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ANNOTATED CHECKLIST OF ARIZONA CONVOLVULACEAE DANIEL E AUSTIN' DepartmentofBotany ArizonaState University Tempe. AZ85287. U.S.A. ahsi"rac;t Specimensexamined in 1-1 iierbariaintlicatethatthereare30nativeant! naturalized speciesotConvolvtiiaceaeinthestateofArizona. Typesandselectspecimensarecited. Notesaregivenoncountydistribi-itions, habitats,altitudes,andHoweringtlares.Com- mentsaremadeontaxonomicproblems,abundanceandnaturalhistoryotselectedtaxa.A lectorypeforIpo/aoicihtterolihyllttOrtegaisselected. RiuSiimi:n Unarevisiondelasmuestrastie 14lierliariosindicaqueexisten30especiesnativasy naturalizadasparaelestadodeArizona.Secitanlostiposylosejcmplarcscstudiados.Sc notaladistribucionporloscondadcs,elhabitato,ladisrribucionaltitudinal,ylasfechasde floracion.Secommentasobrelosproblemasraxonomicos, lahistorianatural,ylaabun- danciatielostaxa. Seseleccionaunlectoti|io|âraIpomoeiiheterophylliiOrtega. Duringpreparationofthefamily("onvolvulaceaefortheVascularPlants ofArizona, certaintaxonomicnoteswerefound necessary(Austin 1990). The following annotated checklist, which includes all correct names of species known for the state, provides notes on several aspects of sys- tematics, abundanceandnaturalhistoryofArizonan morningglories. In somecases notesaregivenon the livingcolorsoffloral partssince these details are normally not part ot floristic surveys. Included are the correspondingnamesfromKearneyandPeebles(1951)andKearneyetal. (1960)andsomeadditionalsynonyms. Where types havenotbeenseen, theusageisbasedonotherauthorswhohaveseenauthenticmaterial, except in afew caseswhere it is based on the protologue. CALYSTEGIA 1. 1. Cai.ys'if.c;ia i,c>nc,ifi;s (S. Watson) Brummitt, Ann. Missouri Bot. Gard. 52:215. 1965. Tvpi;; NEVADA:in 1872,W'lwc/n-s./r (US').Co>2- viili'ulu.slonîJK'iS. Watson,Amer. Naturalist1\M)1. 1873. 'Fermanencatldress:DepartmentofBiologicalSciences,FloridaAtlanticUniversity,BocaRaton,FL 33431,U.S.A. SiDA l4(3):443-457. 1991. 444 — ConvAolRvIulZuOsNlAin:earM!atozbaittszaEalsrMwt.s,,PEraosct.ivCoaoldif1.7A2c6a4d.(CSAcSi!.;Seprh.o4r,o2o0f:K470s.pec19i.'mê1n.arNTYy!p)i-.: KnownfromCoconino, Gila, Maricopa, MohaveanciYavapaicos. The speciesgrowsinchaparralalthough ithasbeenfoundextendingintothe UpperSonoranzone; 609~1706 m; flowering March toOctober. Thisrarespecies, havingbeencollectedfewtimessincethe 1930sand 1940s, is endemic to the Southwest, from California(San DiegoCo. to InyoCo.andfromKernCo.toSanLuisObispoCo.),sNevada(ClarkCo.), Utah (Washmgton Co.) and Arizona. The species appears to be rare throughoutits range. Corollasarewhitewithacream-coloredthroat,andhavepinkpatcheson the limb around the nectar guides (interplicae). Styles, androecia and gynoeciaare white, but the nectary is yellow. Flowers arepollinated by Osmiasp. (Megachilidae). 2. Calysti-giasi-pujm(L.)R. Br. ssp. ancjulataBrummitt, KewBull. 35(2):328. 1980. Typi:: IDAHO. Canyon Co.: Macbride 318 (NY!). Calysteîaseptum(L.)R.Br.var.angulata(Brummitt)N.HolmgreninA.Cronquist ctal.,IntermounrainFl. Vase. Pi. IntermountainWest,U.S.A. 4:77. 1984. Known from a single specimen collected in 1882 from a garden in CochiseCo.; ca 1524 m; flowering inJune. TheseplantsareeasilyconfusedwiththeGreatPlainstaxonC.sylvatka (Kit.)Griseb. ssp.fraterniflora(MackenzieandBush)Brummittasshown (b1y97t8he).inTthereprqeutaadtriaotnesisninTursyeosnin(1t9h3e9)l,eaCfobrarseelslaofndCCosryrlevlalt(ka19a7l2l)owansdepLaerhar- tion from the V-shaped orU-shapedsinusesofC. sepium. 3. Calysthgiamac:ounii(Greene)Brummitt,Ann. MissouriBot. Gard. 52:215. 1965. Ti'PE:CANADA.Saskatchi:wan:Assiniboia,MilkRiver, Aug 1905,Macoun11883(notseen).ConvolvulusniacoumiGreene,Pittonia3:326. 1898. ConvolvulussepiumsensuKearneyandPeebles. — Convolvulus interior House, Bull. Torrey Bot. Club 32:140. 1905. Typi;: COLORADO:nearFortCollins,Crandall/625(NY!,US!). Known from Apache, Coconino, NavajoandYavapai cos. Growing in moist sites, near lakes and streams; 1950—2042 m; floweringJune to July. ThisrarespeciesisaGreatPlainsendemicthathadnotbeencollectedin Arizonasince 1971 untilitwasrelocatedin 1990{Austin&Austin7661, ASU). Corollas, stamens and gynoecia are white, although the nectary is yellow. Bees(notyetdetermined)visit the flowers. 445 2. CONVOLVULUS — 1. Convolvulus arvensls Linnaeus, Sp. PI. 153. 1753. Type: SWEDEN;specimen218.1(LINN, microfiche!). This introduced European weed was firstcollectedinArizonain 1843 neartheNavajoOrdinanceDepotinFlagstaffbyLt. R Shallert, anarmy surgeon.Ashorttimelater,in1882,LemmoncollecteditintheHuachuca MountainsnearFt. Huachuca.ThespeciesisnowknownfromallArizona countiesexceptLaPaz. AcollectionfromGreenleeCo. {Austin&Austin 7634,ASU)isanewcountyrecord.Commonindisturbedsites,roadsides, andcultivatedfields;341—2346m;floweringApriltoOctober.Common name: "Bind"weed." — 2. Convolvulus equitans Benth., PI. Hartweg. 16. 1839- Type: MEXICO: 1837,Hartweg9«(presumablyK). C. incanussensuauctt., nonVahi. KnownfromallArizonacountiesexceptYumaandLaPaz. Frequentin grasslandsandplains;762—1981m;flioweringMarchtoNovember.Local commonnamesare: "SilverBind"weed"(Little37, ARIZ), and"Desert Bindweed" {Wilcoxs.n., ARIZ). Thefilamentsarewhite, theantherspurple.Stylesandthebasesofthe stigmalobesarewhite,butthetipsofstigmalobesarepurple.Thenectary is yellow and cup-shaped. Visited by honeybees {Apis mellifera) and bumblebees {Bombussp.)(PimaCo., Austin&Austin 736], ASU). 3. CRESSA 1. Cressa TRuxiLLHNSis H.B.K., Nov. Gen. Sp. PI. 3:93. 1819. Typh: PERU; Trujillo,Humboldt&Bonpland3727(B:microfiche!, isotype: F!). — C,depressaGoodd.,Bot.Gaz.37:58. 1904. Type:NEV—ADA:Goodding726(UC!). C.insuiarisHouse,Bull.TorreyBot.Club33:315. 1906. Type;MEXICO:Revilla- gigedoIsls,Barkeleiv252(US!,UC!). — C.erectaRydb.,Bull.TorreyBot.Club40:466. 1913. Type;UTAH:Garrett870 (NY!). — C. mwrnuiHeller,Muhlenbergia8:140. 1913- Type:NEVADA;Hei/er&Kennedy 8663a(NY!).C.truxillensisH.B.K.var.minima(Heller)Munz,Aliso4:96. 1958. C.pumilaHeller,Muhlenbergia8;l42. tab. 17. 1913.—nomennudum. C.vaUicolaHeller,Muhlenbergia8;l40.tab. 17. 1913. Type:CALIFORNIA;Hel- ler8936a UC!). C. truxillensisH.B.K. var. vallicola(Heller) Munz, Aliso4:96. 1958. C.creticaL. var.truxillensis(H.B.K.)ChoisymDeCandolle,Prodr.9:440. 1845. Known from Coconino, Maricopa, Mohave, Navajo, Pinal and Yuma COS. Occasional in saline desert; 30—1524 m; flowering May to November. 446 PlantswerelastcollectedinMohaveCo. in 1912ijepsori3026, ARIZ); inPinalCo. in 1937{Peehles13233,ARIZ);inMaricopaCo. in 1964{Rea s.n., ASU); and in Navajo Co. in 1978 {Pinkava et al. 13839 ASU). ADElSt)h,outghhelpasltanctosllweectreedrienloYcuatmeadCion.1i9n8919{7A1us{tHiannu&ltoAruisst.inn.,7A3R86I,Z,ASAUS)U., Previouslytheplantshadbeenseparatedintospeciesorvarietiesonthe basisofgrowingerectorprostrate. Bothgrowthformsweregrowingside by side inYuma. Plants were not found in flower. 4. DICHONDRA 1. DicnoNDRA ARCiF.NiF.A WiUd., Hort. Bctol. 297. t. 81. 1806. Typi;:COLOMBIA:Tolimaca. Honda,Bo?il>land{l\'). Known from a single collection from Bisbee in Cochise Co. in 1931 {Hcirrisori8236, ARIZ). InNewMexicotheplantsgrow inChihuahuan desert scrub and Madrean oak woodlands; ca. 1615 m; flowering in September, earlierin New Mexicoand Mexico. ThespeciesisfrequentinTexasandNewMexicoasfarwestastheOrgan Mts. Perhaps the Arizona collection represented a western limit to the naturalrangeofthespecies,orperhapsitwasintroducedintoBisbeedur- ingtheminingoperations. PlantshavenotbeenrelocatedinArizonaeven though several people have searched for them. 2.DioioNDRABRACHYPooAWooton&Standlcy,Contr.U.S.Natl.Herb. 16:160. 1913. Type:NEWMEXICO.DonaAnaCc3,:OrganMountains, FilmoreCanyon, 23Sep 1906, Woofo/i&Slandleys.n. (US!). Known from Cochise and Santa Cruz cos. Occasional inMadrean oak woodlands and lower ponderosa pine zones; 1219~ 1889 m; flowering July toOctober. Although themorphological traitsgiven byTharp&Johnston(1961) arc sufflcient to distinguish species, they also differ by colors offlower parts.Thecorollalobesarecream,greenatbase.Filamentsandstigmasare green.Anthersarewhitewithapurplestripeandthenectaryisdarkgreen. Theovaryislightgreenandthestylesarecream.Maturefruitscollectedon 9 Sep 1989(Cochise Co. RuckerCanyon. Austin &Austin 7611. ASU). Fruits were fully maturebyOctober. — 3. DicHONDRAseric:i;aSwartz,Prodr.Veg.Ind.Occ. 54. 1788. Typh: JAMAICA:Swartz(S?). D. repensForster&Forstcrvar.serum(Swartz)Choisyin DeCandolIc,Prodr. 9:451. 1845. /). n'pcnsscn.suaucft., nonFor.srcr&Fonstcr. In Arizona the species isknown exclusively from the PajaritoMts. in , 447 SantaCruzCo. Rareinstreamsidevegetation;ca. 1112m;floweringMay to December. ThesingleArizonalocationisSycamoreCanyon(SantaCruzCo. from 1936,Goodding6620ARIZto 1962,Barr62-863ARIZ)neartheMexican border.Thispopulationwasrelocatedafteranhiatusof19years{Austin& Austin 7604, ASU), but its size has apparently dechned. Based on herbariumspecimensflowersandfruitsareuncommoninthispopulation. FlowerswerefoundinApril. Petals, anthersandgynoeciaarepalegreen {Austin&Austin 7604, ASU). No flowervisitorswereseen. 5. EVOLVULUS 1. EVOLVULUS ALSiNou:)r.s L. var. angustifolia Torrey, Bot. Mex. Bound. 150. 1858. Typi.:TEXAS.Pri-siuioCix:ca.the"GrandCanyon" oftheRioGrande.August,Parry(notfoundinCM,GH,ISC,MO,NY,PH,USor YU). U.iUiltsrifeicoihdtesL1.4:v3ar4..ac1a9p3u4l.cet—nnT(vWpiil-l:d.M)EOXoIstCsrOr..,GMueedredr.erBoo:t.caM.usA.caHpeurlbc.o,RiWjiklslUdneinvo.w 6128(B). KnownfromCochise, Pima, PinalandSantaCruzcos., andreportedly in GilaandMaricopa(Kearney and Peebles 1951, 1960). Occasional in pine-oak woodlands, saguaro desert scrub, and desert grasslands; 762—1828m;floweringFebruarytoOctober. Thecommonname: "Dio DI-:vibora"(Lehr 1978)mustrepresentanerrorinspelling, oranortho- graphicvarianteitherof"Tiodl:vniORA"or"Diosdl.vusora,"although1 havefound neitherofthesecommon names to be in use. Flowersopen8:00-9:00a.m.,andcloseca.4:00-5:00p.m.(PimaCo., Austin&Austin7398, ASU). Corollasareblueonthelimb,whiteinthe throat. Stamensandgynoeciaarewhite. Noflowervisitorswereseen. 2. Evo—lvulusarizonicu.sa. Gray, Syn. Fl. N. Amer. 2, 1:218. 1878. Type:MEXICO:Sonora:sandyprairies,Sep 1857,Th//r/?er1023(GH!).See Austin(1990)oncomplexitiesoftypification. — Evolvidi/slactinA.Gray,Proc.Amer.Acad.Arts17:228. 1882. Type:ARIZONA: 1881,Pring/es.n.(F!,GH!,US!).H.anzdniciLSA.Grayvar.ladus{K.Gray)Ooststr. Meded. Bot. Mus. Herb. Ri]ksUniv. Urrechr 14:76. 1934. Known from Cochise, Gila, Graham, Greenlee, Mohave, Navajo, Pinal, Pima, SantaCruzand Yavapai cos. Occasional to frequent indif- ferent partsofthe state; more common in thesouthern counties. Plants grow in chapparal, Madrcan oak woodlands, and mesquite grasslands; 883~ 1828m;floweringApriltoOctober.Commonname:"Falseflax" {McGinmes4, ARIZ). Flowers, which open 7:00-8:00a.m. and close 3:00-4:00 p.m., have 448 bluelimbs,whitethroats,andwhitestamensandgynoecia,andarevisited bybees(notcaughtbutpossiblyHalictidae).Pollenwasgoneandsomeof the anthers were stripped from flowers by 11:30 a.m. {Austin &Austin 7388, ASU). 3. EvoLvui.us NUTTALLiANiJs Roem. & Schult., Syst. Veg. 6:198. 1820. Typk:onthebanksoftheMissouri,Nuttall(Br*—). £.pilosiLsNiirr.,Gen.N.Amer.PI. 1:174. 1818,nom.superfl. Typf,:onthebanks II.oonfnpthhieliMniGsrsoeuernie,,NLueatftla,llBo(r^.^)O.bscrv.Grit. 1:151. 1905.—Type:NEWMEXICO: MetcalfeI22H(NMG, NY!,UC!,US!). Known from Apache, Cochise, Coconino, Gila, Maricopa, Mohave, Navajo,Pima, Pinal,SantaCruzandYavapaicos.Occasionalinchaparral, Madrean oak woodlands, ponderosapine zone, pinon-juniperzone, and rockygrasslands; 822—2438 m; floweringApril toSeptember. SomeherbariumspecimensaredifficulttoseparatefromE.sericeus.The speciesusuallymaybeseparatedbythespreading-villosetrichomesonthe sepalsofE. nuttallianus(SantaCruzCo.,Austin&Austin1512,ASU)and appressed-pilose trichomesonE. sericeusalthough thereare intermediate specimens{Metcalfe 1228, NMC, NY, US). Corollalimbsarepalebluechangingtowhitenearthebase; theshort tubeispaleyellowwithin. Androeciaandgynoeciaarewhite. Noinsect visitorswere seen. — 4. EvcM.viJLiis.sr.Ric:F.usSwartz,Prodr. Veg. Ind.Occ. 55. 1788. Typf: JAMAICA:Sin/rtz(M,S). li. wilcoxuini/5House, Bull. TorreyBot. Club33:315. 1906.—Typf: ARIZONA: Wilcox96(US!). Known from Cochise, Gila, Graham, Greenlee, Pima, Pinal, Navajo, SantaCruz andYavapai cos. Frequent inchaparral, Madrean oakwood- lands, and desertgrasslands; 975—1889m; flowering May toOctober. For thosewishing to recognizethem, twovarietieshavebeen named: var. discolor(Bcnth.) A. Gray, with upper leafsurfaceglabrous and var. sericeus, withleavessericeousonbothsurfaces. Sincetheseplantsmaybe foundgrowingtogetheroutsidetheU.S.A.,Idonotrecognizethedistinc- tion. Specimens that wereseparatedasE. oreophilusGreeneweretreated by Ooststroom(1934)asE.sericeusvar.discolorformB.Thesearebettertreated as E. nuttallianusbecauseoftheir habit, corollashapeand colorand in- dumentum on both leafsurfaces. The androecium and thegynoecium arewhite(CochiseCo. Austin & Austin 1511, ASU). No flowervisitorshavebeenseen. 449 6. IPOMOEA 1. Ipo-mof.abarbatisepalaa. Gray,Syn. Fl. N. Amer. 2, 1:212. 1886. Typi-:TEXAS:Wni^k507(GH!,US!). KnownfromCochise,Gila,Graham,Greenlee,Maricopa, Pima,Santa Cruz,andYavapaicos.OccasionalinmesquitegrasslandsandMadreanoak woodlands; 853~2438 m; floweringJuly toDecember. Flowers open atdawn, and areat first blueon the limbwithawhite throat.Theoutsideofthetubeiswhiteonthefolds(plicae)andpalegreen ontheunfoldedarea(interplicae). Assenescencebeings, thecorollaturns pink and then reddish. Anthers are white, but the filaments are pale yellow.Thestyleisgreen,thestigmawhiteand2-lobed,theovarygreen, and the disc yellow and cup-shaped (Pima Co., Austin &Austin 7394, ASU). 2. Ipomof.a c:apiu.acea (H.B.K.) G. Don, Gen. Syst. 4:267. 1838. Typf:COLOMBIA:Bonp/ani/(miCTofichd). I.muna/taOdv.,IconesPI.5;52.pi.478.f.2. 1794,nonL.(1763),nonjacq.(1789). Known from Cochise, Coconino, Pima, Santa Cruz and Yavapai cos. Occasional m Madrean oak woodlands, desertgrasslands, andponderosa pinezones; 1524—2499 m; floweringJuly toSeptember. Flowersopenatdawn. Thecorollalimbislavenderandthetubewhite, within and without. Androecia and gynoecia are also white. Beeflies (Bombyliidae)visittheflowers(CochiseCo.,Austin&Austin7569,ASU). 3. Ipomof.acardiophyllaA. Gray, Syn. Fl. N. Amer. 2, 1:213- 1886. Tyfi::TEXAS:Wnt^ht511(GH!). NoArizonaspecimenswereseenbyKearneyandPeebles(1951, I960) although theysuggested thisspecies' presence inArizonaasprobable. A populationwasreported nearTombstonebyMasonetal. (1986)andwas relocatedin 1989(CochiseCo.,Austin&Austin7608,ASU). Thespecies isrepresentedbyscatteredplantsextendingalongtheroadforabout0.8 milesatanaltitudeof1127 m in mcsquite-creosote bush scrub(Chihu- ahuandesertscrubsensuBrownandLowe1980).Nofruitswerepresenton 7 Sep 1989;somematurefruitswerefoundon26Sep, butwereheavily parasitized by insects. Rareand local. Flowersopenat6a.m.; mostlywiltedby 11 a.m. Thecorollatubeis paleyellowontheoutside;theHmbispaleblue(asm/,tricolor). Stamens arewhitetopalecream;thestyleisgreen,thestigmawhite. Noodorwas detectable, andnoinsectsvisitedbetween6:00and6:30a.m. McDonald (1982)found nopollinatorsonthespeciesinTexasandNewMexicoand 450 had a90% seed set oncultivated plants. Thespecies may beconsidered autogamous. — 4. IpomoI'A (x)s'iiiLAiA Toff., Bot. Mex. Bound. 149. 1859. Tvpi;: Ti;XAS: WV/t;/)/ 505(GH!,US!). — \.jiitiiisA. NcLson,Univ.WyomingFnbl.Sci. 1(3):65. 1921. Typi;:ARIZONA: UcifisiiN lOKi(RS, notseen;photoFAU!). Known from Apache, Cochise, Coconino, Gila, Greenlee, Mohave, Navajo, Pima, Final, SantaCruzandYavapaicos. Commoninchaparral, Madreanoakwoodlands,andponderosapinezones;975—213.^m;flower- ingJuly toOctober. Flowersbegintoopenat8:15a.m.andbegintowiltat 10:30a.m.The corollalimbislavender,thetubewhitewithinandwithout.Filamentsand anthersarewhite;whitetrichomcsextendalongthefilamentsfrombaseto apex; theovary iscream, andthestyleisgreen. Astigmawas3-lobedon one flower, 2-lobed on others. The flowers are visited by bumblebees (Cochise Co. Aiistni &Austin 75S4, ASU). 5. Ipomoi-a cRi.sriii.ATA H. Hallier, Med. Rijksherb.—Leiden 46:20. 1922.Anewname{orQuamoditgracilisH.Hallier. Typi::MEXICO: basedonsyntypesincludingBoiirgeau1061(G-DC!).Q/m///(h//!gracilisH.Hallier, Bull. Herb. Boiss. 7:116. 1899. /. ciianieiiaucrr., nonL. Known trom all Arizona counties except La Paz. Plants grow in chaparral,Madreanoakwoodlands,andponderosapinezones;731—2773 m; flowering May toNovember. This isprobablythemostcommonand widespread species inthestate. "Siakclok-i"{Blakeley&Marshall368, DES) Flowers,whichareopenfrom6:00a.m.intothelateafternoon,arevisi- tedbyhummingbirds.Sincethestigmaisheldbelowthelevelofstamens, the plants may be facultatively allogamous (Pima Co., Austin & Austin 7593, ASU). Accordingtoananonymousreviewerthespeciesisprobably predominantlyselhngandtheremaybecleistogamousflowersunderlow light conditions. Even when chasmogamous the anthers dehisce before anthesis. — 7. Ifc)M()i;a Hi;nikac:i;aJacq., Collect. Bot. 1:124. 1786. 'ISpi:based on Dillenius, Horr. Elth. t. 80, fig. 92(plate.selectedlectotype!byVerdcourt, 1957). — /.deierton/mHouse,Ann. NewYorkAcad.Sci. 18:203. 1908. Typi::ARIZONA: ThonikT29(ARIZ!, NY!). Although not reported for Arizona by Kearney and Peebles (1951, , 451 1960),certainspecimensidentifiedas"/.hinnttila"arcofthisspecies,the others are /. purpurea(Austin, 1990). Known from Cochise, Coconino, Gila, Graham, Maricopa, Pima, Pmal, SantaCruz, Yavapai andYumacos. Commoninvariousdisturbed sites, especially cotton fields; 883—1859 m; flowering August to November. Thecorollasopenatdawn,someclosingby8:15a.m.,andallarcclosed by 11 a.m. Honeybees were seen bypassing flowers after pausing. A bumblebeebypassedflowersof/. hederaceabutvisitedthoseof/, costellata and Convolvulusequitans. However, flowersof/. hederacea werevisited by short-tailedblackswallowtailbutterflies{Papilioindra). Corollalimbsare blue early afteropening but begin to include reddish pigments as they begintowiltandturnmorepurple. Thecorollatubeiswhitewithinand without. Thestamensand thestylesarewhite; theovaryiscream(Pima Co., Austin &Austin 7396, ASU). — 8. Iix)moi;a i.i;ptotoma Torr., Bot. Mex. Bound. 150. 1859. Typi.: MEXICO. SoNfiRA: T/mr/m-977(OH'). — /.leptotomavanwootoniiE.Kel.so,Rliodora39:151. 1937. Typi::ARIZONA:10Sep 1914, Wooton 10(US!). /. leptotomaTorr. f. wootonii(E. Kelso)Wiggins, Contr. DudleyHerb. 1:21. I9'3(). Known from Cochise, Gila, Graham, Pima, Pmal, Santa Cruz and Yavapai cos. Occasional in plains, Sonoran desert scrub; 609~ 1.^71 m; floweringJune toOctober. Thecorollaopensasthesunraystouchit,between7:30and8:00a.m. andclosebetween 10:30and 11:00a.m. Corollalimbsarelavender, and there isawhitezonebetweenthelimbandpaleyellowbaseofthetube. Thetubeiswhitewithout.Orangetrichomesadornfilamentsfrombaseto apex; the androecium, gynoecium, style, ovary, and disc are white. Skippers(Hesperiidae)visitflowers(GilaCo.,Austin&Austin7601,ASU, which is anew county record; PimaCo. Austin &Austin 7595, ASU). — 9. Ipc:)MOi:a X leucaniha Jacquin, Icones Rar. 2:t. 318. 1788. T^'pe:nospecimenknown;illustrarionchosenaslectotypebyAustinin 1978. /. trilolhisensLiauctt.,nonL. (17')3). /, Idi'/iuoscisensiiShinners(1965),nonL. (1733). Knownfrom Maricopa, PimaandYumacos. Plantsgrow indisturbed sites; ca. 701 m; flowering March toNovember. Known from three old collections (Santa Cruz Co., Pringle in 1884, ARIZ;PimaCo., Thornherin 1912, ARIZ;countyunknown, LeRoys.n., NY); one in 1945 (PimaCo. Goodding&Lusher 128-45, NY), and two recentones(MaricopaCo.4Oct 1979,Heathmans.n.,ARIZ,ASU;Yuma 452 Co.7Nov 1985,Tuttles.n.,ARIZ).Thehybridsareprobablynotasrareas collections seem to indicate since they areweeds incotton fields. 10. Ip{)mc)i:a LiNOHi-iMi-Ri A. Gray, Syn. Fl. N. Amer. 2, 1:210. 1886. Typi;:TEXAS: WV/^j/ 50«(GH!, US!). KnownfromCochiseCo. (Gleeson, 25Aug 1927,Thornbers.n.,ARIZ; Bisbee, 30 Sep. 1930, Thombers.n., ARIZ); reportedly from Pima Co. (KearneyandPeebles 1951;Kearneyetal. I960). PlantsgrowinMadrean oak woodlands, and Chihuahuan desert scrub zones; 1066—1371 m; flowering August toSeptember. Anextremelyrarespecies;itscontinuedexistenceinArizonaisproble- matical. Probablyrelatedtoandeasilyconfusedwith/.puhescensLam.,the twocollectionsfromArizonaaresomewhatintermediatebetweenthetwo on the basisofsepal shapeandpubescence. — 11. Ip()mc)i;a l()nc;ii()i,ia Benth., PI. Hartweg. 16. 1839- Tvi'i:: MEXICO:hhirlwe^(K). KnownfromCochiseandSantaCruzcos. LocallycommoninMadrean oak woodlands; 975— 1828 m; floweringJuly toSeptember The nectary is cream-colored, and theandroecium andgynoecium are white. Early in the evening the flowers have a slight sweet fragrance. Flowersarepollinatedbymoths(Austin 1986).Flowersopen3:00to4:00 p.m.andcloseneardawn;afewwerestillopenat7:30a.m.(CochiseCo., A//\tni &Austin 7382, ASU). 12. Ifomoi-a hlummirai-: A. Gray, Syn. Fl. N. Amer. 2, l:suppl. 434. 1886. Tvpi:ARIZONA:Lemmo//2H,V)(GH!). — /. cTiiyypmij;o:iuAiRAI.ZGOrNayA,:Flr.oec.immAnme2r8.39Ac(aFd!,.GAHrt!.,s 1U9S:!9)0.. 1883, nonMeisner(1869). /. fiinîiiHouse,Torreya6:124. 1906, noin. nov. for/. ciincijoliaA. Gray. Known from Apache, Cochise, Coconmo, Gila, Graham, Greenlee, Pima,SantaCruzandYavapaicos.Occasionalintheponderosapinezone; 1219—2743 m; flowering April toOctober. Thecorollasopen byca. 6:30a.m., areclosingat 10:30a.m. andare completelyclosedat 11:50a.m.Corollalimbsarelavender,thetubewhite withinandwithout.Theovaryandstylearegreen,thestigmaandstamens white(CochiseCo., Austin &Austin 75H1, ASU). — 13. Iix)M()i;a pubi;s<;i;ns Lam., Encycl. Meth. Bot. 1:265. 1791. ISpr: AMERICA:collectorunknown(K!). — /. IwliTophyt/aOrtega, Nov. Fl. Descr. Dec. 1:9. 1797. T^pi:: MEXICO: Horto Regio., 1797,Orlegî(iixtotyph:MA222592, phoroFAU!).