Table Of Content

Journal of Applied Ecology Volume 43 2006 EDITORS PS. Oilier P. Stephens R.P. Freckleton D. McCracken RE. Hulme S.J. Thirgood MANAGING EDITOR Gillian Kerby ASSOCIATE EDITORS J.S. Brashares F. He J.S. Richardson Y. Buckley M. Hebblewhite J.A. Royle M. Cadotte K.U. Karanth A. Sheppard J. Cossentine D. Kleijn I. Steffan-Dewenter C.R. Dickman J.J. Lennon D. Thompson C.S. Elphick J. Leps RM. Thompson A. Erhardt E.J. Milner-Gulland W.H. van der Putten M. Frederiksen S.J. Milton M.J. Walpole C.L.J. Frid W. Montevecchi M. Whittingham J.A. Gill A. Punt C. Wilcox R. Hails R.F. Pywell J. D. Wilson Published for the British Ecological Society by Blackwell Publishing Ltd OXFORD EDINBURGH BOSTON MELBOURNE BERLIN TOKYO This journal is covered by Current Contents, BIOBASE/ vided the appropriate fee is paid directly to the RRO. This Current Awareness in Biological Sciences, Cambridge consent does not extend to other kinds of copying such Scientific Abstracts. Chemical Abstracts, Geo Abstracts, as copying for general distribution, for advertising or GEOBASE and Science Citation Index promotional purposes, for creating new collective works or for resale. OnLineOpen articles are made available in Journal compilation © 2006 British Ecological Society. accordance with the terms of the Creative Commons All rights reserved. No part of this publication may be Deed, Attribution 2.0 (further details available from reproduced, stored or transmitted in any form or by any www.creativecommons.org), which allows Open Access means without the prior permission in writing from the dissemination of the article, but does not permit com¬ copyright holder. Authorization to photocopy items for mercial exploitation or the creation of derivative works. internal and personal use is granted by the copyright Special requests should be addressed to Blackwell Publishing holder for libraries and other users registered with their at: Journalsrights(goxon.blackwellpublishing.com. local Reproduction Rights Organisation (RRO), e.g. Copyright Clearance Center (CCC), 222 Rosewood Drive. Typeset by Graphicraft Limited, Hong Kong Danvers. MA 01923, USA (www.copyright.eom), pro¬ Printed in Singapore by KHL Printing Co Pte Ltd. iii Contents Volume Contents N'olume 43, Issue 1, February 2006 Priority contribution 1 R. WOODROFFE. C. A. DONNELLY, D. R. COX, F. J. BOURNE, C. L. CHEESEMAN, R. J. DELAHAY, G. GETTINBY, J. P. MCINERNEY & w. 1. MORRISON Effects of culling on badger Meles meles spatial organization: implications for the control of bovine tuberculosis Environmental change and fragmentation 11 N. BROWN, s. BHAGWAT & s. WATKINSON Macrofungal diversity in fragmented and disturbed forests of the Western Ghats of India 18 N. p. CHACOFF & M. A. AiZEN Edge effects on flower-visiting insects in grapefruit plantations bordering premontane subtropical forest 28 G. MOREAU, E. s. EVELEIGH, c. J. LUCAROTTi & D. T. QUIRING Stage-specific responses to ecosystem alteration in an eruptive herbivorous insect 35 R. ESCHEN, H. MULLER-SCHARER & u. SCHAEFNER Soil carbon addition affects plant growth in a species-specific way Aquatic habitats 43 p. E. HULME & E. T. BREMNER Assessing the impact of Impatiens glandidifera on riparian habitats: partitioning diversity components following species removal 51 F. VAN DE MEUTTER, R. STOKS & L. DE MEESTER Rapid response of macroinvertebrates to drainage management of shallow connected lakes 61 A. GARDMARK. N. JONZEN & M. MANGEL Density-dependent body growth reduces the potential of marine reserves to enhance yields 70 D. PONT, B. HUGUENY, U. BEIER, D. GOFFAUX, A. MELC HER, R. NOBLE, C. ROGERS, N. ROSET & S. SC’HMUTZ Assessing river biotic condition at a continental scale: a European approach using functional metrics and fish assemblages Avian ecology 81 J. D. HART, T. P. MILSOM, G. FISHER, V. WILKINS. S. J. MOREBY, A. W. A. MURRAY P. A. ROBERTSON The relationship between yellowhammer breeding performance, arthropod abundance and insecticide applications on arable farmland 92 M. KLAASSEN, s. BAUER, J. MADSEN & 1. TOMBRE Modelling behavioural and fitness consequences of disturbance for geese along their spring flyway 101 M. KERY & H. SCHMID Estimating species richness: calibrating a large avian monitoring programme 111 R. A. RONCONi & c. CASSADY ST. CLAIR Efficacy of a radar-activated on-demand system for deterring waterfowl from oil sands tailings ponds .Agricultural sy stems 120 E. KNOP. D. KLEIJN, F. HERZOG & B. SCHMID Effectiveness of the Swiss agri-environment scheme in promoting biodiversity 128 s. PEARCE & M. p. ZALUCKi Do predators aggregate in response to pest density in agroecosystems? Assessing within-field spatial patterns 141 E. K. KLEIN, c. LAViGNE, H. PICAULT. M. RENARD & P.-H. GOUYON Pollen dispersal of oilseed rape: estimation of the dispersal function and effects of field dimension Biological crusts 152 .M. A. BOWKER. J. BELNAP, D. w. DAVIDSON & H. GOLDSTEIN Correlates of biological soil crust abundance across a continuum of spatial scales: support for a hierarchical conceptual model Statistical methods 164 H. K. PREiSLER, A. A. AGER & M. J. WISDOM Statistical methods for analysing responses of wildlife to human disturbance 173 G. s. BROWN, w. J. RETTIE & F. F. MALLORY Application of a variance decomposition method to compare satellite and aerial inventory data: a tool for evaluating wildlife-habitat relationships 185 A. R. SOLOW & w. K. SMITH Using Markov chain successional models backwards 189 N. o'dea, r. j. WHITTAKER & KARL 1. UGLAND Using Spatial heterogeneity to extrapolate species richness: a new method tested on Ecuadorian cloud forest birds 199 Editors' Note (List of Referees) 204 Instructions to Authors \6lume 43, Issue 2, .April 2006 Habitat connectivity 209 p. F. DONALD & A. D. EVANS Habitat connectivity and matrix restoration: the wider implications of agri-environment schemes 219 G. s. HAMILTON, p. B. MATHER & J. c. WILSON Habitat heterogeneity influences connectivity in a spatially structured pest population .Agricultural sy stems 227 B. KRUK, p. INSAUSTI, A. RAZUL & R. BENECH-ARNOLD Light and thermal environments as modified by a wheat crop: effects on weed seed germination 237 M. D. THORSTED, J. WEINER & J. E. OLESEN Above- and below-ground competition between intercropped winter wheat Triticum aestivum and white clover Trifolium repens 246 R. H. GIBSON, I. L. NELSON, G. w. HOPKINS. B. i. HAMLETT & J. MEMMOTT Pollinator webs. plant communities and the conservation of rare plants: arable weeds as a case study iv Invasive species Volume 258 s. R. WANGEN & c. R. WEBSTER Potential for multiple lag phases during biotic invasions: reconstructing an invasion of the exotic tree Acer ptalanoicles Contents 269 M. J. HANSEN & s. 1). WILSON Is management of an invasive grass Agropyron cristatum contingent on environmental variation? V'egetation dynamics 281 R. A. BRAOSTOCK. M. BEDWARD & J. s. COHN The modelled effects of differing fire management strategies on the conifer Callitris verrucosa within semi-arid mallee vegetation in Australia 293 s. w. TODD Gradients in vegetation cover, structure and species richness of Nama-Karoo shrublands in relation to distance from livestock watering points 305 c. SMIT, J. DEN OUDEN & H. MUELER-SCHARER Unpalatable plants facilitate tree sapling survival in wooded pastures 313 M. PEEIFER, K. wiEGAND, w. HEINRICH & G. JETSCHKE Long-term demographic fluctuations in an orchid species driven by weather: implications for conservation planning Environmental impact on invertebrates 325 H. L. NOEL, s. p. HOPKIN, T. H. HUTCHINSON, T. D. WILLIAMS & R. M. siBLY Towards a population ecology of stressed environments: the effects of zinc on the springtail Folsomia Candida 333 K. JOHST, M. DRECHSLER, J. THOMAS & J. SETTELE Influence of mowing on the persistence of two endangered large blue butterfly species 343 R. p. PRASAD & w. E. SNYDER Polyphagy complicates conservation biological control that targets generalist predators Vertebrate management 353 p. DELATTRE, R. CLARAC, J. p, MELis, D. R. J. PLEYDELL & p. GIRAUDOUX How moles Contribute to colonization success of water voles in grassland: implications for control 360 F. J. L. KRAAIJEVELD-SMIT, R. A. GRii EITHS, R. D. MOORE & T. J. c. BEEBEE Captive breeding and the fitness of reintroduced species: a test of the responses to predators in a threatened amphibian Sampling populations . 366 H. B. RASMUSSEN, G. wiTTEMYER & 1. DOUGLAS-HAMILTON Predicting timc-spccific changes in demographic processes using remote-sensing data 377 s. T. BUCKLAND, R. w. SUMMERS, D. L. BORCHERS & L. THOMAS Point transect sampling with traps or lures Volume 43, Issue 3, June 2006 Special Profile: Making better biogeographical predictions of species' distributions 385 Editor’s Note 386 A. GUISAN, A. LEHMANN, S. EERRIER. M. AUSTIN, J. M. C. OVERTON, R. ASPINALE & T. HASTIE Guest Editorial: Making better biogeographical predictions of species’ distributions 393 s. EERRIER & A. GUISAN Spatial modelling of biodiversity at the community level 405 J. L. PEARCE & M. s. BOYCE Modelling distribution and abundance with presence-only data 413 s. BARRY & J. ELiTH Error and uncertainty in habitat models 424 J. M. DRAKE, c. RANDIN & A. GUISAN Modelling ecological niches with support vector machines 433 P. SEGURADO, M. B. ARAUJO & w. E. KUNIN Consequences of spatial autocorrelation for niche-based models Climate change and'fisheries 445 A. RINDORE & p. LEWY Warm, windy winters drive cod north and homing of spawners keeps them there Wetland ecology 454 J. SMART, J. A. GILL, w. J. SUTHERLAND & A. R. WATKiNSON Grassland-breeding waders: identifying key habitat requirements for management 464 N. H. K. BURTON, M. M. REHETSCH, N. A. cIj\rk & s. G. DODD Impacts of sudden winter habitat loss on the body condition and survival of redshank Tringa totanus 474 w. RECKENDOREER, c. BARANYi, A. FUNK & E. SCHIEMER Floodplain restoration by reinforcing hydrological connectivity: expected effects on aquatic mollusc communities Grazing management 485 R. D. JACKSON & B. ALLEN-DIAZ Spring-fed wetland and riparian plant communities respond differently to altered grazing intensity 499 D. M. EVANS, s. M. REDPATH, D. A. ELSTON, s. A. EVANS, R. J. MITCHELL & p. DENNIS To graze Or not to graze? Sheep, voles, forestry and nature conservation in the British uplands Modelling and monitoring 506 E. VAN DEN BOSCH, G. AKUDiBiLAH, s. SEAL & M. JEGER Host resistance and the evolutionary response of plant viruses 517 K. SHEA, A. SHEPPARD & T. wooDBURN Seasonal life-history models for the integrated management of the invasive weed nodding thistle Cardans nutans in Australia 527 R. M. EWERS & R. K. DIDHAM Continuous response functions for quantifying the strength of edge effects 537 M. SIMS, s. WANLESS, M. p. HARRIS, p. IAN MITCHELL & D. A. ELSTON Evaluating the power of monitoring plot designs for detecting long-term trends in the numbers of common guillemots Population viability and management 547 M. BEGUM, G. M. GURR, s. D. WRATTEN, p. R. HEDBERG & H. I. NicoL Using Selective food plants to maximize biological control of vineyard pests 555 F. RASCHE, H. VELVis. c. ZACHOw, G. BERG, J. D. VAN ELSAS & A. SESSiTSCH Impact of transgenic potatoes expressing anti-bacterial agents on bacterial endophytes is comparable with the effects of plant genotype, soil type and pathogen infection 567 M. A. c. NICOLE, c. G. JONES & K. NORRIS The impact of harvesting on a formerly endangered tropical bird: insights from life-history theory V 576 M. LINKIE, G. CHAPRON, D. j. MARTYR. J. HOLDEN & N. LEADER-WILLIAMS Assessing the viability of tiger subpopulations Volume in a fragmented landscape 587 c. RODRIGUEZ, K. JOHST & J. BUSTAMANTE How do Crop types influence breeding success in lesser kestrels through prey Contents quality and availability? A modelling approach 598 Editorial Announcement Volume 43, Issue 4, August 2006 The Tansley Lecture 599 w. J. SUTHERLAND Predicting the ecological consequences of environmental change: a review of the methods Forum 617 W. J. SUTHERLAND. S. ARMSTRONG-BROWN, P. R. ARMSWORTH. T. BRERETON, J. BRICKLAND. C. D. CAMPBELL. D. E. CHAMBERLAIN, A. 1. COOKE, N. K. DULVY. N. R. DUSIC, M. LITTON. R. P. ERECKLETON. H. C. J. GODERAY. N. GROUT. H. J. HARVEY. C. HEDLEY, J. J. HOPKINS. N. B. KIET, J. KIRBY, W. E. KUNTN, D. W. MACDONALD, B. MARKER. M. NAURA, A. R. NEALE. T. OLIVER, D. OSBORN, A. S. PULLIN, M. E. A. SHARDLOW, D. A. SHOWLER. P. L. SMITH. R. J. SMITHERS, J.-L. SOLANDT. J. SPENCER. C. J. SPRAY, C. D. THOMAS, J. THOMPSON, S. E. WEBB. D. W. YALDEN & A. R. WATKINSON The identification of 100 ecological questions of high policy relevance in the UK Birds and resource distribution 628 G. M. SIRIWARDENA. N. A. CALBRADE, J. A. VICKERY & w. J. SUTHERLAND The effect of the spatial distribution of winter seed food resources on their use by farmland birds 640 M. J. WHiTTiNGHAM. c. L. DEVEREUX. A. D. EVANS & R. B. BRADBURY Altering perceived predation risk and food availability: management prescriptions to benefit farmland birds on stubble fields 651 E. DE LA MONTANA, J. M. REY-BENAYAS & L. M. CARRASCAL Response of bird communities to silvicultural thinning of Mediterranean maquis Fisheries management 660 A. FENTON, T. HAKALAHTi, M. BANDILLA & E. T. VALTONEN The impact of variable hatching rates on parasite control: a model of an aquatic ectoparasite in a Finnish fish farm 669 M. M. HANSEN, D. BEKKEVOLD. L. F. JENSEN, K.-L. D. MENSBERG & E. E. NIELSEN Genetic restoration of a Stocked brown trout Salmo tnitta population using mrcrosatellite DNA analysis of historical and contemporary samples Predicting species' responses to management 680 R. p. BLACKSHAW & R. s. VERNON Spatiotemporal stability of two beetle populations in non-farmed habitats in an agricultural landscape 690 J. LOMAN & B. LARDNER Does pond quality limit frogs Rana analis and Rana temporaria in agricultural landscapes? A field experiment 701 D. HAWLENA & A. BOUSKILA Land management practices for combating desertification cause species replacement of desert lizards 710 E. HUNTER, F. BERRY, A. A. BUCKLEY, c. STEWART & J. D. METCALFT Seasonal migration of thornback rays and implications for closure management 721 J. M. MILNER, C. BONENFANT, A. MYSTERUD, J.-M. GAILLARD, S. CSANYI & N. CH. STENSETH Temporal and Spatial development of red deer harvesting in Europe: biological and cultural factors influence of environmental factors 735 D. BONTE, L. LENS & J.-P. MAELFAIT Sand dynamics in coastal dune landscapes con.strain diversity and life-history characteristics of spiders 748 N. GOVENDER, w. s. VC. TROLLOPE & B. w. van wilgen The effect of fire season, fire frequency, rainfall and management on fire intensity in savanna vegetation in South Africa 759 w. HARDTLE. M. NTEMEYER. T. NIEMEYER. T. ASSMANN & s. FOTTNER Can management Compensate for atmospheric nutrient deposition in heathland ecosystems? 770 A. J. ELMORE, s. J. MANNING, J. F. MUSTARD & J. M. CRAINE Decline in alkali meadow vegetation cover in California: the effects of groundwater extraction and drought 780 B. G. MCKIE. z. PETRIN & B. MALMQViST Mitigation or disturbance? Effects of liming on macroinvertebrate assemblage structure and leaf-litter decomposition in the humic streams of northern Sweden Diversity and distributions 792 s. CHANDY, D. J. GIBSON & p. A. ROBERTSON Additive partitioning of diversity across hierarchical spatial scales in a forested landscape 802 M. LUTOLF, F. KIENAST & A. GUISAN fhe ghost of past species occurrence: improving species distribution models for presence-only data Ecological methods 816 M. LEiRA, p. JORDAN, D. TAYLOR, c. DALTON. H. BENNTON, N. ROSE & K. IRVINE Assessing the ecological status of candidate reference lakes in Ireland using palaeolimnology 828 H. L. HOOPER. R. CONNON, A. CALLAGHAN, S. J. MAUND, M. LIESS, S. DUQUESNE, T. H. HUTCHINSON, J. MOGGS & R. M. siBLY The use of image analysis to estimate population growth rate in Daphnia magna Volume 43, Issue 5, October 2006 Special Profile: Biological Invasions Editor's Perspective 835 P. E. HULME Beyond control: wider implications for the management of biological invasions Review 848 Y. M. BUCKLEY, S. ANDERSON, C. P. CATTERALL, R. T. CORLETT. T. ENGEL, C. R. GOSPER. R. NATHAN. D. M. RICHARDSON. M. SETTER, o. SPIEGEL, G. VTVIAN-SMITH. F. A. VOIGT, J. E. s. WEIR & D. A. WESTCOTT Management of plant invasions mediated by frugivore interactions Special Profile: Biological Invasions (cont) s. P. WORNER & M. GEVREY Modelling global insect pest species assemblages to determine risk of invasion A. KULMATiSKi. K. H. BEARD & J. M. STARK Soil history as a primary control on plant invasion in abandoned agricultural fields E. JONGEJANS. A. w. SHEPPARD & K. SHEA What Controls the population dynamics of the invasive thistle Carduus nutans in its native range? E. K. DODSON & c. E. EIEDLER Impacts of restoration treatments on alien plant invasion in Piniis ponderosa forests. Montana. USA R. E. o'dell & V. p. CLAASSEN Relative performance of native and exotic grass species in response to amendment of drastically disturbed serpentine substrates K. E. STOKES & s. A. CUNNINGHAM Predictors of recruitment for willows invading riparian environments in south-east Australia: implications for weed management s. L. HALPERN & N. UNDERWOOD Approaches for testing herbivore effeets on plant population dynamies F. LEPRIEUR. M. A. HICKEY, c. J. ARBUCKLE. G. p. CLOSS. s. BROSSE & c. R. TOWNSEND Hydrologieal disturbance benefits a native fish at the expense of an exotic fish T. c. INGS. N. L. WARD & L. CHITTKA Can commercially imported bumble bees out-compete their native conspecifics? Priority Contribution 949 R. E. GREEN. M. A. TAGGART, D. DAS. D. J. PAIN, C. S. KUMAR, A. A. CUNNINGHAM & R. CUTHBERT Collapse of Asian vulture populations: risk of mortality from residues of the veterinary drug diclofenac in carcasses of treated cattle Ecosystems and management 957 c. LOEHLE Control theory and the management of ecosystems 967 S. BENEDICK. J. K. HILL, N. MUSTAFFA. V. K. CHEY, M. MARYATI, J. B. SEARLE. M. SCHILTHUIZEN & K. C. HAMER Impacts of rain forest fragmentation on butterflies in northern Borneo: species richness, turnover and the value of small fragments Avian ecology 978 M. YASUE & p. DEARDEN The potential impact of tourism development on habitat availability and productivity of Malaysian plovers Charadrius peronii 990 G. BLANCO, J. A. LEMUS & J. GRANDE Faecal bacteria associated with different diets of wintering red kites: influence of livestock carcass dumps in microflora alteration and pathogen acquisition Vegetation and management 999 A. H. HEGGENSTALLER, F. D. MENALLED, M. LIEBMAN & p. R. WESTERMAN Seasonal patterns in post-dispersal seed predation of Ahutilon theophrasti and Setaria faberi in three cropping systems 1011 F. KIRCHNER. A. ROBERT & B. COLAS Modelling the dynamics of introduced populations in the narrow-endemic Centaurea cotymbosa: a demo-genetic integration Methodological insights 1022 R. INGER. G. D. RUXTON, J. NEWTON, K. COLHOUN, K. MACKIE. J. A. ROBINSON & s. BEARHOP Using daily ration models and stable isotope analysis to predict biomass depletion by herbivores 1031 H. WANNER. H. GU, B. HATTENDORE, D. GUNTHER & s. DORN Using the Stable isotope marker ^Ca to study dispersal and host-foraging activity in parasitoids 1040 B. ERAPPIER & M. J. DUCEY Line-intercept laboratory subsampling estimates periphyton biovolumes more efficiently than standard methods V'olume 43, Issue 6, December 2006 Predators and predation Priority contribution 1049 F. SERGIO. 1. NEWTON, L. MARCHESI & p. PEDRiNi Ecologically justified charisma: preservation of top predators delivers biodiversity conservation 1056 c. CARROLL & D. G. MiQUELLE Spatial viability analysis of Amur tiger Panthera tigris altaiea in the Russian Far East: the role of protected areas and landscape matrix in population persistence 1069 M. A. BEKETOV & M. LIESS The influence of predation on the chronic response of Artemia sp. populations to a toxicant 1075 L. PALMA, p. BEJA, M. PAIS & L. c. DA FONSECA Why do raptors take domestic prey? The case of Bonelli’s eagles and pigeons Ungulates and land management 1087 A. ZANNESE, N. MORELLET. C. TARGHETTA, A. COULON, S. FUSER. A. J. M. HEWISON & M. RAMANZIN Spatial Structure of roe deer populations: towards defining management units at a landscape scale 1098 L. M. MARTIN & B. J. wiLSEY Assessing grassland restoration success: relative roles of seed additions and native ungulate activities Land management and diversity 1110 J. WRETENBERG, A. LiNDSTROM, s. SVENSSON, T. THiERFELDER & T. PART Population trends of farmland birds in Sweden and England: similar trends but different patterns of agricultural intensification 1121 M. RUNDLOE & H. G. SMITH The effect of organic farming on butterfly diversity depends on landscape context 1128 S. M. SMART, R. H. MARRS. M. G. LE DUC, K. THOMPSON, R. G. H. BUNCE. L. G. FIRBANK & M. J. ROSSALL Spatial relationships between intensive land cover and residual plant species diversity in temperate farmed landscapes 1138 A. R. YALLOP, J. I. THACKER. G. THOMAS. M. STEPHENS, B. CLUTTERBUCK, T. BREWER & C. A. D. SANNIER The extent and intensity of management burning in the English uplands vii Forest management Volume 1149 s. H. ROXBURGH, s. w. WOOD, B. G. MACKEY, G. WOLDENDORP & p. GIBBONS Assessing the carbon sequestration potential of managed forests; a case study from temperate Australia Contents 1160 A. E. EYCOTT, A. R. WATKiNSON & p. M. DOLMAN Ecological patterns of plant diversity in a plantation forest managed by clearfelling 1172 L. CAYUELA, D. J. GOLICHER, J. M. R. BENAYAS, M. GONZALEZ-ESPINOSA & N. RAMIREZ-MARCIAL Fragmentation, disturbance and tree diversity conservation in tropical montane forests 1182 K. R. BLANCHE, J. A. LUDWIG & s. A. CUNNINGHAM Proximity to rainforest enhances pollination and fruit set in orchards 1188 T. RIJKERS, w. OGBAZGHi, M. WESSEL & E. BONGERS The effect of tapping for frankincense on sexual reproduction in Boswellia papyrifem Gene dispersal 1196 K. E. HAYTER & J. E. CRESSWELL The influence of pollinator abundance on the dynamics and efficiency of pollination in agricultural Brassica napiis: implications for landscape-scale gene dispersal 1203 A. N. CURETON, H. J. NEWBURY, A. F. RAYBOULD & B. V. FORD-LLOYD Genetic Structure and gene flow in wild beet populations: the potential influence of habitat on transgene spread and risk assessment Models and methods 1213 M. KILJUNEN, J. GREY, T. SINISALO, c. HARROD, H. IMMONEN & R. 1. JONES A revised model for lipid-normalizing 5'^C values from aquatic organisms, with implications for isotope mixing models 1223 o. ALLOUCHE, A. TSOAR & R. KADMON Assessing the accuracy of species distribution models: prevalence, kappa and the true skill statistic (TSS)