlschnerB,KramerEJandMahajanS(Eds)EncyclopediaofMaterials Science and Technology. ISBN: 0-08-043152-6 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronicormechanical,includingphotocopy,recording,oranyinformationstorageandretrievalsystem,without permission in writing from the publishers. PermissionsmaybesoughtdirectlyfromElsevier’sRightsDepartmentinOxford,UK:phone(þ44)1865843830, fax(þ44)1865353333,e-mailpermissions@elsevier.com.Requestsmayalsobecompletedon-lineviatheElsevier homepage (http://www.elsevier.com/locate/permissions). First edition 2004 Library of Congress Control Number: 2003113256 A catalogue record for this book is available from the British Library ISBN: 0-12-145160-7 (set) This book is printed on acid-free paper Printed in Spain EDITORIAL ADVISORY BOARD EDITOR-IN-CHIEF Jeffery Burley University ofOxford Oxford UK EDITORS Julian Evans Imperial College London Ascot, UK John AYoungquist Forest Products Consultant Verona,WI, USA SaidAbuBakr JabouryGhazoul Western MichiganUniversity Imperial College London Kalamazoo,MI, USA Ascot, UK LABruijnzeel Stephen Harris VrijeUniversiteit University ofOxford HV Amsterdam,TheNetherlands Oxford, UK RowlandBurdon John LInnes NewZealandForest ResearchInstitute University ofBritish Columbia Rotorua,NewZealand Vancouver,BC,Canada JamesBarrackCarle Madhav Karki Forest ResourcesDevelopmentService Medical and AromaticPlants Programin Asia(MAPPA) Rome,Italy New Delhi,India Dennis Dykstra Simon R Leather BlueOxForestry Imperial College London Portland,OR, USA Ascot, UK vi EDITORIALADVISORYBOARD HGydeLund Fergus LSinclair Forest InformationServices University ofWales Bangor Gainesville,VA,USA Gwynedd, UK Lawrence AMorris Universityof Georgia Harold Steen Athens,GA,USA NC, USA JohnParrotta USDA ForestService Research K FreerkWiersum PR,USA Wageningen University Wageningen,The Netherlands DavidRook GallowHill, UK Michael JWingfield PeterSSavill University ofPretoria Universityof Oxford Pretoria, Republic of SouthAfrica Oxford,UK Stephen RJSheppard Robert Youngs TheUniversity ofBritish Columbia Virginia PolytechnicInstituteand StateUniversity Vancouver,BC,Canada Blacksburg, VA,USA FOREWORD T his is a most timely publication because of the vast amount of new information on forest sciences that hasbeenproducedoverthelastfewyearsandbecauseofthegrowingrealizationofthevitalimportance offoreststotheworld.Fortunatelyforourfuture,theworldisbeginningtorealizethatforestsarebothvital for our survival and that they offer many benefits. These benefits vary from the more obvious ones such as timber and fibers for paper pulp, to the environmental aspects such as the sequestration of carbon, the protection ofwatershedsandthe prevention offlooding inmany areas. Also there is agrowingemphasis on the production of non-timber forest products because of their role in sustainable management of forests. Sinceforestsaresocrucialtoourfuturetheirsustainablemanagementisessentialandthisrequiresagreat amount of expertise and information. As I look at the list of the authors and the advisory board of this Encyclopedia, it reads like a who’s who of forest science. These experts have put together a collection of informationandup-to-datecontributionsthatwillbeaninvaluableresourceforanyoneinvolvedwithforests in any way. I am sure that students at all levels, their teachers and lecturers, professional researchers, policy makers, and even the interested layman or amateur forester will find these volumes of great use. AsIlookthroughthecoverageIfinditmostcomprehensiveandcontemporary.Itincludessuchimportant modern topics as the molecular biology of forest trees, the role of forests in the carbon cycle, computer modeling and the use of recently developed methods such as geographic information systems. More traditionalaspectssuchasforestbiologyandecology,theprocessingofforestresourcesintoawiderangeof products,forestrymanagementandpracticeandtheeconomicandsocialaspectsofforestryarebroughtup- to-datehere.Notonlyarethecontributionsthemselvesuseful,buttheyalsodirecttheseriousinvestigatorto morein-depthoradvancedmaterialoneachtopic.Itisalsomostusefulthatthisfineworkwillbeavailablein an electronic version that will facilitate cross-referencing to related topics and references. I know that I will find these volumes most useful and frequently used and I am sure that they will be a standard reference work on forest sciences for at least the next decade. How timely at a period of human history when there is a desperate need to stop deforestation, re-forest many destroyed areas and develop better methods for sustainable use of forests and to conserve the many species of plants and animals which they hold. Prof. Sir Ghillean Prance FRS School of Plant Sciences, University of Reading, Reading, UK INTRODUCTION A tthestartoftheThirdMillenniumthelevelsofpublicandpoliticalattentiontoforests,theirbenefitsto mankind, and their management are at their highest. National and international institutions, governmental and non-governmental organizations, all forms of media and representatives of civil society are searching for socially equitable methods of managing forests toobtainall theirmultiple benefits. Underlying this search is the need for precise and relevant information about the forests, their uses and management together with the political and social institutions that can best effect sustainable management. Our audience for this reference work includes libraries, governmental and non-governmental organiza- tions, universities and individuals involved in research on forests, forest products and services, and relevant topics,local,nationalandinternationaldecision-makingauthoritiesandadministrations,forestland-owners and other forest-dependent individuals. The ranges of biophysical and socio-economic aspects of forests, forestry, forest products and forest services are extremely large; correspondingly, past and current research cover large numbers of scientific disciplines and policy issues. Systematic research has been undertaken for over a century in some forest sciencessuchassilvicultureandforestmanagement;inothertopicsnewlyemergingtechniques,suchasthose of molecular genetics, are being developed to aid understanding of physiological and environmental characteristicsoftrees andforests or toassist selectivebreedingof trees forplantations. AnEncyclopedia of ForestSciencethereforehastoencompassabroadspectrumofpureandappliedsciences,ancientandmodern technologies, and old and recent knowledge. In this Encyclopedia we have obtained outstanding contributions of some 200 specialists covering 250 topics that have wide implications for forest conservation, management and use worldwide. Of course, it is notpossibletocovereverypossiblesubjectofrelevancetoforestsbuttheonesselectedaregenerallyofglobal interest;andeveniftheyareoflocal,nationalorregionalcharacter,theyareimportanttoallthoseconcerned with forest management, research, education, training, policy-making or public information. Becauseofthegreatbreadthofexpectedreadershipwehaveaskedawiderangeofcontributingexpertsto produceuptoapproximately4000wordssummarizingcurrentviewsoftheirtopic.Thecontributionsarenot written in the traditional form of a scientific journal article with detailed bibliographic references for all major statements. Rather each is a continuous, highly readable description based on an author’s personal view of the state of knowledge in her/his area of expertise. Selected major references are given at the end of eachcontributiontofacilitateandencouragefurtherreadingonthesubject.Whereverpossiblephotographs, other graphical illustrations and tables are used to make the material more concise and visualized. Cross- referencingbetweencontributionsandtheprovisionofdummyentriesinthetableofcontentsfacilitateafull coverage of material relevant to each topic. Some contributions are short because it proved difficult to identify an author with the appropriate experience and willingness to write full articles. However, these may be enlarged in future editions of the Encyclopedia and in the web-based version of it. The availability of modern information technology facilitatesnotonlythepreparationofsuchaworkbutalsothemaintenanceofitstimeliness,thespreadingof its availability and the ease of searching and downloading selected material. x INTRODUCTION As Editors we thank the authors for their contributions, the editorial advisors for their specialist support, and the staff of Elsevier for their prompt and effective actions that have allowed the four volumes of the Encyclopedia tobepublishedwithin twoyearsofthe initialcommissioningofthis referencework.Wehope thattheEncyclopediawillprovetobeavaluabletoolandsourceofinformationformanyyearstocome.In particularwehopeitwillencourageagrowingpublicandadedicatedprofessiontounderstandthefactsand institutions necessary for wise management, use and conservation of the world’s forest resources for the equitable benefit of all mankind. Jeffery Burley Julian Evans John A. Youngquist GUIDE TO USE OF THE ENCYCLOPEDIA Structure of the Encyclopedia The material in the Encyclopedia is arranged as a series of entries in alphabetical order. Most entries consist of several articles that deal with various aspects of a topic and are arranged in a logical sequence within an entry. Some entries comprise a singlearticle. TohelpyourealizethefullpotentialofthematerialintheEncyclopediawehaveprovidedthreefeaturestohelpyou findthe topicof yourchoice: a ContentsList, Cross-References andanIndex. 1. Contents List Yourfirstpointofreferencewillprobablybethecontentslist.Thecompletecontentslistwillprovideyouwithboththe volumenumberandthepagenumberoftheentry.Ontheopeningpageofanentryacontentslistisprovidedsothatthe fulldetails of thearticles within theentry areimmediately available. Alternativelyyoumaychoosetobrowsethroughavolumeusingthealphabeticalorderoftheentriesasyourguide.To assist you in identifying your location within the Encyclopedia a running headline indicates the current entry and the currentarticle within thatentry. You will find ‘dummy entries’ where obvious synonyms exist for entries or where we have grouped together related topics. Dummyentriesappear inboththe contents listandthe body ofthe text. Example Ifyouwere attempting tolocate material on yield tables andforecasting viathe contentslist: YIELDTABLESseeMENSURATION:ForestMeasurements;GrowthandYield;TimberandTreeMeasurements;Yield Tables,Forecasting, Modelingand Simulation. ThedummyentrydirectsyoutotheYieldTables,ForecastingandSimulationarticle,intheMENSURATIONentry.At the appropriatelocation inthe contents list,the page numbersfor articles underMensuration are given. IfyouweretryingtolocatethematerialbybrowsingthroughthetextandyoulookedupYieldTablesthenthefollowing information wouldbeprovided in the dummyentry: YieldTablesseeMensuration:ForestMeasurements; Growthand Yield;TimberandTreeMeasurements; Yield Tables, Forecasting, Modelingand Simulation. Alternatively, ifyou were lookingupMensurationthe followinginformation could beprovided: MENSURATION Contents Forest Measurements Timberand TreeMeasurements Growth andYield Yield Tables,Forecasting, Modelingand Simulation Tree-Ring Analysis
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