ANNOTATED CHECKLIST OF THE VASCULAR FLORA OF THE BEECH CREEK UNIT OF THE BIG THICKET NATIONAL PRESERVE, TYLER COUNTY, TEXAS PDF
Preview ANNOTATED CHECKLIST OF THE VASCULAR FLORA OF THE BEECH CREEK UNIT OF THE BIG THICKET NATIONAL PRESERVE, TYLER COUNTY, TEXAS
ANNOTATED THE VASCULAR FLORA OF THE CHECKLIST OF BEECH CREEK UNIT OF THE THICKET NATIONAL PRESERVE, BIG TYLER COUNTY, TEXAS Brown MacRoberts and MacRoberts Michael Barbara Larry H. R. E. Vm, Bog 740 Columbia Spring Branch Science Center Research, 8856 Westview Drive Shreveport, Louisiana 71 104, U.S.A. Suzanne Birmingham Walker Department c Highway 87 South 14671 State Texas 75973. Texas 77005, Sheibyviile, U.S.A. iston, U.S.A. ABSTRACT 5S£ i^^ RESUMEN INTRODUCTION This manuscript the 5th part of an intended inventory, based on voucher specimens, of the vascular flora is we of the Big Thicket National Preserve, southeastern Texas. In 4 previous papers, provided an annotated Sandy and Lance (MacRoberts checklist of the Hickory Creek, Turkey Creek, Big Creek, Rosier units et we 2002; Brown 2005, 2006a, 2006b). In paper, provide an annotated checklist of the Beech this et al. al. Creek Unit. AND METHODS SITE West The Gulf Big Thicket located mostly within the longleaf pine region of the Coastal Plain in south- is & & Harcombe Marks Watson Marks Harcombe Harcombe eastern Texas 1979; 1979; 1979; 1981; (Ajilvsgi 1993; Diggs 2006 and references therein). The Big Thicket National Preserve consists of 12 et et al. al. units scattered over 7 Texas counties. The units range from 222 to 10,100 hectares and total about 34,000 The and hectares (Peacock 1994). Beech Creek Unit consists of 2,059 hectares located in western Tyler is County (Fig. relatively with poor drainage over most of the area. Elevation ranges from about 1). It is flat 43 meters 62 meters. Beech Creek and Beech Creek run through the unit. The Beech Creek Unit to Little was chosen inclusion in the Big Thicket National Preserve because of extensive stands of beech forest for its & & Harcombe Marks Watson Marks Harcombe dominated by lower (Ajilvsgi 1979; 1979; 1979; 1981). is It slope hardwood-pine forest (mesic lower slopes and terraces), with lesser amounts of baygall, mid-slope hardwood and oak-pine floodplain hardwood-pine forest (mesic stream bottoms), floodplain forest forest, & & Harcombe Van (Harcombe Marks Marks Kley (seasonally flooded river floodplain) 1979; 1981; 2006). A map of community types can be found in Harcombe and Marks (1979). Fire suppression over the past 30 years has resulted in extensive midstory development and thus reduction of herbaceous vegetation that t Journal of the Botanical Research Institute of Texas 2(1) L®- »51^ \ ALABAMA COUSHATTA - INDIAN RESI •4®^"""*" \J| Beech { \CreekUnitjl SANDYj^ V BIG creek gBjj UNrr w AR ^ \ 1\\ MENARD CREEK (CORRIDOR \ I UNIT F JACK GORE * y* w fer V ^ BAyDU CORRIDOR UNIT] ^^— \j k><5)- n-^ HARDIN 9CT\ |\ 5? r,.^ £ -^L 4 it *--~--' 1 Location of the Beech Creek Unit, Big Thicket Nation, 1. on September adapted to frequent Hurricane Rita, a category 3 hurricane, passed over the unit 24 is fires. decompose. 2005 damaging and thousands probably take decades toppling of will for this litter to trees. It Watson Details about topography, and climate can be found in Deshotels (1978); (1979, 1982, 2006); soils, Harcombe and Marks Marks and Harcombe and Diggs (2006). (1979); (1981); et al. been Like most units of the Big Thicket National Preserve, the Beech Creek Unit has not the subject of a detailed inventory. floristic Larry Brown 290 specimens during 9 days beginning 9 November 2006 and ending 17 collected field October 2007. The MacRobertses collected 154 specimens during 4 field days in 2004 and 1 day in 2007. 2004 and day Sandi Suzanne Walker collected 101 specimens during 2 days in additional in 2007. Elsik 1 and Warren Pruess together gathered 82 specimens during 2005. Geraldine Watson collected 25 specimens 3 2 3 1 Brown et a Vascular flora of the Beech Creek Unit, Big Thicket 653 I., over several years in the 1970s and early 1980s. Warren Pruess collected another 11 specimens over a late A&M We 2 year period. Other collectors totaled 12 specimens. conducted a search of the Texas University (TAMU) herbarium consortium base (http://www.csdl.tamu.edu/FLORA/tracy/main/html/) ad- data for ditional specimens but found no additional taxa not reported here. known Because our purpose was to produce a of taxa to occur on the Beech Creek Unit, a vouchered list specimen was considered be the only evidence acceptable inclusion in the In 675 herbarium to for list. all, specimens form specimens have been annotated by Larry Brown. the data for this report. All In most cases, nomenclature follows Jones et (1997), Kartesz (1999), Turner et al. (2003), and Diggs al. (2006). In cases of multiple collections, no more than 4 are listed for each taxon. et al. RESULTS The following an annotated of the Beech Creek Unit taxa. Abbreviations used are given below. is list SW thaceae Chapm., coriacea (Pursh) 01 Ilex Chapm. PH 002 ovata (Walter) Lindau lanceolata (Chapm.) R.W. Ilexlongipes exTrel., la var. MM SW 46 7056 ung, llexopaca Aiton, 67, MM SEWPLB LBSH 7058 r Gmel.) Steud., 488; llexvomitoria Sol. in Aiton, (J.F. 2151 Araceae MM 50 a humilis Nutt., 71 SW Ahsaema triphyllum Schott, (L) 0, 1 MM ubrum 7078 L, u 5EWp 245 spjnQsa ArajJQ GW SW 2261 Aristolochiaserpentaria L, 05; scopallinumL,SBNP2' Asclepiadaceae SEWP 230 Asdepios perennis Walter, LBRH 32061 variegata Asclepias L, SW Aspleniaceae bra (Michx.) Dunal, 03; LB 3 321 11 SW Ambrosia artemisii 03 Centella erecta Fernald, (Lf.) SEWP WP Ambrosia 265 L, Chaerophyllum Hook. trifida tainturieri var. tainturieri, 1 1 SW LBRH 05 SEWP Baccharis halimifolia 32071; maculata 414 L, Cicuta 1 L, 33034 LB & Bidens aristosa (Michx.) Briton, lum leptophyllum Sprague ex Britton (Pers.) P. SEWP 33030 SW LB Bidens bipinnata L, 252; 58 Wilson, SW Bidens frondosa LB 33003 DC, L, Eryngium prostratum ex 68 Nutt. MM 99 SEWP 32783 Boltonia diffusa 71 Hydrocotyle Thunb., 262; LB Ell., verticillata MK SEWPLB +Bradburia Semple, pilosa 491, 501; LBSH 32122 (Nutt.) x.)Raf.,LB 3 1994; SW 97 71 Sanicula canadensis L, 71 ^ WP Sanicula smallii E.R Bicknell, 1 1 ^^ Apocynaceae W^mMLKronquist, J^ LB 32466 ^ Trachelospermum difforme (Walter) Gray, LB 3 998 ^J.^ A. 1 5EWp LBJL 6a 335; (L) } SW Aquifoliaceae 063 Coreopsis lanceolata L, SEWP llexambigua (Michx.) Torn, PH 001 Coreopsis tinctoria Nutt., 253 6 5 3 97 Journal of the Botanical Research Institute of Texas 2(1) SW 32959 Elephantopus carolinianus Raeusch., 74; LBJL Elephantopus nudatus LB 33002 Gray, A. Elephantopus tomentosus LB 28598a, 28353; 327; L, LBJL 3261 SEWP DC, 342 ex Erechtites hieraciifolio Raf. (L.) 943 *Cardamine LB hirsuta L, 3 1 MM LBRH 32060 Muhl. ex 7139; Erigeron strigosus Willd., Cardamine parviflora arenicola Schultz, LB var. (Britt.) L. 32457 I 31942 & LBSEWP Eupatorium lancifolium (Torn A. Gray) Small, 3 899; MM Bromeliaceae 7187 SEWP 395 usneoides SEWPLB Tillandsia L, 496 (L.) Eupatorium mohrii Greene, MM SW 47 Eupatorium perfoliatum L, 83; 71 Buddlejaceae 33004 Eupatorium xpinnatifidum LB PM Elliott, Polypremum procumbens LBJL 32601 L, 04; SW 33039 Eupatorium LB rotundifolium L, 39a; Burmanniaceae MM SW 33009 LB Apteria aphylla Barnhart, 7224; 88; PM SW43; 33016 Eupatorium serotinum LB Michx., 02; SEWP 422 BurmanniabifloraL, & Greene Euthamia ex ieptocephala Gray) Porter (Torn A. LBSEWP 31894 Callitrichaceae Britton, MM LBRH 32048; 7683 LB 3201 retusa (Lam.) Sch. Bip, Callitrichepeploides Nutt., 'Facelis SEWP 255 puichella Foug.. var.pulchella, Gaiilardia Campanulaceae MM Gamochaetacoarctata Kerguelen, 7106 MM (Willd.) SW 7209 Lobelia cardinalis L, 84; SW061 Gamochaeta purpurea Cabrera, MM (L) DC, LBSH 49 appendiculata 7673; 321 Lobelia A. MM amarum amarum, Heienium Rock 35 71 GW (Raf.) H. var. SW 934 puberula Michx, 2249, 2336; LB 3 Lobelia 34; SEWP 33023 Helianthus angustifolium 403b; LB L, & LBRH bra Pavon) Greene, LB 32009; 32059; (Ruiz 32494 Hieracium gronovii L, LB MM 7674 Hymenopappus DC. artemisiifolius LBRH 32059 Nieuwl. Triodanisperfoliata (L.) 7148 WP DC, marginata LB 32487; (Thunb.) A. 21 lia 1 ; Cab *Hypochaeris microcephala (Sch. albiflora Bip.) re ar. SW110 MM LBRH (Kuntze) Cabrera, 32075; 7661 Caprifoliaceae LBRH Chambers, LB 32001; 32051 Krigiacespitosa (Raf.) MM MM Thunb, 7162 *Lonicerajaponica 7678 Willd., .) SW Sambucus SW nigra subsp. canadensis 60, R. Bolli, (L.) Mikania scandens 86 L. (L) Willd., 33, BT050 66 Lactuca canadensis LB 321 L, SW MM Viburnum 57 acerifolium L, Pluchea camphorata DC, 721 721 LB 32804 (L) 2; 1 , SW MM SW Viburnum dentatum 48 L, DC, 92 Pluchea 7236; foetida (L.) nudum Viburnum nudum, 2260b; 321 Wooten & var. Pyrrhopappus LB L. carolinianus (Nutt.) Standi., ( Celastraceae WP SW Pyrrhopappus Don) DC, 25 pauciflorus SEWP 4' (D. 9; 1 1 1 L, MM LBRH 32074 Rudbeckia hirta L, 71 2; 1 SW Solidago LB 3301 altissima L„ 00; 1 SEWPLB 502 Solidago ludoviciana Gray) Small, (A. LBSEWP Solidago rugosa aspera (Aiton) Fernald, Mill. var. SEWP 33024 31887; 331; LB MM SW SW SEWP *Sonchus asper (L) LB 32006; 544; Clethra alnifolia L, 20; 71 Hill, 1 1 1 MM Nesom, 7206 (L) G.L. i Commelinaceae MM WP & Symphyotrichum lateriflorum A. D. Love, 120; (L.) Commelina LBSH 26 321 erecta L, GW SEWPLB 7668; 2248; 481 MM Commelina LBSH 20 321 virginica 71 55; L, LBSEWP Symphyotrichum racemosum G.L Nesom, (Elliott) Convolvulaceae 31886; LB 33001 MM Michx, Dichondra carolinensis 71 61 Berberidaceae Ipomoea Dennst cordatotriloba cordatotriloba var. LB 33031 350; Ipomoea pandurata Mey, LB 32493 G. (L.) L,SRNP3t Betula nigra CarpinuscarolinianaW< Ostrya virginiana {M\\\) SEWP NyssaaquaticaL, 362 SEWP 397 Nyssa Walter, biflora MM SEWP Marsh, 7081 Nyssa 275; sylvatica 6 8 9 5 1 1 Brown et al., Vascular flora of the Beech Creek Unit Big Thicket Crassulaceae Penthorum 32805 sedoides LB Cupressaceae MM MM SW Juniperus virginiana L var. virginiana, 7076; LB 33032 lilinum Kuhn, 96; 71 33 (L.) ;r/c/7um(L.)L.CRich,SW42 Cucurbitaceae Matsum & lonatus (Thunb.) Nakai LB 3301 *Citrullus h,SW27;MM7654 Cuscutaceae MM Cuscuta compacta Juss. ex Choisy, 7231; LB 33007 Cyperaceae Carex albicans Willd. ex Spreng. var. australis (LH. Bailey) LB31936 Rettig, SW Carex (Dewey) Mack, ex 27 brevior Lunell, 1 32004 Carex cherokeensis Schwein., LB MM complanata & Carex LB 7677 Jorr. Hook., 31 953, 31 989; Carex corrugate LB 3201 Fernald, SEWP wa(L)A.Gra Leucothoerac 359 :emt ^Schkuhr ex Willd. var. fioridana (L.H. Bailey) Naczi y, MM MM DC, 23 & LBSH L) 71 Bryson, LB 31947, 32007; 32133; 7672 MM16 Don, Carex fiaccosperma Dewey, LB 3201 3201 la (L) D. SE^m 4, raL,SW89;f 33035 LB 71 32175 17; Carex/ran/c//Kunth, LB MM Rhododendro hx.)Sweet,WP112 Carex giaucescens 721 LBJL 32600 Elliott, 0; MM SW MM SW Rhododendrc LBSH 7122; escens Rudge, LBRH 32046; 7663; rorr., 18; 29; 32129 PH005 10 ZumSw mi 31950 130,45; LB 19, Euphorbias GW Acalyphagrac 2255 99; tUSmc SEWP Chamaesycer 247 ulata ill, Cyperuse Sme SEWP ns 251 (Lag.) ill, SEWPLB MM *Cyperm 503 >eck., 7200 Croton capita. rfichx., Cyperus 32794 erythrcrhizos Muhl., LB SEWP Croton glandt us var. sep:tentrionalis Muell.-Arg., L. SEWP Cyperus haspan LB 32481 L, 356; 270 SEWP 400 Cyperus odoratus L, SEWP Croton monanthogynus 259 Michx., Cyperus pseudovegetus Steud, LB 32165 32490 Croton willdenowii Webster, LB G.L. MM Cyperus 7666 virens Michx., 32807 Euphorbia LB coroliata L, LBSH 32134a Eleocharis microcarpaTorr., SEWP 32460 Euphorbia dentata LB Michx., 254; Roem&Schult., berculosa (Michx.) 895; LB: 32475 Phyllanthus caroliniensis Walter, LB MM MM LBRH 7662 MM 7233; 32047; *Phyllanthus urinaria 71 L, 1 32484 KyllingaodorataVahl, LB SW jticosa (W. Bartram) Fernald, 14 SEWP Kyllinga pumila Michx., 399 SW *Triadica sebifera Small, 01 (L.) MM Rhynchosporacaduca LBSH 32150 7127; Elliott, SEWP Rhynchospora corniculata (Lam.) A. Gray, 231 >5/nDurazz.,MM7107 +Rhynchosporadebilis LBSH LBRH 32087 32140; Gale, lum Centrosema LBJH 32604 Benth, virginiar (L.) Rhynchospora LBSH 32214 A. elliottii Dietr., w. SEWP L* canadensis canadensis, 271 Cercis Rhynchospora 32479 LB fascicuiaris (Michx.) Vahl, SEWP Chamaecrista fascic ulata (Michx.) Greene, 276; LBJL Rhynchospora (Chapm.) LBSH globularis Small globuiaris var. 32605 32132,32220 m Moench, Chamaecrista LB 32461, 321 32486, SW nictita (L) 79, Rhynchospora glomerata LB 32472, 3301 Vahl, 98; (L.) 32806 MM Rhynchospora inexpansa (Michx.) Vahl, 71 1 32488 MM ..,SW 65; LB Rhynchospora 33038 mixta 7074; LB Britton, Desmodium Auhl ex DC, SEWPLB 497 ciliare Willd.) {I Rhynchospora plumosa LB 32022; LBRH 32078; Elliott, m GW Desmodium DC, 2337 glabeliu (Michx.) SW 7676; 109 7ftimDC,MM7189 MA Rhynchospora Chapm. LBSH 3214 ex pusiila Curtis, SEWP 41 21 ., 32489 LB MM Britton, Scleriaoligantha Michx., LB 31 995; 7671 SEWP 43 MM T(Jacq.) Harper, merata Michx., 7192; LB 32019; LBJL 32593; *Kummerowia (Thumb.) LBSH 32 28 Schlindler, striate SW120 i 1 8 2 3 Journal of the Botanical Research Institute of Texas 2(1) SEWPLB 495 Carya aquotica (Michx. Nutt., f.) SEWP Carya texana Buckley, 41 31941 SW 28 & LBSH Carya tomentosa (Lam. ex Nutt., ex Rose) B.L Turner, Poir.) hystricina (Small Britton so 32119 LBRH 32050 Rhynchosialatifoiia Nutt, LBRH LBSH Juncus acuminatus Michx., LB 32005; 32069; SEWP Strophostyles umbellata (Muhl. ex Willd.) Britton, 347; 32142a LBSH LB 32809 321 18; 31997 (Wiegand) LB Juncus antheiatus R.L Brooks, SEWP & 235 Poggenb., Styiosanthes blflora (L) Britton, Sterns, SW 40 us Mack., MM LBSH 321 52 campestre *Trifolium Schreb., 71 51 LBRH 32070, 32077; LBSH 321 ; sissimus Buckley, 23; L.SWU4 latum 32473 LB lappaceum 72 LB 321 *Trifolium L, 31938 Juncus effususL, LB rem LB 999 MM 31 (L) Poir., LBRH 32049 Juncus marginatus Rostk, 7130; s)Swe SEWP 355 Michx., MM 7665 LB 32002; subsp. nigra (L rt, 32799 Lam., LB aides 32020 Juncus LB tenuis Willd., BRH: 2088 SEWP 33021 LB 408; Wood) 935 Smyth, LB bulbosa 31 Luzula (A.W. Lamiaceae SEWP 404 Shinners, Hyptis alata (Raf.) SW :a Bartr. ex Willd., 54, 55; L SEWP Moench, 398 Lycopus rubeilus MM SW 32797 LB 7227; licus L, 80; MM Monarda ex 7203 citriodora Cerv. Lag., & SEWPLB 499 Pycnanthemum albescens Torn A. Gray, Pycnanthemumte LB 321 76 MM 7096 L, SEWP LB 3201 SEWP 33010 Muhl. ex Spreng., 239; 340; LB Scutellaria elliptica rell, um(Sw.)Druce,SW108 69 WP Geranium caroiinianum L, 1 1 Grossulariaceae SEWP 403a; LB 32024, 32784; ilea virginica L, IV Haloragaceae SEWPLB 489 _., Han 32594 7054 Sara, michauxii LBSH 321 36 to t Lilium Poir., /I MM 2289 33022 Nothoscordum 7216; LB bivalve (L) Britton, Hypericaceae texanum (Planch.) Fernald, Britton var. i.) & & Hook.) Torr. A. v. .MM 2259;LBJL32617 LBSH 46 321 71 74; GW 2257 Hypericum 7207; galioide VI MM SW Aiton 7067; LB 07; ?ns (L) f., SW SEWP 31896; 95 MM SW Hypericum hypericoides Crantz, 73; (L.) & iray,SW75;LB32802 Gmel.) Mitreola petiolata Torr. A. (IF. 32599 32063; LBJL SW Lygodiaceae Hypericum mutilum 25 L, 66, MM SW SW Sw., 87; ray) Triadenum Gmel.) Gleason, 85 waiter! 35, (J.G. SEWP 543 campestre Sisyrinchium Bicknell, E.P. SW WRC *Cuphea carthagenens /lacbr, 30; LB 31 9i ell, 3201 Bicknell B E.P. i; tj 0;SW 1 7 3 Brown etal., Vascular flora of the Beech Creek Unit Magnoliaceae MM SEWP 437 Magnolia grondi 70* linata flora L, Mill., Magnolia virginianaL,S\N 15 MM LBRH 32079 P/antago 7094; w/g/n/cfl L, SEWP occidental 363 Piatanus L, Poaceae & 5EWPLB 504 Agrostis hyemalis (Walter) Britton, Sterns Poggenb., edarac/7 L, MM 7675 32085; LBRH 32054 *Aira caryophyllea var. capillars Mutel, L. MM 7' _.,SW101 iropogon Virginia MM SW LBSH Morella caroiiniensis Small, 21; 7090; 32121 32790 Morella Small, LB cerifera (L.) Oleaceae SEWP 409 Chionanthus virginicus L, GW MM SW Fraxlnus americana L, 721 7063; 2252; 3, 1 1 MM LBRH 7658 *Bromuscatharticus 32072; Vahl, Fraxinus pennsylvanica Marsh., LB 32795 MM MM Chasmanthium laxum 04 (L) H.O. Yates, 71 7146 *Ligustrum sinense Lour., SEWP 68 Ludwigia altemifolia L, 261 236; LB 321 , GW Ludwigia decurrens Walter, 2333; LB 32801 33000 LB 32459, :u/are, SEWP Ludwigia glandulosa 401 Walter, & Gould (Sw.) C.A. Clark var. ) SEWP 358 Ludwigia leptocarpa Hara, (Nutt.) LBRH LBSH 32142 !2089; n, SEWP &CA MM Ludwigia pilosa Walter, 406; LB 32796, 3301 Gould 7165; LB Clark, ) Ludwigia LB 31 993 palustris (L) Elliott, 31990,32180 SEWP Oenothera 392 biennis L, SW & LBRH 06 Oenothera 32052; laciniata Torr. A. Gray, LBSEWP 1 31949 tatum 31891; LB LBRH 32056 Nutt, dichotomum Gould dichotomun hantheiium subsp. (L) MM SEWP LBSH 34 Ophioglossaceae 332; 71 37, 7071; 321 SW Underwood, 78 Botrychium biternatum (Sav.) Freckman & LBRH (Muhl. ex Lelong, 32083; L Elliott) Orchidaceae SW LBRH praecox Watson., 32066, Spiranthes (Walter) 41; S. G v n dichot (L.) & Freckman LBSH 32131 (Ashe) Lelong, Gould Dichanthelium oligosanthes oligosanthes, (Schult.) var. LBRH 32064 LB 32018, 32471; MM 7136 Dichanthelium polyanthes Mohlenbr., (Schult.) & Gould turn C.A. Clark, (Elliott) LBRH 32086 MM 33005 Dichanthelium scoparium (Lam.) Gould, 721 LB 8; & Freckmann LBRH Lelong, 32044; ) LBSH 32138,32123a, 32121a MM vr/^M GW Gray) Turner, 7095; (A. 32458 KoeL, 2330; LB 32476, Digitariaciliaris (Retz.) SEWP 248 *Eleusineindica Gaertn., (L.) 32177 Elymus LB virginicus L, 33040 Nees, LB (Michx.) MM 7230 Eragrostisrefracta (Muhl.) Scribn., 33025 LB Eragrostisspectabilis (Pursh) Steud., 5 Journal of the Botanical Research Institute of Texas 2(1) 32010 mNutt., LB V LBSEWP Mild, 31892; LBRH 32073 L, MM 5EWP 237 7168 Diodia virginiana L, alter, m SEWPLB 492 Alton, »//us(L)P.Beauv.subsps< PLB 483 Houstonia pusilla Schoepf, LB 31 933 32485 LB Michx., GW SEWP 2256; LB 364; i., SEWP 396 es, MM WRC1 33006 Panicum verrucosum MuhL, LB 402; 71 93; 1 32463 *Paspaium diiatatum LB Poir., SEWPLB 494 Paspaium Nash, langei Fourn.) (E. MM SEWP 233 *Paspalum nototum 7158 Lam.) Fluegge, Terrell, MM SW Paspaium plicatulum 90 Michx., 71 64; 32478 Paspaium setaceum LB Michx., 7132 MM GW SW *Paspalum 7202; Steud., 70; urvillei Scrophulariaceae % LBSH 3: MM SW SEWP Bacopa monnieri Pennell, 243; 7681 112 avena 992 (L.) Piptochaetium ; GratiolapilosaMchx., LB 32021 L,WP117 annua *Poa 31938 SW LB Gratiola virginiana L„ LBSEWP *Sacciolepis indica (L) Chase, 31897; 0' 33014 Crustacea MuelL, LB *Lindernia (L) F. Dumort, 320 *Schedonorusarundinaceus LB (Schreb.) SW Mecardonia acuminata (Walter) Small, 76; 7223; IV 32803 Schizachyrium scoparium (Michx.) Nash, LB MM Sphenopholis intermedia (Rydb.) Rydb., LB 31 985; SEWP umbrosum Micra'nthemum Gmel.) 352; Blake, (J.G. LBSH 32145 Sphenopholis obtusata (Michx.) Scribn., MM GW 7667 SW 2329 Sporobolus indicus 02; (L.) R. Br., 1 Mimulus LB 32787 SW alatus Aiton, 03 Tridens flavus Hitchc. var. flavus, (L.) 1 LBRH LB 31944; Nuttallanthus texana (Scheele) Sutton, LBRH 32057 Vulpia octofiora (Walter) Rydb., 32062 Polygalaceae MM GW SW 2254 3360a, Scoparia dulcis L, 7221; 79; Polygala mariana LBJL 3261 Mill., LB 940 "Veronica arvensis L, 31 polygama LBRH 32045 Polygala L, Polygonaceae SEWP 268 Smilaxbona-noxl., SEWP 402a Brunnichiaovata (Walter) Shinners, MM 7145 Smilaxglauca Walter, SEWPLB Polygonum punctatum 485 Elliott, SW SEWP 24 a 41 L, 7; Rumexhastatulus LBRH 32055 MM Baldw., 7070 Smilaxpumila Walter, 71 5, 1 SEWP 274 Polypodiaceae Smilaxrotundifolia L, //Morong,MM7086 pod poly bides Pleopeltis f vs&Windham,GW53;MM7105; ara(Weatherby)Ar SW SEWP 402b; 81 Primulaceae *AnagallisarvensisL,M MM Symplocos 71 tinctoria Her., (L.) L. 1 < SEWPLB 505 Halesiadiptera Ellis, SW 04 Styraxamericanus Lam., Morton, Thelypteris kunthii (Desv.) C.V. L 7156 Crataegus marshallii E GW 2335 laevigata Willd., Celtis LBSH Crataegus ok., 3: GW SW MM Ulmus 2334 atata Michx., 56; 7064 Crataegus spathulata Michx., 7166, SEWPLB 493 Ulmus americana L, 7Gf/ra(Andr.)Focke,MM7215 MM 7068 liana Aiton, (Mill.) SEWP 426 taEhrh., SEWPLB 482 Link, > 3 : Brown et Vascular flora of the Beech Creek Unit, Big Thicket al., primulifoliaL,SWU Ware, rodiata Dufr. parviflora (Dyal) Egg. Viola (L.) f. GW 2292 LBRH 32053 Viola sagittata Alton, 31986 ild.,GW 2294; LB Verbenaceae MM americana 7084 Callicorpo L, SW SEWP 256 Ampelopsisarborea Koehne, 29 (Sweet) Troncoso, ilchella (L.) SW SEWP 250 Greene, 94 Parthenocissus quinquefolia (L) Planch., ..) MM GW SEWP 2428 258 7079; HiensisVeW., chx., MM MM 7055 7143 Verbena Small rotundifolia Michx., halei Vitis SEWP 257 *Verbena Spreng., rigida Xyridaceae LBSEWP Chapm. Violaceae 31893, Xyris difformis var. difformis, MM MM 7237 LB 3301 7204, lanceoiata 7092; Viola L, MM GW PM SEWP 32462 LB Viotapatmata 7159; 2295; 05 +Xyris]upicai L.C. Rich., 41 354, 328; L, 9, Addendum (whorled pogonia) and sugar maple (probably either Acer barbatum or Acer leucoderme) Isotria ve ai rticillata on Beechwoods Guide, mimeographed guide available the entrance, b\ listed for the unit the Trail a at trail we them during our two- year survey, and could not find herbarium material of sami failed to collect field making Geraldine Watson (2006) also indicated that sugar maple (Acer barbatum) a remarkable recover is with removal in this unit the of cattle grazing. Carex The specimen was immature The section ovales. in this collection too for specific identification. was Perhaps was an immature only other Carex in section ovales C. brevior. this C. brevior. We Bartonia texana. Currently Beech Creek the only Big Thicket unit to have this rare Texas endemic. is made two sphagnum mat Beechwoods and one from any near collections, one in a along the Trail, far trail, Beech Creek with Beech Creek. the intersection of Little We An name Bradburia pilosa. are following Semple (2006). equally correct is Chrysopsis pilosa Nutt. Brown Rhynchospora This Texas and the the Pineywoods. See debilis. the third collection for for first is (2007) additional information. et for al. Xyris jupicai. There no consensus the native or introduced status of this yellow-eyed-grass. as to is DISCUSSION There are 103 families and 229 genera represented for the 470 taxa (418 native) on the Beech Creek Unit list. Lamiaceae Cyperaceae Fabaceae Eight families: Poaceae (59 taxa), Asteraceae (53 taxa), (40 taxa), (26 taxa), 48.9% and account Juncaceae (15 taxa), Euphorbiaceae (13 taxa), Scrophulariaceae (11 taxa), (11 taxa) for The Rhynchospora of the taxa collected. largest genera are: Carex (15 taxa), Dichanthelium (13 taxa), (11 taxa), and Juncus (10 taxa), Quercus (9 taxa), Cyperus (7 taxa). may While is the case that our list is incomplete, as are all floras, and that a few taxa reported here no it longer exist on the unit, specimens of most taxa that have grown in the Beech Creek Unit during the past We 95+ Beech Creek Unit has about quarter century have been collected (probably %). estimate that the 450 425 native species/taxa. to ACKNOWLEDGMENTS CA Agreement 14001004 No. This study was supported in part by a National Park Service Cooperative to from Harcombe and by an Taxa Inventory (ATBI) grant the author the Paul in part All Biological to first The The were from author Big Thicket Association of Saratoga, Texas. page charges paid for this grant. first Deanna Boensch and Zhou, Thicket his passage across appreciates Lily Big ecologists, for facilitating fire him private land the northeast corner of the unit and then walking with along Little Beech Creek south to Journal of the Botanical Research Institute of Texas 2(1) new with Beech Eleven near the confluence Creek. additic to Thanks Ray and Keith all-day hike. also to C. Telfair Eric for II careful reading of the paper. A&M and western Texas College 1979. Wild flowers of the Big Thicket, east Texas, Louisiana. Press, G. Ajilvsgi, Station. and Annotated W.W. 2005. Brown, MacRoberts, M.H. MacRoberts, RA. Harcombe, Pruess, D. Johnson. L.E, B.R. I.S. Elsik, and Hardin of the vascular of the Turkey Creek Unit of the Big Thicket National Preserve, Tyler checklist flora 807-1 counties, Texas. Sida 21:1 827. W.W. and 2006a. Annotated Brown, MacRoberts, M.H. MacRoberts, RA. Harcombe, Pruess, S.D. Jones. Elsik, L.E., B.R. I.S. Sandy County, checklist of the vascular flora of the Big Unit of the Big Thicket National Preserve, Polk Texas. 22:705-723. Sida PA W.W. and 2006b. Annotated Brown, MacRoberts, M.H. MacRoberts, Harcombe, Pruess, S.D. Jones. Elsik, L.E., B.R. I.S. checklist of the vascular flora of the Lance Rosier Unit of the Big Thicket National Preserve, Hardin County, 175-1 Texas. Sida 22:1 189. and and 2007. Notes on the of Texas with additions other significant Brown, E.L DJ. Rosen, flora Keith, Liggio. L.E., J. Texas 1:1255-1264. records. Bot. Res. Inst J. USDA Conservation Deshotels, J.D. 1978. Soil survey for the Big Thicket National Preserve, Texas. Soil Service, College Station, Texas. Diggs, G.M., B.L Lipscomb, M.D. Reed, and RJ. O'Kennon. 2006. Illustrated flora of east Texas. Sida, Bot. Misc. 26:1-1594. PA Unpublished Harcombe, and PL Marks. 1979. Forest vegetation of the Big Thicket National Preserve. report: New Park Service, Santa Mexico. U.S. Fe, Harcombe, PA, Glitzenstein, R.G. Knox, Si. Orzell, and E.L. Bridges. 1993. Vegetation of the longleaf pine region J.S. of the West Gulf Coastal Plain. Proc. Ann. Tall Timbers Fire Ecol.Conf. 18:83-103. and Montgomery. 1997. Vascular plants of Texas. Univ. Texas Press, Austin. Jones, Wipff, P.M. S.D., J.K. Asynonymized and with the vascular of the United KarteszJ.T 1999. checklist biological attributes for flora atlas CA Meacham. American Canada, and Greenland. ed. Kartesz and Synthesis of North flora. States, st In J J. 1 North Garden. Chapel Version Carolina Botanical Hill. .0. 1 and Annotated of the vascular of the Hickory MacRoberts, M.H. MacRoberts, LE. Brown. 2002. checklist flora B.R., 20:781-795. Creek Unit of the Big Thicket National Preserve, Tyler County, Texas. Sida PL Monogr. Marks, and PA. Harcombe. 1981. Forest vegetation of the Big Thicket, southeast Texas. Ecol. 51:287-305. A&M Peacock, H.H. 1994. Nature lover's guide to the Big Thicket. Texas Press, College Station. New Semple, J.C 2006. Bradburia. ln:The flora of North America. Vol. 20. Oxford University Press. York. Turner, B.L, H. Nichols, G. Denny, and O. Doron. 2003. Atlas of the vascular plants ofTexas, Vol. Dicots, Vol. 2: Ferns, 1 : gymnosperms, monocots. Sida, Bot. Misc. 24. and O'Kennon, Van 2006. The pineywoods. G.M. Diggs, Lipscomb, M.D. Reed, RJ. Illustrated flora Kley, J.E. In: B.L. 76-106. 26:1-1594. of east Texas. Sida, Bot. Misc. Pp. Watson, G.E. 1979. Big Thicket plant ecology: an introduction. Big cket Mus. Publ. Ser., No. 5, Saratoga, Texas. Denton, Watson, 2006. Thicket plant ecology, an introduction, 3rd ed. University of North Texas Press. G.E. Big Texas. Mimeographed unpublished report. Big e.
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