W.K. Parker's collection of Foraminifera in the British Museum (Natural History)

34 Pages·1992·15 MB·English

by  R L Hodgkinson

Bull. Br. Mus. nat. Hist. (Geol.)48 (2): 45-78 Issued26November 1992
W. K. Parker's collection of foraminifera in
Museum
the British (Natural History)
R.L. HODGKINSON
DepartmentofPalaeontology, British Museum (NaturalHistory), CromwellRoad, London SW75BD
CONTENTS
TheCollection 46
TheCatalogues 46
T.R. Jones'approachtoidentifications 47
Provenanceofmaterial 47
Generalremarks 47
H.B. Brady'suseofmanuscript names 48
Thenumberingsystem 48
Thetypesofnewspecies 48
Anomalina 48
Articulina 48
Discorbina 49
Hauerina 50
Hippocrepina 50
Lagena 50
Lituola 54
Miliolina 55
Nodosaria 55
Nonionina 55
Nubecularia 56
Orbitolina 56
Planorbulina 58
Planularia 58
Polymorphina 59
Polystomella 61
(
Pullenia 61
Pulvinulina 61
Rotalia 62
Sphaeroidina 63
Textularia 63
Triloculina 65
Trochammina 65
Truncatulina 68
Uvigerina 68
Valvulina 68
WilliamKitchenParker;ashortbiography 69
Acknowledgements 75
References 75
Systematicindex 76
Synopsis. William KitchenParker(1823-1890)wasassociatedwith H.B. Brady, W.B. Carpenter,T.R. Jonesand
W.C. Williamson, and amassed in his lifetime a considerable collection ofRecent and fossil foraminifera. It was
purchasedbytheBritishMuseum(NaturalHistory)in1892fromhisexecutors,theretobeidentifiedandcatalogued
byProfessorThomasRupertJones.ThisarticleprovidesinformationontheCataloguesandthenewspecieserected
byParkerandhisassociates. Lectotypeshave here beenselectedandillustrated forthe followingtaxa: Anomalina
coronata,Articulinamultilocularis,Hauerinaplicata, Hippocrepinaindivisa, Lagenacrenata, L. distoma, L. sulcata
var. distomapolita, L. sulcatavar. marginatasubvar. squamosomarginata, L. sulcatavar. tetragona, L. sulcatavar.
tubiferosquamosa, Lituola cassis, L. findens, L. nautiloidea var. soldanii, Polymorphina elegantissima P. frondi-
,
formis, P. regina, P. variata, Polystomella arctica, Pulvinulina concentrica, Rotalia beccarii var. annectens, R.
beccariivar. dentata, R. schroeterianaand Uvigerina (Sagrina) dimorpha. Several 'lectotypes'chosenelsewhere by
other authors, for Discorbina rimosa, D. turbo var. polystomelloides, Lagena sulcata var. striatopunctata and
Trochammina squamata var. gordialis, shown to be invalid because they were not syntypes, are all regarded as
neotypes. AbriefbiographyofParkerisgiven.
46 R.L. HODGKINSON
THE COLLECTION 1. 'Very often the figured specimens, though present in the
collection cannot be easily defined'.
2. 'Thecollectionwasthe basisofthe "NotesontheNomen-
William Kitchen Parker's sudden death on July 3rd, 1890set clature ofthe Foraminifera", the papers ofW.B. Carpen-
in train a series of events which resulted in his collection of ter in the Philosophical Transactions ofthe Royal Society
foraminifera beingdeposited in the British Museum (Natural of1856, 1859and 1860andtheIntroductiontotheStudyof
History), London. His executors attempted to realize as the Foraminifera, 1862'.
much money as possible from the estate, and his son W.N. 3. 'In naming the species, care has been taken to follow as
Parker,onadvicefromProfessorT.R. Jones,approachedthe closelyaspracticable the methodofnomenclature usedby
Trustees of the British Museum (Natural History) offering W.K. Parker. Modernnamesforsomegenera,speciesand
the collection of foraminifera for sale. The Director, W.H. varieties are adopted when of real importance, having
Flower, authorized the purchase, funded jointly by the been founded on knowledge obtained since Prof. Parker
Departments ofZoology and Geology (now Palaeontology), left off working on his collection'. (I do not know when
for£100. Thecollectioncametothe Museumin 1892andwas Parker 'ceased' to work on foraminifera in his lifetime, if
said to comprise about 2000 fossil and 1000 Recent slides, he ever did, but his last taxonomic paper containing new
most containing many genera and species. The number of species seems to have been published in 1872, when he
specimensthis added to the collection isdifficult to estimate, was 49 years old, eighteen years before his death in
but was considerable. T.R. Jones, a long term friend and 1890-R.L.H.).
collaborator of Parker, was keeping a wary eye on the 4. 'The Published Lists and the Catalogue do not always
specimens, forhe wrote to the Keeper ofZoology A.C.L.G. correspond since some specimens may have been lost,
Gunther that he would be ... 'glad to carry on the arrange- others not noticed in the published lists either on account
ment' (of the specimens). In June 1892 Jones met R. Kirk- of their not being regarded as notable varieties or, not
patrick, a curator at the Museum, to discuss the collection. being contained in that part of the collection which was
Thisled,onJune25th,tobothDrGuntherandtheKeeperof first studied when the lists were originally made. Some
Geology Dr A.S. Woodward jointly requesting authority to cannot be fully identified in the closed cells of the slide
employ Jones in 'naming, labelling and arranging the collec- since only one side orface ofthe foraminiferis visible'.
tion of Recent and fossil foraminifera purchased from the
executors of the late Professor W.K. Parker'. The Director AlthoughJonesmakesnocommentintheCataloguetothe
recommended andthe Trustees approved, but stipulatedthat Recent foraminifera, in the Fossil Catalogue he does write:
the payment for this work was not to exceed £80 (being Wote-The namesgiven in the Catalogue are to be taken, not
dsliivdiedsecduerqautaeldlwyobueltdwepernovtihdeetJwooneDsewpiatrhtm£e1n1t7s/)6dan(£d1e.8v8epr)y.1B0y0 aansysumoethetrhensaammeesawprpliitetseneqounalltyhetosltihdeesnatmheemsseglivveesn.in. .t'heI
the middle of 1894Jones seems to have completed his work, Recent Catalogue. Who wrote the names on the slides still
for records ofGeology Department payments cease on June remains to be discovered.
4th. The resultofhislaboursisawell-arrangedcollectionand Within the Recent Catalogue, the Introduction is followed
a manuscript catalogue for both the Recent and Fossil parts by a General Geographical List and each locality is given a
ofthecollection. Thiscollectionanditscataloguesarehoused numberin Roman numerals; forexample:
in the Micropalaeontology Section of the Department of North AtlanticOcean
Palaeontology, British Museum (Natural History), London. Britain
South Coast
Poole, DorsetVII
and the slides from those localities were also numbered (for
THE CATALOGUES
example Poole, Dorset VII.1 and 2 etc.). Each group was
then placedin boxesspeciallysupplied forthe purpose (PI. 6,
There are two catalogues, herein referred to simply as the fig. 5, p. 73).
Catalogue,FossilandRecentbeingspecifiedwhennecessary. In 1937, Edward Heron-Allen, who as an unpaid curator
bInosuindedthuentRilece19n3t7,volpousmseibtlhyereatistahneinidnisctaitgaitoinotnhaotfitEwdawsanrodt wMausseulomo,kimnagdeafatenrottehein tchoellCecattiaolnogoufe (fsoereamPIi.ni4f,efriag. a2t) tthhaet
Heron-Allen(Hodgkinson 1989);theFossilvolumemayhave he had removed the slidesfrom the boxes and arranged them
been boundearlier. Both catalogues are boundin blue, titled by their numbers in 58 drawers so that they could be more
alongthe spine in gold lettering, thus: easily examined. The only numbers which now need to be
known are those written by Jones in pencil on the reverse of
CATALOGUEOFFOSSILFORAMINIFERA EC. theslideandthe individualMuseum registernumber. Heron-
PARKERCOLLECTION T. RUPERTJONES Allen preserved the labelsfrom the boxes by gluing them on
INDEXTO BOXES I-XLV 1894 to sheets ofpaper and filing them in a loose leafring binder
labelledonthe spine 'ORIGINALLABELS from Boxes and
and
Special slides'. Additional labels concerned with the H.M.S.
CATALOGUEOFPARKERCOLLECTION Firebrand material are kept in the back ofthe Catalogue.
RUPERTJONES INDEXTO BOXES I-XXVII Following the General Geographical List is the List ofthe
Recent Foraminifera in the Parker Collection, which tells the
The Introduction to the Catalogue, in what appears to be species content ofeach slide from a major and a sub-headed
Jones' handwriting, is to be found in the front ofthe Recent geographical locality, forexample:
manuscriptandwasprobablywritten atthecompletionofthe 'A. BRITISH (North Atlantic Ocean) I. Southend on the
work. In itJones makes thefollowingcomments. Thames [6 slides, 4 good], see column 5 in Table VII
W.K. PARKERCOLLECTION OFFORAMINIFERA 47
p. 422ec Philos. Trans 1865 slide no. 1. . . no. 2. . . etc.'. PROVENANCE OF MATERIAL
(Foractual examplesofthissee PI. 4, figs 3, 4).
The Recent slides were catalogued (see below) between
1960-70 by Mr I.D.J. Burdett, a Scientific Assistant in the To those who are interested in the provenance of material,
DepartmentofZoology. theCataloguesprovideavaluablesourceandreferencetothe
The date of binding of the Fossil Catalogue is not known, cruises ofthe oceanographic vessels, which were well known
but the date ofits compilation is given on the spine as 1894. to the savants of Parker's time, but less so today. For the
The Introduction is not in Jones' handwriting and differs remaining samples, Jones sometimes gives the names of
slightlyfrom thatatthebeginningofthe Recent Catalogue in individual donors. Below, in alphabetical order, is a list of
that it mentions publications in the Quarterly Journal ofthe contributors.
Geological Society and The Geological Magazine and states Recent: H.M.S. Cyclops (Lieut. J. Dayman); H.M.S.
that'AGeologicalandGeographicalorderhasbeenfollowed Cyclops (Capt. W.J.S. Pullen); H.M.S. Firebrand (Com. J.
as nearly as circumstances permit'. 'The order ofthe species Dayman); H.M.S. Fly (Capt. F.P. Blackwood; Jukes collec-
in the several lists is approximately in accordance with the tion); W.E. Parry Expedition; H.M.S. Plumper (Capt. G.H.
zoologicalarrangementadoptedin H.B. Brady's"Challenger Richards); H.M.S. Porcupine 1862.
Report", 1873-1876', published in 1884. L. Barrett, W.B. Carpenter, H. Cristy, Mr Cummings,
Between 1960and 1970these Fossil slidesweretakenfrom G.M. Dawson, G. Dines, W.J. Hamilton, A. Hilton, Sir
theirboxesandplacedindrawersandthecompletecollection Everard Home, W. King, R. McAndrew, Signor Meneghini,
was catalogued (see below) by Miss H. Patterson, an Assis- J. Millar, F.W. Millett, Colonel Pelly, Capt. T.A.B. Spratt,
tant Scientific Officerin the ZoologyDepartment, with some Marchese C. Strozzi, Dr Sutherland, G.C. Wallich, J. Weth-
assistance from myself, then an Assistant Experimental erell, J. Wright, W.C. Williamson and S.P. Woodward.
Officer. Unfortunately, becausetheMuseum'smicropalaeon- For a discussion of the provenence ofspecimens from the
tological collection is arranged stratigraphically, by collec- HundeIslandsandtheirincorporationintotheParkerCollec-
tors, theFossilCollectioncouldnot be kepttogether, anditis tion, see under Textularia agglutinans var. biformis (p. 63).
thus more difficult to retrieve than the Recent Collection. Throughout the Catalogue Jones mis-spells Abrolhos as
The boxes have been kept, however, and all the labels are 'Abrohlos' (following mis-spelling in Brady, Parker & Jones
stillin place (see theexample on PI. 6, fig. 5). 1888).
The information on the entire collection is filed in the Fossil: D.J. Anstead, J.F. Blake, L. Barrett, J. Brown, Mr
retrieval system (indices to genera and species etc.) of the Burton, Cesare de Ancona, O. Fisher, DrHaidinger, M. von
then Protozoa Section, now Micropalaeontology Section, Hantken, R. Harkness, W. Harris, Colonel Henekin, H.B.
DepartmentofPalaeontology, B.M.(N.FL). Holl, J.W. Hulke, Captain Hutton, M. Jokoyama, T.R.
Jones, F. Karrer, A. Lennox, C. Lyell, Dr Meneghini, F.W.
Millett, Sir Roderick Murchison, Mr Nugent, S.P. Pratt, Dr
Prestwich, J. Purdue, A.W. Rowe, D. Sharpe, T.A.B.
T.R. JONES? APPROACH TO Spratt, J. Sinzow, H.C. Sorby, Dr Southby, J. Sowerby, R.
Stachey, Marchese C. Strozzi, J. Szabo, Mr Thomas, Dr
IDENTIFICATIONS Thomson, M.E. de Verneuil, Mr White, S.V. Wood and J.
Wright.
In the main, H.B. Brady'swork on the Challenger Foramin-
ifera provided the basic framework on which Jones, in his
Catalogue, based his revision ofParker's generic and specific GENERAL REMARKS
names. Parker & Jones (1865: 335) adhered 'as strictly as
possible to the plan of study laid down by Williamson 1858,
and Carpenter 1862' and in their own papers of 1859-1863 During the 1939-1945 World War, the specimens were
(The Nomenclature ofthe Foraminifera) and they identified stored, but in 1953 when the Museum was almost fully
on the 'near approach to identity' method. This probably recovered, A.R. Loeblich jr and H. Tappan examined the
encouraged their frequent descriptions of new species, sub- collection whilst preparingthe Treatise (1964). Over a period
species, varieties and subvarieties which seem to cause so of35 days, they sifted through all the type slides, and finally
muchconfusiontopresentdaytaxonomists. Itisalsopossible selected lectotypes for22species.
that Jones, 20 years later when he came to prepare the A century has now elapsed since Parker died and T.R.
Catalogue, was rather rushed and was unable to spend time Jonesstarted his Catalogue. Manyofthe genera and species,
identifying new, unfigured, varieties, which in his opinion both calcareous forms and those with agglutinated tests, are
and hindsight he might have considered junior synonyms of currently being restudied (Jones 1984, Bronnimann & Whit-
better known species. This is especially noticeable in the taker 1988, Gibson etal. 1991 and Kaminski et al., in press)
section dealing with the Orbitolina concava group. Some- and it now seems opportune to examine the complete collec-
timeshefailedtonoticesyntypesonslidesandthustheywere tion again and where possible update some information. I
not recorded in the Catalogue. It has been my policy to have tried, first, to find the syntypes, select lectotypes (with
carefully examine these slides for missing types, often with illustrations if necessary) of the remaining species and pro-
success. vide information on the type specimens; and secondly to list
thecollectors ofParker's material.
A third aim has been to prepare reference lists of the
publications used by T.R. Jones to identify genera, species,
varietiesetc. Asfaras I am aware, there were never any lists
accompanying the collection and the identifications were all
48 R.L. HODGKINSON
undertaken for the Catalogue by Jones himself from these (1984) forLagenasulcatavar. striatopunctata, Bolli, Loeblich
publications. Though not given here, these publication lists & Tappan (1957) for Pullenia obliquiloculata, Berggren &
are available for private study at the Museum, in unedited Kaminski (1990) for Trochammina squamata var. gordialis
form; the references given here (p. 75) are for the present andGibsonetal. (1991)for Textularia (Bigenerina) tubulifera
textonly. - have all previously selected what they called 'lectotypes'
from the Parker Collection from among specimens that were
not in fact syntypes, the original syntype series being lost or
H.B. BRADY'S USE OF MANUSCRIPT NAMES unrecognized. The selections are, however, tantamount to
designating neotypes as I here regard them. Unfortunately,
all so-called 'paralectotypes' of these species are without
Brady worked in close collaboration with Parker and Jones. statusin taxonomy.
Brady, whose manuscript on the Shetlands' Rhizopoda was
readinFebruary 1864, possibly attributedthe namesofthree Anomalina coronata Parker & Jones, 1857
species (Lagena distoma, Polystomella arctica and Pul- PI. 1, figs 1-3
vinulina concentrica) to Parker and Jones because they had
told him that they had already seen, named and illustrated Lectotype (here designated). From Norway, locality II {ex
them in amanuscript whichwas tobe read on 12th May 1864 1894:4:3:216), ZF4913.
(eventually published 1865). Since Parker & Jones' paper is Type reference. Parker& Jones 1857: 294; pi. 10, figs 15,
farlonger and more detailed than Brady's, itisreasonable to 16.
assumethat they had thought upthe names at a much earlier
date. Bradyacknowledgestheirhelpin thepreparationofhis Localities. Recentspecimenswererecordedfromlocalities
paper (1864: 466), so he was clearly prepared to concede I, II, IV, VII and VIII, along the Norwegian coast; material
authorship; in these cases, therefore, authorship is quoted as now only exists from localities II (XXVI), 1894:4:3:216; IV
'Parker& Jones, in Brady 1864'. (XXVI.2), 1894:4:3:219 and VIII (XXVI.2), 1894:4:3:227.
Therearenootherspecimensextantinthe ParkerCollection.
See pages 38, 39, 41, 113, 124ofJones' Catalogue.
THE NUMBERING SYSTEM Remarks. This species can still be assigned to the Anoma-
linidae ofCushman. Loeblich & Tappan (1988: 604) synony-
mize the family with the Alfredinidae ofSingh & Kalia, and
Register numbers reject Anomalina in favour of Epistomaroides Uchio, which
The British Museum (Natural History) (abbreviated in the would not, in my opinion, contain A. coronata since it does
not possess the sutural bars and supplementary chambers.
text to 'BM(NH)') register numbers for Recent material are Loeblich & Tappan's decision, however, has proved prema-
variously prefixed 1894:, 1963:, etc., and ZF. The prefix P
ture as an application to the Commission on Zoological
psiaglnaiefioenstoalfoogsysilS.ecAtlilonthoefstpheecDiempenasrtamreenhtooufsePdalianeotnhteolMoigcyr.o- Nomenclature,byHansen& Rogl (1980),forthesuppression
ofAnomalina(onwhichitdepends) hasnowbeenwithdrawn
(seeHaynes, 1990: 528). Barker(1960: pi. 97,figs 1,2) refers
Slide numbers the species to Paromalina Loeblich & Tappan, and indeed it
ThefossilandRecentslidesarealsonumbered (inadditionto has been figured by Kihle & L0faldli (undated circa 1975) as
the Museum Register number) with Jones' own numerical Paromalina, from the Norwegian shelfwhere it has a strati-
srRleoifdmeeraetnoncbneeusmfyeosrutaneldmsqw(uiXitcChkiIlnVy.)eaFkcoehryCetaxhtaeamllpoolcgaeuleiX,tCy,1aVnt.dh2et,hairisanbaiwlchlio(cw2hs) ttthhheee g(b1re9ae8pn8h:ipcl6a3rc7ae)n.dgeiMneodfDiiPosllceiiasn&toomcSaeclonitenttao(1AR9se7ac8e)n,noti.bnyPtahLeroioerbmlasitlcuihdnya&ohfaTsaDipssipcnaacn-e
nomalina in an area off Nova Scotia, concluded that the
slide.
species biserialis, coronata, semipunctata andjaponica inter-
grade; thuscoronata becomesasubjectivejuniorsynonymof
semipunctata (Bailey) with a stratigraphic range of Miocene
THE TYPES OF THE NEW SPECIES to Recent.
&
The following section provides information on the type Articulina multilocularis Brady, Parker Jones, 1888
specimens described by Parker, either alone or with col- PI. 1, figs 4, 7. Fig. 1.1
leagues (usuallyJones and Brady). Arrangementisalphabet- Lectotype (here designated). From Plumper station 2 (ex
ical, givingthe type reference, type locality, detailsofextant 1894:4:3:1033), ZF4914.
materialandanynecessary remarks. Itisunfortunate that, in
most cases, it has not been possible to isolate the original Type reference. Brady, Parker, & Jones 1888: 212; pi. 40,
figured specimens from the syntypicseries. Where lectotypes fig. 10.
have been chosen, however, those specimens named in the Type locality. Recent, fragments of nullipore, H.M.S.
Csealteacltoegdu.e,Noclosseusbtjetcotitvheelyoriidgeinntailfifeidgursepse,cihmaevnesalhwaavyes bbeeeenn P19l°u4m7p'eSr37s°t5a8t'ioWn, s2,ounindin3g1 wfiaththsoomsmeofvienre sAabnrd,olohfofscoBasatnko,f
used. Once a type specimen has been selected all other
Brazil.
originallycited localitiesbecome merely 'localities'.
Severalauthors-Loeblich&Tappan(1964)forDiscorbina Remarks. This species is not recorded in Jones' Catalogue
rirrtosa and D. turbo var. polystomelloid.es, R.W. Jones (p. 146), but I found a specimen in the Parker Collection
W.K. PARKERCOLLECTION OFFORAMINIFERA 49
(LXXXV.9) 1894:4:3:1033 from the Abrolhos Bank, in 31 Localities. Fossil, Tertiary: Grignon, France. Recent:
fathoms, which I have designated lectotype andfigured here. Australiancoral reefs.
coFnhreaommbsepthreecsitmtyehpanendwemasyscriflpoetcuitnoodnt.yitpTei,sheubnucotlreiaagrli,nlaoblwuitfnigpgousfrsoeirblsaeh,odwtehgsartemeoonrloeyf fRreommarthkes.oriTgihnearleloacraelitnioes.spMeycirmee-nesxalmiistneadtiinonthoefaClaltGarlioggnuoens
artistic licence, it could be argued that they are one and the and Australian reef slides failed to find any specimens. It is
sameandthat mylectotypeshould be moreproperlyreferred recorded in the Fossil Catalogue (pp. 92, 94) only from
'Hauteville' (Hautteville), Departement Manche, France
toasthe holotype.
Pearcy (1914) does not record this species from a sample (LXXIX.40, 43(?) P47227, P47230; also 73-75 see below),
gathered from the type area. collectedbyLyell, andin the RecentCatalogue (pp. 206, 207
and 218, XCIII 45-48 and C.8) 1894:4:3:1373-6 and 1517.
Discorbina biconcava Parker & Jones, in Carpenter L(Po4e1b6l7i0c)h,&aTsapyeptanun(f1i9g6u4r:edC,592a-n5d94')padreasliecgtnoattyepdesa''l(ePct4o1t6y6p9e)'
1862 from the non-topotypic locality at 'Hauteville', France
Lectotype. Designated by Loeblich & Tappan (1964), (LXXIX.73-75). The former should properly be a called the
ZF3646. 'neotype' (see p. 48), and is formally designated as such
herein to stabilize the nomenclature, as the species is type of
Type references. Carpenter et al. 1862: 202, text-fig. 32g thegenusEpistomariaGalloway. The twofigured 'paralecto-
(p. 201); also Parker&Jones, 1865: pi. 19, fig. lOa-c. types' (Loeblich & Tappan 1964: fig. 472.1, 2) are from a
suite of six specimens exchanged with the U.S. National
Typelocality. Recent, shoresand, Melbourne, Australia. MuseumofNatural History (Smithsonian Institution).
Remarks. Recorded in the Recent Catalogue, p. 219 This species is widely recorded from localities in the Paris
(CII.l), 1894:4:3:1520. A lectotype was designated, but not Basin by Le Calvez (1970: 153; pi. 32, figs 1-3).
figured, by Loeblich & Tappan (1964: C584). It (ZF3646),
andthe two paralectotypes (ZF3645) have beenre-registered Discorbinaturbo (d'Orbigny) var. globigerinoides
&
from aslide labelled 'Melbourne'. Parker Jones, 1865
BaDr.kerbi(c1o9n6c0a:vapi.is91t,hefigt.yp2e),srpeefceiresstohfe Pslpaencuileisnotiod.DeisscPoarrbr.- Lectotype. Designated by Loeblich & Tappan 1964,
inellaCushman & Martin. P41661. Grignon.
Type reference. Parker & Jones 1865: 385-6, 421; pi. 19,
Discorbina dimidiata Parker & Jones, in Carpenter fig. 7a-c.
1862
Localities. Fossil, from the Cretaceous (locality unspeci-
Lectotype. Designated by Loeblich & Tappan (1964), fied) and Eocene, Calcaire Grossier at Grignon, ParisBasin,
ZF3651. France.
Type references. Carpenter et al. 1862: 201, text-fig. 32b Remarks. It is the type speciesofNeocribrellaCushman. A
(p. 201); also Parker&Jones: 1865: pi. 19, fig. 9a-c. lectotype and paralectotypes were designated by Loeblich &
Tappan from Grignon (1964: C680), respectively registered
Typelocality. Recent, coastofMelbourne, Australia. P41661 and P41660 (probably both from LXXXIV.125 ex
Remarks. Recorded in Recent Catalogue, p. 219 (CII.2), P47723, asthisslide nolongercontainsspecimens);seeFossil
P1a8r9r4:4(:139:3125:21,22o9n) atoslibdee lvaebreyllecdo'mMmeolbnoursnoeu't.h IotfisSsyadindeyb,y LCaXtXaXlIoVgu.e9,6-p9.81)05r.egTishteerreedaPr4e76a9d4d-i6t.ioLnaelCaslpveeczim(e1n9s70:(f1r7o7m;
Australia. A lectotype (ZF3651) was designated by Loeblich pi. 42, figs2, 6) recordsits distribution in the Lutetian ofthe
& Tappan (1964: C580) but was unfigured by them; their ParisBasin.
figuredparalectotypes(1964: fig. 456.5-6), whichconstituted
part of an exchange with the BM(NH), are in the U.S. Discorbina turbo (d'Orbigny) var. polystomelloides
&
National Museum of Natural History (Smithsonian Institu- Parker Jones, 1865
tion). The remaining specimens are in the BM(NH) and Neotype. Theso-called 'lectotype'fromJukes'dredgingno.
regLiosetbelriecdhZ&F36T5a0p.pan (1964: 580)regardD. dimidiataastype 2,selectedbyLoeblich&Tappan 1964, isinfacttheneotype,
species of Lamellodiscorbis Bermudez, but later (1988: 559) ZF3603; see p. 48. The 'paralectotypes' (see below) have no
type status.
they placed this genus, without comment, into Trochulina
d'Orbigny. Type reference. Parker & Jones 1865: 421; pi. 19,
fig. 8a-c.
&
Discorbinarimosa Parker Jones, in Carpenter 1862 Typelocality. FromtheRecentcoralreef,Jukes'dredging
Neotype. The so-called 'lectotype,' selected by Loeblich & (Jukes' no. 2), August 6th 1844, 17 fathoms, white shelly
Tappan 1964, is in fact the neotype, P41670: see p. 48. The mud, Australia.
'paralectotypes' (see below) have no status as types. For Remarks. The only reference to this species (as Discorbina
localitysee under'Remarks' below. polystomelloides) occurs in the Recent Catalogue, p. 235. It
Type reference. Carpenteretal. 1862: 205. isnotfromthetypelocalitybutfromJukes'dredgingNo2, 14
fathoms (CX.20), probably from near (south of) Raine Islet.
Typefigure. In Parker&Jones 1865: pi. 19, fig. 6a-c. In the absence ofsyntypes, Loeblich & Tappan (1964: C594)
50 R.L. HODGKINSON
designated, but did not figure, a 'lectotype' (properly neo- G.M. Dawson from 16 fathoms only; there are none from
type, ZF3603) from this slide. As this species is the type of 18-20 fathoms. Were H.B. Brady's specimens (1884: pi. 26,
Epistomaroid.es Uchio, this specimen is formally designated figs 10-14) also supplied by Parker? Few of the nine speci-
neotype herein in order to stabilize the nomenclature; see mens are in good condition; the best show external annular
p. 48. rings without matching internal partitions. The agglutinated
Evidence that Anomalina and Epistomaroides are conge- wall is of the trochamminaceous type (J.E. Whittaker, per-
nericisput forward byHansen & Rogl (1980), who maintain sonalcommunication).
thatAnomalinapunctulata d'Orbigny (the typespeciesofthe H. indivisa (the type species ofHippocrepina) still lives in
former) may be a badly drawn Epistomaria punctata Said Canadianwaters (Schafer& Cole 1978: pi. 1, fig. 8).
(1949: 37), which they claim is an Epistomaroides. Although
accepted, rather prematurely, by Loeblich & Tappan (1988: Lagena crenata Parker & Jones, 1865
604), Hansen & Rogl's application to the ICZN for the PI. 1, figs 14, 15
suppression of Anomalina d'Orbigny and the placing of its
nameonthe listofinvalidzoologicalnames, andtheconcom- Lectotype (here designated). From Malaga (ex P47925),
itant application to place the genus name Epistomaroides P52783.
Uchio on the list of valid zoological names, have subse- Type reference. Parker & Jones 1865: 420; pi. 18, fig. 4a,
quently been withdrawn (see Haynes, 1990: 519, 528). The
b.
twogeneraremaindistinct.
Localities. Fossil, of Middle Tertiary age, Bordeaux,
Hauerinaplicata Parker & Jones, 1865 France and Malaga, Spain. Recent, from the Swan River
shore sand, Australia.
PI. 1, figs5, 6
Lectotype (here designated). From Jukes' dredging no. 1 Remarks. The only extant fossil syntypes are from the
{ex 1894:4:3:1686), ZF4915. Miocene (?)ofMalaga(Catalogue: 124, CX.8),P47925;one,
selected here as lectotype, is most like Parker & Jones'
Typereference. Parker&Jones 1865: 437; table 10. original figure (although the base is slightly damaged). The
Typefigure. InCarpenter 1862: pi. 6, fig. 35. omnuldy,rec6o-r7d ifnatthheomRseceinnt CtahtealoSgwuaen(p.R2i1v9e)r,,fr(oXmCIaXn.c4h)o,r
Typelocality. FromtheRecentcoralreef,Jukes'dredging 1894:4:3:1503, does not precisely match the details of the
(Jukes' no. 1), August 6th 1844, 17 fathoms, white shelly locality as originally given. The species is said to be still
mud, N.E. Australia. commonly found along the south-western coast of Australia
(Albani & Yassini 1989: 377).
Remarks. This species is not mentioned in the Catalogue,
but from its type locality there are 3 incomplete specimens Lagenadistoma Parker & Jones, in Brady 1864
which are conspecific, referred to Hauerina ornatissima Kar-
PI. 1, figs 12, 13
rer (originally described as Quinqueloculina) by Jones; they
are from the Jukes' no. 1 dredging (CIX.25), 1894:4:3:1686. Lectotype (here designated). From Norway (ex
Intheabsenceofagoodoriginaldescriptionofplicata, andas 1894:4:3:230), ZF4917.
tsph.e oonflyCadripaegnntoesirs(w1e862p:oss8e1s;spiis.f6r,omfigi.ts3s5y),notnhyem,diHfafeureernicneas Typereference. Brady 1864: 467; pi. 48, fig. 6.
between these two species could be easily overlooked when Localities. Sub-fossil, from the fens near Peterborough,
examined superficially. Karrer's Q. ornatissima is fossil, England. Recent, along the coast of Norway, the Shetland
however, so these specimens determined as such by Jones Islands (82 fathoms) and the coast of Northumberland,
couldwell beplicata,whichisonlyknownfrom the Recent, a England.
fact not considered by Brady (1884: 192). Since the best
specimen isso like Carpenter'sfigure, I have no hesitation in Remarks. This species was first cited, as Lagena laevis
designating it the lectotype. If the two species are subse- (Walker & Montagu) var. striata (Montagu), by Parker &
quently found to be conspecific the name plicata, which has Jones, 1857: pi. 11, fig. 24 from the Norway Recent (top
nottomyknowledge been usedrecently, would havepriority paragraph, p. 283), where it is still found (Kihle & L0faldli
over ornatissima. There are no Hauerina listed in Murray & [undated, circa 1975]). There are sub-fossil specimens from
Taplin's unpublished Catalogue (1983) of the Carpenter sandy clay alluvium, on the border of Cambridgeshire and
Collection. Huntingdonshire, Peterborough Fen, one mile from itswest-
ern boundary, (CXL.38-40), P48441-3, Catalogue p. 167.
Hippocrepinaindivisa Parker, in Dawson 1870 There is one specimen from Shetland (Catalogue p. 21,
PI. 1, figs8-10 XloXcIaIli.t3y),VII1I8,943:40:-32:01029faatnhdomosne(Csatpaelcoigmueen: f4r0,om41t,heXXNVoIr.w5a)y,
Lectotype(here designated). From Gaspe Bay, 16fathoms 1894:4:3:230. My lectotype from this last-named locality is
(ex 1894:4:3:812), ZF4916. most like the originalfiguredspecimen. The sub-fossil prepa-
rations are balsam strews and no specimens are visible on
Typereference. Dawson 1870: 177, text-fig. 2. P48441 (CXL.38) orP48442 (CLX.39).
Localities. Recent, from Gaspe Bay; 16 fathoms, also The authorship of distoma was cited by Brady, 1864 as
18-20fathoms; GulfofSt Lawrence, Canada. 'Parker and Jones MS' (see p. 48). R.W. Jones (1984: 125)
placed this species into his own subfamily the Phialineiinae
Remarks. Jones' Catalogue, p. 126 (LXV.2, 6) andnewgenusPhialinea, but now (personalcommunication)
1894:4:3:808 and 812, records specimens given to Parker by preferstoreferitto Procerolagena Puri (aseniorsynonymof
.
W.K. PARKERCOLLECTION OFFORAMINIFERA 51
Hyalinonetrion Patterson & Richardson). He regardsLagena Type reference. Parker & Jones 1865: 357; pi. 13, fig. 21
gracilis Williamson, 1848 (the type-species by original desig- and pi. 18, fig. 8 (asdistoma-polita).
nation of Procerolagena) as arguably a senior synonym of
Lagenadistoma Parker&Jones, in Brady 1884: 461. Localities. Fossil, Crag ofSuffolk, England. Recent, Nor-
way dredgings, 30-200 fathoms between North Cape and
Lagenasul&cata (Walker & Jacob) var. distomaaculeata SDerao,ntihnetihme,Gfurlofmof69S°uteoz6(3N°oN. (1NoofrwPualylelno'csalMitSy VliIsItI))a; lRieghdt
Parker Jones, 1865 yellow (ochreous) clay, close to Island Shadwan off S.E.
Type reference. Parker & Jones 1865: 348, 420; pi. 18, fpoaitnhtomesn,trSawnacenoRfivJeurb,alMeStlrbaoitu,rn37e2,fAautshtroamlsi;a;coJausktess'anddr,e7dgoirng8
fig. 5 (asdistoma-aculeatd) (no. 1, August6th 1844), 17fathoms,awhiteshellymudnear
Type locality. Fossil, from theTertiary, Eocene, Calcaire RaineIslet, 11°30'S 144°E,northernpartoftheGreatBarrier
Grossierat Grignon (in the Paris basin). Reef; on the Durham coast, England.
Remarks. There are Catalogue records of L. sulcata from Remarks. The recordsofLagenapolita (Catalogue, p. 165)
the Eocene, Calcaire Grossier of Grignon, France from the Pleistocene, Valley of the Nene, Wisbeach [a
(LXXXIV.59, 60), P47659-60, but these are not var. dis- misspelling of Wisbech, Cambridgeshire] (CXXXVII.2)
tomaaculeata. Asthereare norecordsintheCatalogue under P48400 and Pleistocene, Boston, Lincolnshire, eastern
aculeata, distoma, distoma-aculeataorsulcatavarieties, and a England (CXXXVIII.l) P48401 are misidentifications.
search ofall the Grignon slides has revealed nothing, it must Though never recorded under its full varietal name but as
be assumed thatthese rare specimenshave been lost. Lagena distomapolita (Recent Catalogue), it is possible that
the specimen illustrated on pi. 13, fig. 21 ofParker & Jones,
Lagena sulcata (Walker & Jacob) var. 1865 is that from the Norway locality VIII (XXVI.24)
& 1894:4:3:249, sinceinthe typedescriptiontheword 'Arctic' is
distomamargaritifera Parker Jones, 1865
written after the figure number. It is the correct size and the
?P1. 1, figs20, 21 Catalogue (p. 43) refers to this figure, but it is not clear
Type reference. Parker & Jones 1865: 348, 357; pi. 18, whetherJonesismakingacomparisonoradefinitestatement
fig. 6a, b (asdistoma-margaritifera). of fact. Jones obviously had a problem in distinguishing
distomapolita from Amphorina gracillima Seguenzar, 1862
Typelocality. Recent,spongesand('surf-washedsponges' (gracillima has a more rounded fusiform side view), for it is
inplate explanation), from Melbourne, Australia. recorded as the latter fromJukes' dredging no. 1 (Catalogue
p. 230, XIX.25) 1894:4:3:1686; it is from this slide that a
Remarks. Not recorded in the Catalogue under its full lectotype has been chosen asit comparesfavourably withthe
original name, itcan be found neither under L. margaritifera specimen in pi. 18, fig. 8 of the original figures. There are
nor distoma-margaritifera. There is, however, a record of other specimens from the Red Sea (LXXXIX.3)
Lagena distoma from Melbourne shore sand, Australia, 1894:4:3:1085, not listed in the Recent Catalogue, p. 157.
(CII.2), 1894:4:3:1521 (4th specimen from left) which is not The Jukes' dredgings come from an area which was exten-
typical of that species (compare with illustration of Lagena sively recollected by H.M.S. Challenger (see co-ordinates
distoma above), and is illustrated here. The original figured given above). The specimens in the Parker Collection from
specimen (pi. 18, fig. 6a) (the description reads as if there Melbourne (CII.l, 2) 1892:4:3:1520, 1521, I consider to be
was more than one specimen) was ornamented with 'pearl gracillima. Jones does not list 'on the Durham coast' in his
like grains'). My illustrated specimen (ZF4918) bears elon- Catalogue, and there are no specimens in the Dogger Bank
gateraisedbars,someanastomosing, andissimilartothetwo slides (XVIII and XIX) 1894:4:3:122-124; there are however
specimensfiguredbyAlbani&Yassini(1989: 381,figs3B,C) specimens labelled gracillima from this area in Brady's
as Procerolagena distoma margaritifera from Port Stephens, BM(NH) collection, which need fuller investigation. The
eastern Australia. Even though this is the only specimen fossil record from Sutton, listed as gracillima, (CXXX.22)
extant in the ParkerCollection which shows any similarity to P48311, is indeed that species. Whittaker & Hodgkinson
the type illustration, it differs enough in ornament to make (1979: 43) considered that specimens similar to those of
the designation of a lectotype unwise without knowing the Parker & Jones (1865: pi. 18, fig. 8) formed part of a series
range of variation; furthermore, it is not clear whether it is whichcould be referred to Lagena clavatad'Orbigny.
strictly a syntype. There are records from Melbourne of The species distomapolita was placed by Albani & Yassini
Lagena sulcata s.s. (CII.l) 1894:4:3:1520 (this has randomly 1989 into Procerolagena Puri (see remarks above, under
arranged pits) and L. sulcata ? (CIII.I) 1894:4:3:1569, which Lagenasulcata var. distomamargaritifera).
arenot conspecific.
witThheLaggeennusaPbryocLeoreobllaigcehna&PuTriapwpaasnco(n1s9i6d4:ereCd51s8y)nobnuytmowuass Lag(eMnoantsaugluca)tasu(bWvaalr.kesrqu&amJoascoomba)rvgairn.amtaargPianraktear &
reinstated (1988 416) without explanation (see also discus-
: Jones, 1865
sionunderLagena distoma, p. 50).
PI. 1, figs 18, 25; Fig. 1.2
Lagena sulcata (Walker & Jacob) var. distomapolita Lectotype (here designated). From Jukes' dredging no. 1,
Parker & Jones, 1865 N.E. Australia {ex 1894:4:3:1686), ZF4920.
PI. 1, fig. 22 Type reference. Parker & Jones 1865: 348, 356; pi. 18,
Lectotype (here designated). From Jukes dredging no. 1, fig. 2a, b (assquamoso-marginata)
N.E. Australia {ex 1894:4:3:1686), ZF4919. Localities. Fossil, in the Middle Tertiary beds of San
52 R.L. HODGKINSON
W.K. PARKER COLLECTION OFFORAMINIFERA 53
Domingo. Recent, living on the coral-reefs of Australia fathoms, in 'South-east or Disco Bay', Davis Straits, west
(Jukes). coast ofGreenland, 68°50'W53°N.
Remarks. The Lagena sulcata group from the Middle Ter- Remarks. There are no syntypes. No fossil specimens have
tiaryofSanDomingoisnotrecordedinthe Catalogue. There beenfound. The RecentCatalogue (pp. 10, 30) refersonlyto
is, however, a single specimen from Jukes' dredging (no. 1, Lagena striatopunctata. The type figures were of specimens
August 6th 1844), 17 fathoms, white shelly mud near Raine from the Arctic (west coast of Greenland) Hunde Islands,
Islet, 11°30'S, 114°E, northern part of the Great Barrier No. 3, 30-40 fathoms, with a shelly sandy mud bottom, a
Reef); it is labelled 'Lagena squamosomarginata ?' (Cata- sample not mentioned in Jones' Catalogue. R.W. Jones'
logue p. 230, CIX.25) 1894:4:3:1686. I have designated this 'lectotype' (ZF3416) is from the Hunde Islands No. 2, 28-30
aslectotype. Ithasaslit-like aperture and 'entosolenian' tube fathoms (Catalogue 30, XXV.2c) 1894:4:3:175, with a
:
('endosolen' of Knight, 1986), which would place it in the 'gravel bottom'. Since it is not in fact a syntype his specimen
Ellipsolageninae ofSilvestri (Loeblich & Tappan 1988: 428); is here referred to asthe neotype (p. 48).
thepresenceofashortneck, non-radiateaperture,endosolon The perforations on the ribs suggest this form should be
and peripheral keels, and absence of a phialine lip indicate included within the genus Cushmanina R.W. Jones (1984:
the genus Lagenosolenia McCulloch (Loeblich & Tappan 105). Barker (1960: pi. 58, figs 37-40) raised the taxon to
1988: 428). Barker (1960: pi. 60, fig. 24) raised squamo- specific level within Oolina. Le Calvez (1970: 82; pi. 16,
somarginatato specific level within Fissurina Reuss. fig. 1), as Lagena, records it from the Lutetian of Montjav-
oult (south of Gisors) in the Paris basin, but not from
Lagena sulcata (Walker & Jacob) var. striatopunctata Grignon.
&
Parker Jones, 1865
Lagena sulcata (Walker & Jacob) var. tetragona Parker
Neotype. The so-called 'lectotype' selected by R.W. Jones, &
Jones, 1865
1984, is infact the neotype: see p. 48.
PI. 1, figs 16, 17
Type reference. Parker & Jones, 1865: 350; pi. 13, figs
Lectotype (here designated). From Grignon (ex P47660),
25-27.
P52784.
Linosciadleitaisehse.llF(oasslitlh,ouEgohctehneespdeecpioessitiss nooftGfriiggunroedn,frFormantchei,s Type reference. Parker & Jones 1865: 352, 420; pi. 18,
locality the note in the type reference, 1865: 419, may mean fig. 14a, b.
that this was one of the species found within the Cerithium Type locality. Eocene, the Calcaire Grassier; Grignon,
giganteum). Recent, IndianOceanonaclamshellandat2200 France.
fathoms, a fine white calcareous mud with Polycystinae,
5°37'S 61°33'E; the Red Sea at 372 fathoms, a light yellow Remarks. Recorded as Lagena tetragona (slides
clay, close to Island Shadwan, off SE point at entrance of LXXXIV.59-61), P47659-61 from the Calcaire Grassier of
Jubal Strait; the South Atlantic at 2700 fathoms, pale mud, the Paris Basin, Grignon, near Paris; one of the best speci-
halfaluminous, 26°45'S 32°52"W; the Hunde Islandsat 30-40 mens is made lectotype. The 'ridges/spiral ribs' in the neck
PLATE
1
Figs1-3 AnomalinacoronataParker&Jones. LectotypeZF4913, here designated; Norway. Dorsal, edge andventralviews,respectively.
AH x 40.
Figs4,7 ArticulinamultilocularisBrady, Parker&Jones. Lectotype ZF4914, heredesignated; AbrolhosBank. Side andobliqueapertural
views, x 125. SeealsoFig. 1.1 (p. 56).
Figs5,6 HauerinaplicataParker&Jones. LectotypeZF4915, heredesignated;N.E. Australia. Sideandoblique-aperturalviews, x 95.
Figs8-10 HippocrepinaindivisaParker. Figs 8,9, Lectotype ZF4916, heredesignated; Gaspe Bay. Side andobliqueaperturalview, x 45;
Fig. 10, Paralectotype ZF4916a, brokenspecimenfromsame locality,viewshowinglackofinternalstructures, x 130.
Figs11, 19 Lagenasulcata(Walker&Jacob)var. trigonomarginataParker&Jones. P52785, from the EoceneofGrignon, France;
oblique-aperturalandsideviews,respectively. Bothviewsshowingthe threeribsonasinglemarginonly, x 195.
Figs12, 13 LagenadistomaParker&Jones. Lectotype ZF4917, heredesignated; Norway. Fig. 12,detailofornament, x 395;Fig. 13,
entirespecimen, x 40.
Figs14, 15 LagenacrenataParker&Jones. LectotypeP52783, heredesignated;MiddleTertiary. Malaga. Basalandsideviews,
respectively, x 100.
Figs16, 17 Lagenasulcata(Walker&Jacob)var. tetragonaParker&Jones. LectotypeP52784, heredesignated;CalcaireGrossier,
Grignon. Sideandoblique-aperturalviews,respectively, x 135.
Figs18,25 Lagenasulcata(Walker&Jacob)var. marginata(Montagu)subvar.squamosomarginataParker&Jones. LectotypeZF4920,
heredesignated;N.E. Australia. Oblique-aperturalandsideviews,respectively, x 160. See alsoFig. 1.2(p. 56).
Figs20,21 PossiblyLagenasulcata(Walker&Jacob)var. distomamargaritiferaParker&Jones. ZF4918, fromspongesand, Melbourne,
Australia. Fig. 20,detailofornament, x 300;Fig. 21, entirespecimen, x 40. Seetextp. 51.
Fig.22 Lagenasulcata(Walker&Jacob)var. distomapolitaParker&Jones. Lectotype ZF4919, heredesignated; N.E. Australia. Side
view, x 40.
Figs.23,24 Lagenasulcata(Walker&Jacob)var. tubiferosquamosaParker&Jones. LectotypeP52786, heredesignated;Calcaire Grossier,
Grignon. Fig. 23,sideview, x 75;Fig. 24,detailof'tubes'on neck, x 400.
Fig.26 LituolanautiloideaLamarckvar.globigeriniformisParker& Jones. Lectotype ZF3658; northofNewfoundlandBank. Ventral
view, x 330.
Allscanningelectronphotomicrographs.
.
54 R.L. HODGKINSON
region can fork and recombine. If Le Calvez, 1970, ever Remarks. There aresyntypesrecordedasHaplophragmium
observedthisspeciesfrom Grignon she didnot recordit. cassis (Catalogue p. 126, LXV.3) 1894:4:3:809 and a strew
(LXV.7) 1894:4:3:813 with several specimens. This large
Lagena sulcata (Walker & Jacob) var. species is under fuller investigation by Bronnimann & Whit-
trigonomarginata Parker & Jones, 1865PI. 1, figs 11, 19 Tctahakopesprean(n.inaIpntrdeipsstairltlahteliivotensy)p,ienotnshpewechtioyepsseeaoarfdevaAism(eSmcoahtalfeiecurtmo&tyLpCoeoelhbealsi1c9bh7e8e)&n.
Type reference. Parker & Jones 1865: 348, 352, 419;
pi. 18, fig. la, b (astrigono-marginata) Lituolafindens Parker, in Dawson 1870
Typelocality. Eocene, from inside a Cerithiumgiganteum Lectotype (here designated). From Gaspe Bay (ex
shell, Calcaire Grassier; Grignon, France (Parker & Jones 1894:4:3:810), ZF4929.
1865: 419, note).
Type reference. Dawson 1870: 176;text-fig. 1 (p. 177).
Remarks. Recorded as Lagena trigonomarginata in the
Recent Catalogue (p. 10) from Skye and Whitehaven, but Localities. Recent, from Gaspe Bay, 16-20 fathoms; and
not from Grignon (Fossil Catalogue, p.100-107). There is a St George's Cove in Gaspe Bay; Gulf of St Lawrence,
marginata (Mont.) mentioned on p. 103, (LXXXIV.57, 58) Canada.
P47657-8, but the specimens so called are not conspecific Remarks. Recorded as Reophaxfindens (Catalogue p. 126,
since they lack the 'three, meridional, three-edged equal LXV.4) 1894:4:3:810. Of the specimens in the collection,
ribs'. P47656-67 and P47724 (LXXXIV.56-67 and 126), onlyonecouldpossiblyhave been afiguredspecimen (centre
twdphselihuhradrsienepc.eeenhsocP,ttrsa.ihhbrtoasFhkusaeriltrnbodetm&ohehnaneJsevloieninspdueocleslmoaa'bnLntteefeXadirX,oigXnunorlreIfeyed-V,drrs.iesubb6glpusi1cetsaicttsbie(airmPv-eaa4erdnr7niid6Pba/h65bboa1el2rs)de7.m8niaao5nNrtsgaotbpinheenedlaceetiinoclamttl,fheouoetnsiurytnnrpwdafteita;wteciohdtIs trsbdHepoere-epcaricoawolituilnmyusn-sepegAtnelrssalo.ittefeoTndfhtiheesRaexrentre-ebodf,rpiaogrhnE.keaoaerxrn1n)loifassinaiayntdnnndsddteeSynptpsiauseerdxs(aehtPtfneraertrerdsoemkmeeflrCrdSo)yoetlsmlGaiefertgcroehntaeragigpitoalerencee'd,cssoeiBlCnmteoMtpcva(tieisnoNn,tyHyibntp)nuoeh.gtt;e
desTihgenatteadx.on may well belong in Galwayella Patterson & LBoreabdlyi'schC&halTleanpgpearns(p1e9c8i8m:enpsi. o4f7,Refiogpsha7-x10f)inrdeeinlslusftrraotme
Richardson. Gaspe Bay, 18-20fathoms (1884: pi. 32, figs 10a, b and 11a,
Lagena sulcata (Walker & J&acob) var. b),LiBtMuo(lNaHfi)ndZeFn2s2i7s2t.he type species ofProtoschista Eimer &
tubiferosquamosa Parker Jones, 1865 Fickert.
PI. 1, figs23, 24
Lectotype (here designated). From Grignon (ex P47669), Lituola nautiloidea Lamarck var. globigeriniformis
&
P52786. Parker Jones, 1865
Type reference. Parker & Jones 1865: 354, 420; pi. 18, PI. 1, fig. 26
fig. 7a, b (astubiferosquamosa). Lectotype. From north of the Newfoundland Bank,
ZF3658. Designated by Loeblich &Tappan, 1964.
Typelocality. Eocene, from inside a Cerithiumgiganteum
shell, Calcaire Grassier; Grignon, France (Parker & Jones Type reference. Parker & Jones 1865: 407; pi. 15, figs46,
1865: 419, note). 47 (Arctic) andpi. 17, figs96-98 (North Atlantic).
Remarks. Referredtoastubuliferosquamosa(amis-spelling Type localities. Recent. 30-70 fathoms at Hunde Islands,
by Jones) in the fossil Catalogue (pp. 103, 107, in South-east or Disco Bay, Davis Straits, west coast of
LXXXIV.68-72, 126), P47725, P47668-72, from Grignon. Greenland (68°50'W 53°N); from Baffin's Bay, depth not
The figured specimen has not been isolated and would be recorded (75°10'N 60°12'W); also at 314 fathoms (75°25'N
difficult to recognize with certainty. The specimens on slide 60°W) and 220 fathoms (75°N 59°40'W); in the Boreal tract
LXXXIV.126 have been remounted (P47725). There are towards Newfoundland Bank, 1660fathoms (51°30'N 38°W);
numerous, mostly worn and poorly preserved, specimens on from the north of Newfoundland Bank at 145 and 954
slides68 (P47668) and 69 (P47669); few have the characteris- fathoms,NorthAtlanticOcean;inthe RedSea, 557fathoms,
ticneck, which mightaccountforLe Calvez' (1970) failure to (17°49'N 40°02'E) and 678 fathoms (23°30'N 36°58'E); from
report them. One of the most complete individuals on slide the Abrolhos Bank in the South Atlantic at 260 fathoms
P47669 has been designated lectotype. This species is unlike (22°45'S 40°37'W) and 940 fathoms (19°32'S 37°51.5'W);
anylagenid I have seen andpossiblynot even a foraminifer. from the Indian Ocean, 2200 fathoms (5°37'S 61°33'E); and
the Mediterranean sea (no detailsgiven).
Lituola cassis Parker, in Dawson 1870 Remarks. The lectotype selectedand figuredbyLoeblich &
Figs2.1, 2
Tappan (1964: C259, fig. 173:2a-c) is that figured by Parker
Lectotype (here designated). From Gaspe Bay (ex & Jones (1865: pi. 17, fig. 96) from the North Atlantic (ex
1894:4:3:813), ZF4637. 1894:4:3:294), ZF3658. It is from a locality at 49°23'N
48°48'W, 954fathoms, Arctic, northofNewfoundlandBank,
Typereference. Dawson 1870: 177, text-fig. 3. as given in Table 5 of the type reference, but not quoted in
Type locality. Recent, from Gaspe Bay, between 10-17 full in the Ellis & Messina Catalogue.
fathoms; GulfofStLawrence, Canada. L. nautiloidea var. globigeriniformis (asAmmoglobigerina