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Alaska Trees and Shrubs Agriculture Handbook (United States. Dept. of Agriculture) ; title: No. 410 author: Viereck, Leslie A.; Little, Elbert Luther publisher: University of Alaska Press isbn10 | asin: 0912006196 print isbn13: 9780912006192 ebook isbn13: 9780585207391 language: English Trees--Alaska--Identification, Shrubs--Alaska--Identification, Trees--Alaska-- subject Classification, Shrubs--Alaska--Classification. publication date: 1986 lcc: QK146.V54 1986eb ddc: 582.1609798 Trees--Alaska--Identification, Shrubs--Alaska--Identification, Trees--Alaska-- subject: Classification, Shrubs--Alaska--Classification. Page i Alaska Trees and Shrubs by Leslie A. Viereck, Principal Plant Ecologist Institute of Northern Forestry Pacific Northwest Forest and Range Experiment Station U.S.D.A. Forest Service, College, Alaska and Elbert L. Little, Jr., Chief Dendrologist Division of Timber Management Research U.S.D.A. Forest Service, Washington, D.C. Agriculture Handbook No. 410 Forest Service United States Department of Agriculture Washington, D.C. 1972 Reprinted by the University of Alaska Press Fairbanks, Alaska Page ii Library of Congress Catalog Card Number: 86-050727 International Standard Book Number: 0-912006-19-6 Printed in the United States of America. Forest Service, United States Department of Agriculture First Printing, Agriculture Handbook No. 410, 1972 University of Alaska Press Second Printing, 2,000 copies, 1986 Third Printing, 2,000 copies, 1988 Fourth Printing, 2,500 copies, 1991 Fifth Printing, 2,500 copies, 1994 Cover: The scratchboard cover design by William D. Berry is a typical white spruce paper birch stand in interior Alaska in early winter. The shrub layer is alder and willow. Page iii Contents List of species iv Introduction 1 Previous work 2 Preparation of this handbook 3 Plan 4 How to use this handbook 5 Acknowledgments 5 Statistical summary 6 Growth forms 6 Alaska trees 8 Geographic distribution 9 Local and rare species 10 Vegetation of Alaska 11 Coastal forests 13 1. Coastal spruce-hemlock forests 14 Interior forests 15 2. Closed spruce-hardwood forests 15 3. Open, low growing spruce forests 18 4. Treeless bogs 19 5. Shrub thickets 20 Tundra 21 6. Moist tundra 21 7. Wet tundra 22 8. Alpine tundra 22 Vegetation map 23 Keys for identification 24 Key to Alaska trees based mainly on leaves 25 Winter key to deciduous trees of Alaska 28 Key to genera of Alaska shrubs 30 Winter key to Alaska shrubs 34 Alaska trees and shrubs 43 Key to Alaska willows 78 Vegetative Key to Alaska willows 82 Selected references 254 Index of common and scientific names 260 Page iv List of Species 1 Yew family (Taxaceae) 1. Pacific yew, Taxus brevifolia Nutt. (LS-ST, C, R) 44 Pine family (Pinaceae) 2. lodgepole pine, Pinus contorta Dougl. (ST-LT, C) 45 3. tamarack, Larix laricina (Du Roi) K. Koch (ST-MT, I) 48 4. black spruce, Picea mariana (Mill.) B.S.P. (ST-MT, I) 51 5. *white spruce, Picea glauca (Moench) Voss (MT-LT, I-c) 52 6. *Sitka spruce, Picea sitchensis (Bong.) Carr. (LT, C) 54 7. *western hemlock, Tsuga heterophylla (Raf.) Sarg. (LT, C) 58 8. *mountain hemlock, Tsuga mertensiana (Bong.) Carr. (ST-LT, C) 59 9. Pacific silver fir, Abies amabilis (Dougl.) Forbes (MT-LT, C, R) 61 10. subalpine fir, Abies lasiocarpa (Hook.) Nutt. (ST-LT, C, R) 62 Cypress family (Cupressaceae) 11. *western redcedar, Thuja plicata Donn (LT, C) 64 12. *Alaska-cedar, Chamaecyparis nootkatensis (D. Don) Spach (MT-LT, 66 C) 13. common juniper, Juniperus communis L. (PS-SS, I-C) 68 14. creeping juniper, Juniperus horizontalis Moench (PS, I) 69 Willow family (Salicaceae) 15. *balsam poplar, Populus balsamifera L. (MT-LT, I-c) 72 16. *black cottonwood, Populus trichocarpa Torr. & Gray (LT, C) 74 17. *quaking aspen, Populus tremuloides Michx. (SM-MT, I) 76 18. netleaf willow, Salix reticulata L. (PS, I-C) 86 19. Setchell willow, Salix setchelliana Ball (PS, I-c) 87 20. polar willow, Salix polaris Wahlenb. ssp. pseudopolaris (Flod.) Hult. (PS, 88 I-c) 21. skeletonleaf willow, Salix phlebophylla Anderss. (PS, I) 89 22. least willow, Salix rotundifolia Trautv. (PS, I-C) 90 23. arctic willow, Salix arctica Pall. (PS, I-C) 91 1 Size is indicated by letters: LT, large tree; MT, medium tree; ST, small tree; LS, large shrub; MS, medium shrub; SS, small shrub; PS, prostrate shrub. General distribution is given as I, interior, and C, coastal, with small letter where restricted, and R, rare. The 10 tree species producing nearly all the commercial timber are indicated by an asterisk (*). Explanation under Statistical Summary, page 6. Page v 24. Alaska bog willow, Salix fuscescens Anderss. (PS, I-C) 92 25. ovalleaf willow, Salix ovalifolia Trautv. and Salix stolonifera Cov. (PS, 94 i-C) 26. grayleaf willow, Salix glauca L. (MS-ST, I-C) 95 27. barren-ground willow, Salix brachycarpa Nutt. ssp. niphoclada (Rydb.) 97 Argus (SS-MS, I) 28. halberd willow, Salix hastata L. (MS, I) 98 29. low blueberry willow, Salix myrtillifolia Anderss. (SS, I) 99 30. tall blueberry willow, Salix novae-angliae> Anderss. (LS, I) 101 31. Barclay willow, Salix barclayi Anderss. (MS-LS, I-C) 102 32. Hooker willow, Salix hookeriana Cov. (LS-ST, C, R) 106 33. undergreen willow, Salix commutata Bebb (MS, I-C) 106 34. Chamisso willow, Salix chamissonis Anderss. (PS, I) 108 35. park willow, Salix monticola Bebb. (MS-LS, I-c) 108 36. Richardson willow, Salix lanata L. ssp. richardsonii (Hook.) A. Skwortz. 110 (MS-LS, I-C) 37. Barratt willow, Salix barrattiana Hook. (SS, I) 110 38. feltleaf willow, Salix alaxensis (Anderss.) Cov. (LS-ST, I-C) 112 39. silver willow, Salix candida Fluegge (MS, I, R) 115 40. Bebb willow, Salix bebbiana Sarg. (LS-ST, I-C) 116 41. diamondleaf willow, Salix planifolia Pursh ssp. pulchra (Cham.) Argus 118 (PS-LS, I-C) 42. Scouler willow, Salix scouleriana Barratt (LS-MT, I-C) 120 43. Sitka willow, Salix sitchensis Sanson (LS-ST, I-C) 122 44. littletree willow, Salix arbusculoides Anderss. (LS-ST, I-c) 123 45. sandbar willow, Salix interior Rowlee (LS, I) 126 46. Pacific willow, Salix lasiandra Benth. (LS-ST, I-c) 126 Bayberry family (Myricaceae) 47. sweetgale, Myrica gale L. (SS-MS, I-C) 128 Birch family (Betulaceae) 48. dwarf arctic birch, Betula nana L. (SS-MS, I-C) 130 49. resin birch, Betula glandulosa Michx. (SS-MS, I-c) 130 50. *paper birch, Betula papyrifera Marsh. (ST-LT, I-C) 134 51. American green alder, lnus crispa (Ait.) Pursh (MS-LS, I) 140 52. Sitka alder, Alnus sinuata (Reg.) Rydb. (LS-ST, I-C) 142 53. red alder, Alnus rubra Bong. (ST-MT, C) 144 Page vi 54. thinleaf alder, Alnus tenuifolia Nutt. (LS-ST, I-C) 146 Mistletoe family (Loranthaceae) 55. hemlock dwarf-mistletoe, rceuthobium tsugense (Rosend.) G.N. Jones 146 (Parasite, C) Gooseberry family (Grossulariaceae) 56. swamp gooseberry, Ribes lacustre (Pers.) Poir. (MS, I-C) 150 57. stink currant, Ribes bracteosum Dougl. (MS-LS, C) 150 58. northern black currant, Ribes hudsonianum Richards. (PS-MS, I-C) 152 59. skunk currant, Ribes glandulosum Grauer (SS-MS, I) 154 60. trailing black currant, Ribes laxiflorum Pursh (PS-MS, C) 155 61. American red currant, Ribes triste Pall. (PS-MS, I-C) 156 Rose family (Rosaceae) 62. Pacific ninebark, Physocarpus capitatus (Pursh) Kuntze (MS-LS, C) 158 63. Beauverd spirea, Spiraea beauverdiana Schneid. (SS, I-C) 159 64. Douglas spirea, Spiraea douglasii Hook. (MS, C) 160 65. luetkea, Luetkea pectinata (Pursh) Kuntze (PS, i-C) 162 66. Oregon crab apple, Malus diversifolia (Bong.) Roem. (ST, C) 162 67. Greene mountain-ash, Sorbus scopulina Greene (MS-ST, I-C) 166 68. European mountain-ash, Sorbus aucuparia L. (ST-MT, C, introduced) 167 69. Sitka mountain-ash, Sorbus sitchensis Roem. (MS-ST, C) 168 70. Siberian mountain-ash, Sorbus sambucifolia (Cham. & Schlecht.) 169 Roem. (MS, Aleutian Is., R) 71. saskatoon serviceberry, Amelanchier alnifolia (Nutt.) Nutt. (MS, I, R) 171 72. Pacific serviceberry, Amelanchier florida Lindl. (MS-ST, C) 172 73. black hawthorn, Crataegus douglasii Lindl. (LS, c, R) 172 74. American red raspberry, Rubus idaeus L. var. strigosus (Michx.) 175 Maxim. (MS, I-C) 75. salmonberry, Rubus spectabilis Pursh (MS-LS, C) 176 76. western black raspberry, Rubus leucodermis Dougl. (MS, C, R) 177 77. western thimbleberry, Rubus parviflorus Nutt. (MS, C) 178 78. bush cinquefoil, Potentilla fruticosa L. (SS-MS, I-c) 180 79. Drummond mountain-avens, Dryas drummondii Richards. (PS, I-c) 182 Page vii 80. white mountain-avens, Dryas octopetala L. (PS, I-c) 184 81. entire-leaf mountain-avens, Dryas integrifolia Vahl (PS, I-c) 184 82. prickly rose, Rosa acicularis Lindl. (SS-MS, I-c) 187 83. Woods rose, Rosa woodsii Lindl. (SS-MS, I, R) 189 84. Nootka rose, Rosa nutkana Presl (MS-LS, C) 190 Maple family (Aceraceae) 85. Douglas maple, Acer glabrum Torr. var. douglasii (Hook.) Dipp. (MS- 192 ST, C) Elaeagnus family (Elaeagnaceae) 86. buffaloberry, Shepherdia canadensis (L.) Nutt. (MS, I-c) 194 87. silverberry, Elaeagnus commutata Bernh. (MS-LS, I) 196 Ginseng family (Araliaceae) 88. devilsclub, Oplopanax horridus (Sm.) Miq. (MS-LS, i-C) 197 Dogwood family (Cornaceae) 89. red-osier dogwood, Cornus stolonifera Michx. (MS-LS, I-C) 199 Crowberry family (Empetraceae) 90. crowberry, Empetrum nigrum L. (PS, I-C) 201 Pyrola family (Pyrolaceae) 91. pipsissewa, Chimaphila umbellata (L.) Barton (PS, C, R) 204 Heath family (Ericaceae) 92. copperbush, Cladothamnus pyrolaeflorus Bong. (MS, C) 204 93. narrow-leaf Labrador-tea, Ledum decumbens (Ait.) Lodd. (SS, I-C) 206 94. Labrador-tea, Ledum groenlandicum Oeder (MS, I-C) 207 95. Lapland rosebay, Rhododendron lapponicum (L.) Wahlenb. (PS-SS, I) 209 96. Kamchatka rhododendron, Rhododendron camtschaticumPall. (PS, C) 210 97. rusty menziesia, Menziesia ferruginea Sm. (LS, i-C) 212 98. alpine-azalea, Loiseleuria procumbens (L.) Desv. (PS, I-C) 212 99. bog kalmia,Kalmia polifolia Wang. (SS, C) 214 100. red mountain-heath, Phyllodoce empetriformis (Sm.) D. Don (PS, c, 215 R) 101. blue mountain-heath, Phyllodoce coerulea (L.) Bab. (PS, I, R) 216 102. Aleutian mountain-heath, Phyllodoce aleutica (Spreng.) Heller (PS, i- 217 C) 103. four-angled cassiope, Cassiope tetragona (L.) D. Don (PS, I-c) 219 Page viii 104. Mertens cassiope, Cassiope mertensiana (Bong.) D. Don (PS, C) 220 105. starry cassiope, Cassiope stelleriana (Pall.) DC. (PS, I-C) 220 106. Alaska cassiope, Cassiope lycopodioides (Pall.) D. Don (PS, C) 222 107. bog-rosemary, Andromeda polifolia L. (PS-MS, I-C) 223 108. leatherleaf, Chamaedaphne calyculata (L.) Moench (PS-SS, I) 224 109. salal, Gaultheria shallon Pursh (MS, C) 226 110. Miquel wintergreen, Gaultheria miqueliana Takeda (PS-SS, Aleutian 226 Is., R) 111. bearberry, Arctostaphylos uva-ursi (L.) Spreng. (PS, I-C) 230 112. alpine bearberry, Arctostaphylos alpina (L.) Spreng. (PS, I-c) 230 113. red-fruit bearberry, Arctostaphylos rubra (Rehd. & Wilson) Fern. (PS, 231 I-c) 114. mountain-cranberry, Vaccinium vitis-idaea L. (PS, I-C) 233 115. dwarf blueberry, Vaccinium caespitosum Michx. (PS-SS, i-C) 234 116. red huckleberry, Vaccinium parvifolium Sm. (MS-LS, C) 236 117. early blueberry, Vaccinium ovalifolium Sm. (MS, i-C) 236 118. Alaska blueberry, Vaccinium alaskaense Howell (MS, C) 238 119. bog blueberry, Vaccinium uliginosum L. (PS-SS, I-C) 240 120. bog cranberry, Vaccinium oxycoccos L. (PS, I-C) 240 Diapensia family (Diapensiaceae) 121. diapensia, Diapensia lapponica L. (PS, I-C) 242 Honeysuckle family (Caprifoliaceae) 122. Pacific red elder, Sambucus callicarpa Greene (LS-ST, C) 243 123. high bushcranberry, Viburnum edule (Michx.) Raf. (MS-LS, I-C) 244 124. snowberry, Symphoricarpos albus (L.) Blake (SS-MS, C, R) 246 125. twin-flower, Linnaea borealis L. (PS, I-C) 248 126. bearberry honeysuckle, Lonicera involucrata (Richards.) Banks (MS- 250 LS, C, R) Composite family (Compositae) 127. fringed sagebrush, Artemisia frigida Willd. (SS, I) 252 128. Alaska sagebrush, Artemisia alaskana Rydb. (SS, I) 252

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