About the Author Michael Benton is Professor of Vertebrate Palaeontology and was previously also Head of the Department of Earth Sciences at the University of Bristol. His many books include standard reference works and textbooks as well as popular books about dinosaurs and the history of life. Other titles by Michael J. Benton published by Thames & Hudson include: The Seventy Great Mysteries of the Natural World Other titles of interest published by Thames & Hudson include: The Complete Ice Age: How Climate Change Shaped the World The Reign of the Dinosaurs The Story of Fossils: In Search of Vanished Worlds Our Fragile World: The Beauty of a Planet Under Pressure Our Living Earth The Great Naturalists Elephant! The Illustrated Natural History of Selborne See our websites www.thamesandhudson.com www.thamesandhudsonusa.com To Mary, Philippa and Donald for their forbearance and inspiration, as ever ACKNOWLEDGMENTS I am hugely grateful to Colin Ridler of Thames & Hudson for first suggesting that I should write this book, and then for discreetly guiding me through various false starts. Thanks to him, and to his team, for their thorough and extensive work on the book as it went through the publication process. The entire text has been read by Paul Pearson and Richard Twitchett at the University of Bristol, James Lovelock at the University of Oxford, and Paul Wignall at the University of Leeds. I value the time they spent reading the draft manuscript, their wise suggestions about style and content, but especially for picking up some glaring misunderstandings and errors of fact. Thanks also to archivists and librarians at the Geological Society of London, the British Library and the Natural History Museum, for finding ancient letters and notebooks by Murchison, Owen and the others. Finally, the appearance of the book has been enhanced immeasurably by the artwork, and I thank John Sibbick from Bath for his exquisite craftsmanship informed by a real knowledge of the ancient beasts he draws. Finally, I dedicate this edition to Vitaly Georgievich Ochev, our delightful companion in the field in Russia (chapter 10), who died in 2004. CONTENTS PROLOGUE THE GEOLOGICAL TIME-SCALE Chapter 1 ANTEDILUVIAN SAURIA Chapter 2 MURCHISON NAMES THE PERMIAN Chapter 3 THE DEATH OF CATASTROPHISM Chapter 4 THE CONCEPT THAT DARED NOT SPEAK ITS NAME Chapter 5 IMPACT! Chapter 6 DIVERSITY, EXTINCTION AND MASS EXTINCTION Chapter 7 HOMING IN ON THE EVENT Chapter 8 LIFE’S BIGGEST CHALLENGE Chapter 9 A TALE OF TWO CONTINENTS Chapter 10 ON THE RIVER SAKMARA