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Wabua, a new spider genus (Araneae: Amaurobioidea: Kababininae) from north Queensland, Australia PDF

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Preview Wabua, a new spider genus (Araneae: Amaurobioidea: Kababininae) from north Queensland, Australia

WABUA, ANEW SPIDER GENUS (ARANEAE: AMAUROBIOIDEA: KABABININAE) FROM NORTH QUEENSLAND, AUSTRALIA VALERIETODD DAV1ESANDCHRISTINE LAMBKIN Davies, V.T& Lambkin,C.2000 1231: Wabua,anewspidergenus(Araneae: Amaurobioidea: Kababininae)fromnorthQueensland,AustraliaMemoirsoftheQueenslandMuseum46(1): 129-147. Brisbane. ISSN 0079-8835. Eleven newspeciesofWabuagen. nov. aredescribed; theseare W major,thetype species and W.hypipamee, W.kirrama, W.seaview, W.elliol, W. eungella, W. crediton, W.aberdeen, W. Cleveland, W.paluma and W. halifax. All were collected between latitudes 17°16' and 21°13'S.AcladisticanalysisshowsthattheKababininaecontinuestoformawellsupported O monophyletic group though its placement in a family remains problematical. Araneae, Amaurobioidea, Kababininae, Wabua, taxonomy, distribution. Queensland, Australia. Valerie ToddDavies, QueenslandMuseum, POBox3300, SouthBrisbane4101. Australia: ChristineLambkin. DepartmentofZoology& Entomology. UniversityofQueensland, St Lucia4072, Australia: received 10November 1999. This is the fourth genus to be described in the (SF),National Park(NP). SandEaregiveninthe subfamily Kababininae: others are Kababina latitude/longitude dataforthe holotype only and Davies 1995, Carhinea Davies 1999 and omitted forthe paratypes. Malarina Davies 2000. The three described Anatomical abbreviations used in text, figures generaareallconfinedtothe WetTropicsregion andinthelistofcharactersandtheirstates(Table of northern Queensland which extends from 2): AL, abdomen length; ALE, anterior lateral CooktowntoTownsville. Wabuagen.nov.isalso eyes; ALS, anterior lateral spinnerets; AME, found in the Wet Tropics region (between anterior median eyes; APOPH, apophysis; ATP, latitudes 17°16' and 18°36'S) and, unlike the anterior tegular process; AW, abdomen width; 3 others,extendsfurthersouthtoCrediton(21°1 'S), CB,cymbium;CAL,calamistrum;CH,cheliceral; mid-east Queensland. For nomenclatural CL, carapace length; CR, cribellum; CW, cara- purposes Davies is designated the author ofthe pace width; E, embolic; EG epigastrial groove; names ofthe new genus and its species. All the EPIG epigynal; ID, insemination duct; MAP, species are named for their type locality. The major ampullate spigots; MT, metatarsal; P, female epigynum varies little between species patellal; PCR, paracribellar spigots; PE, parem- whereas the male palp is complex and provides bolic; PLD, prolaterodorsal; PLE, posterior most ofthe characters for separation ofspecies. lateral eyes; PLS, posterior lateral spinnerets; The figuresofepigynaand localityofspecimens PME, posterior median eyes; PMS, posterior should aid identificationswhen usingthe key. median spinnerets; RTA, retrolateral tibial MATERIALS ANDMETHODS apophysis; T, tarsal; TRICH trichobothria. SYSTEMATICS Collection methods include litter-sieving followedbyheatextractionin funnel,pitfall(PF) Subfamily KABABININAEDavies 1999 collection, pyrelhrum (PY) spraying of tree trunks and fallen logs, hand collecting from DIAGNOSIS. Medial atrium ofepigynum wider under logs in daylight and night collecting. than long; spermathecae posterior or lateral to Notation of spines follows Platnick & Shadab atrium. Male palp with rounded tegulum with (1975). All material is lodged in the Queensland prolateral groove; course ofsperm duct showing Museum (QM). Cladistic methods are given clearly. Membraneous conductor; median underthe section on relationships ofWabua. apophysis absent. Tibial apophysis with ventral and dorso-retrolateral branches. Abbreviations. Most spiders were collected by G.B. Monteith (GBM) and fellow collectors D. DESCRIPTION. Carapace highest in foveal Cook(DC),D. Yeates(DY),G Thompson(GT), region; posterior eye row straight or slightly H. Janetzki (HJ) and S. Hamlet (SH); other recurved; AME reduced. Chelicera with two collectors include R. Raven (RR) and V.E. retromarginal and two promarginal teeth; Davies (VED). In location data: State Forest prolateral filamentoussetaatbaseoffanglonger , 130 MEMOIRS OF THE QUEENSLAND MUSEUM than other setae. Feathery hairs on legs, ridged Tibia ofmale palp with 2-3 discrete retroventral setae cuticle.Malepalp:emboluswithorwithoutprox- (Fig.4A) 8 imal embolic apophysis. Cribellum (two fields) 6. Embolus with smooth curve from origin to prolateral lparregseenbtrooradabcsoelnutlusinpfreemsaelnets,whaebsnenctriibnelmlaulmesi;s Eemdbgoel(uFsigw.i4dJe)ningatprolateralrum(Fig.6A) enngell7a 7. Embolic apophysis with 2 pointed 'folds' subdistal to absent. embolus(Figs6A.7D) crediton Wabua Davies gen. nov. Embolic apophysis with 2 reduced blunt foldsa(bFeirg.de6eJn) TYPESPECIES. Wabuamajorsp.nov. 8. TTiibbiiaaooffmmaalleepapallppabmouucthaslloonnggearstvhvaindwei.deA.TPTaebgsuelnutm.wi.t9h ETYMOLOGY. Derived from the Aboriginal 'wabtt'. mid-proventralprotrusionanddigitiformATPC(lFeivg.el8aAn)d meaning 'forest' in the Djirbal language of north 9. Embolus with proventral keel (Figs 4A. 5A). RTA Queensland. pointed,digitiform elliol DIAGNOSIS. Female cribellate (cf. Carbinea). Embolus without proventral keel (Fig. 2E). RTA blunt kirrama wEiptihgysnhuamllwoiwthomuetdipaoslteartiroirukmn.obC(ocnfd.uMcatloarrinaan)d\ 10. Elmabteorlaluskeewlitdhistlaalrlgye(Friegt.ro8lJa)t;erpaallpakleetlibiaandasslmoanlglaesrwpirdoe- embolusofmalepalparisingantero-ventrally on paluma tegulum (cf. Kababina); without proximal par- Embolussmoothwithsmalldistalkeel(FigsSO.9D,E); embolicapophysis(cf.CarbineaandMalarvia). palpaltibiaslightlylongerthanwide halifax DESCRIPTION. Small spiders (most less than Wabua major Davies sp. nov. 4.0). Carapace pale with 2 dark longitudinal (Figs 1A-E, 2A-C, I0A; Table 1) bands (see Malarina Davies & Lambkin, 2000: friagd.iat1iAng)fsroommeftoviemae.sAbrdeodmuceendwittohdaaprakttelrinneosf SMI37A9°T2380E'3RS),I.A1PL4A5.R°3A2HT'OYE,LPOEliTStt:Yer,PNE14:Q-l2d0:9.,i6v.\M1a9sj7a8om,resVdEMaDtaa,i,(RS3NR92(0Q4lQ)dM;. pairedspotsgivingwaytolightmedianchevrons 66", 9, same data (S39205); 9, Majors Mm, 7km SE ondarkground(seeKababina,Davies, 1995:fig. Ravenshoe, 1100m, sieved litter, 4.V.1983, GBM, DY ID). Legs banded with darker rings. Conductor (S39206): 49, Maalan SF, litter, 20.iv.1978. VED, RR with stalk broadening to saucer shape with (S39207): 29, Mt Father Clancy, Maalan SF, litter. 21 raggededge.Embolusbroad,narrowingdistally. iv.1978,RR(S39208); 3,MtFatherClancy,9kmSMillaa Millaa, PF, 6-14.xii.1988, GBM, GT (S39209); 29, 6", KEYTO SPECIES OF WABUA Downey Ck, 25km SE Millaa Millaa, I7°39\ 145°47', 1. Posteriormargin ofepigynal atrium closeto epigastrial sPiaelvmeedralittotenr,N4P0,0mE,7m.xairig.i1n9,88,17G°B3M7\,G14T5°(4S63\9241000);m,9,P6F, tfhuarnroIwV(Figs IB. 2A). Leg I equal toorslightl'y.Io.ni7ze2r 2I.Ox.ix.i-il.0lx9i9i5.-179.9i5i,.Gl9B9M6(,S3D92C12()S;399.2131p)e;n4ul6t\.sd,amTeullloycaFlailtlys. 2. RPeoTpsitAgearsoitforrimaalmlaferugrpiranolwpo(wfFiigte.hpi8dgFiy.sKnt)aal.lLaaentdgriIpusrmhooxrwitemelarllthfpaornorjIweaVcrtd.io1no0sf t9Rud0r,0noml,1T0,8k.TmXuJlSlUv98R9aFav,lelnGssBhRMode,,,G11T77,°45H30J\\(IIS443559°°23312I3'');,;s7i65e",0veRdm"e.dlitPtReFrd,, (eFpiigg.yna1lA,aJt)r.iuDmisatnadnacnetebrieotrwleoeonpoafntienrsieomrinmaatriognindueotf 8.xii.1989-511990, GBM, GT, HJ (S39214); 6, 49, lessthanlengtholatrium 3 Upper Boulder Ck via Tully, 17°50', 145°54', 900m, RaiTnnAtseermwiiinotarhtoimuotanrdpguricotnxhiaomlaffl-axtp2rrolijeuenmcgttihaonno,dfattaDrniitsuetmra)inocre lboeotpweeo4nf sNsiaNemveWeddTalutiltalteyr,(,S8325790.2-x11.701)90;8037m,c,?,GPFB9,M,1,7U-pIDp8Ye.,xri.GBloT9u8l4(d,Se3Vr9E2CD1k.6,)G;1B16Mk9,m, 3. RmiTdA-prwoivetnhtrpalropxroitmraulsiobnuolfgteegu(lFuimg. 2C). Witmhaojuotr J(.S3G9a2l1l9o)n;, 9D,C.1G00T0m(,S3s9i2e1v8e)d;li9tt.er,saI7m.exi.dla9t8a4,, 1G0B0M0,m 4. DR(mliTiMosdAoat-palvneweocinfetptiharbnpaesaltlleawpmrreigtoenietnabritupaaisnroitownoeixnrtdiiohumofcarttl2me-xag4srup1glruiueorntmrro(oIlFvfFeiesiagsngt.tsrrlIaieKlunUgm.ttshaeh2ntoyaDdef)p,a.ianptteraWeiimrtiuihetmoeeh.rr PVRl9idEt,Y)teD,Mro,,tn7I.GTmv7y.T°os13so9(2sn8S\,y33.29rI2Dok42cY50km)s°W.(;3S364T9\.u2,xli1l1i1v11.8,20l.)9x01;8i7m39°.,,51959G'.8l,Bi46ttM1(e,r4S,M,53t9°G125FT824i,.1\sV)h.H6;e1J5r9099((,m5KS,,j41e20slG1il0ebB01vem1eMr),dg; Ddliiossotcparneoctfeeiobnresfteowmriemneianntigaonatnecrdoiumocrbt)mxar2gtihneolfenagttrhiuomfaatnrdiuanmte(rFiigo5.r (C(rSSa44t22eIr1,11971)):7;°4297,9',,Mst1a4mF5ei°s2hde9art\,a17(2Sk34m02m1S,1W8)l;itMt4ierl9.l,a2a36.Mx,iiTl.ol1aw9a9e.4r,InG7r°BT3h4Me\ 3G).Malepalpaltibiawith3longretroventralsetae(Fig. 145°34\ 1050m, litter, 27.iv.l982, GBM,DY,DC 3J) seaview (S42120);49,2penult6,samedata. 1100m,(S42121);2 5. Tibiaofmalepalpwith4longretroventralsetaeforminga 6\ Millaa Millaa Lookout, 17°31\ 145°34\ 1000m, PF, comb(Fig.6A) 6 Lxii.1993-25.ii.1994, J. Hasenpusch (S42123); 9, 6\ 5 1 NEW SPIDER GENUS EROM THE WET TROPICS 131 Sluice Ck, 9km WSW Millaa Millaa, 17°33\ 145°33\ V021; II D101, P101, V010, R001; III, D010, 1150m, sieved litter, 5.xii,1988, GBM,GT (S42124); 3, P101,V11LR101; IV D101.P11J, V1I2, ROIL M1a0s0s0me,y sCike,ve1d2lkitmter,S4W.V.M1i9l8l3a,aGMBilMla,a,D1Y7°(3S74\211254)5;°363,\ (MeFtiagtsar1sAi,: 2sBp,iCn)e:d weimtbholduisstacluwrhvoerdls4h-a5r.pl6y wpiatlhp 21km S Atherton, 17°27\ 14518', 1040-1100m sieved litter,5.xi.l983, DY,GT(S42126). , flange-likeapophysis. Shorttibia,slightly longer than wide, 1:0.9; excavatedventrally with distal DIAGNOSIS. Epigynum with broad antero- spur-like apophysis and proximal bulge. lateral gonopores, insemination ducts narrowing Spinnerets:ALSwithonemajorampullatespigot toformanteriorloopbacktospennathecae.Short and a nubbin, about 14 piriform spigots, some malepalpaltibia,excavatedventro-retrolaterally tartipores. PMSwithlargeanteriorspigot(minor with distal spur and proximal bulge. Two long ampullate)and7-8spigotsoftwosizes. PLSwith proventral tibial setae extending half-way up several spigots. Large colulus. Males varied in tegulum. length from 3.1-3.5. DESCARWIPTION. Female. CL 1.4. CW 1.0, AL DISTRIBUTION.(Fig. 10A)Widespreadinrain- 1.6, 1.1.Viewedfromabovetheeyerowsare forests from 400-1230m in the Hugh Nelson, straight; from the front anterior row is straight, CardwellandWalterHillRangestotheimmediate posteriorrowisprocurved. RatioofAME:ALE: south and westoftheAtherton Tableland. PME: PLE is 5:10:10:10. Legs 1423 (Table 1). Notation ofspines. Femora: I, DUO, P001; II, Wabua hypipamee Davies sp. nov. DUO, P001, R001; III, D100, POOL R001; IV, (Figs 1F-K, 2D, 10B) D100, P001,R001. Patellae: I-IV,D001. Tibiae: sIR,p0iV1n01e1;d0;IwVi,IIt,hDP100di01s1,t,alPV00w01h10o,r;lVIII4I,-151D,.0R0E11p0,i1gP.y0n0M1eut,matVa(0rF1siig1:s, (M17QA°fM26E'SRS4,I2A11L4659.)°.2H8PO'AELR,OA9TT5Y0YPmPE,E:Sst:i9c,NkMbQtrludHs:yhpi5ni,gp3a95m.,xe.es1a9Nm80Pe,,dNGatQBaldMa,s hololvpe (S42113); 23, Maalan Rd, 2km S Palmerston 1B-E, 2A): broadshallow atrium, anteriorgono- Highwav, 17°36\ 145°42\ 750m, PF, 26.xi.1994- pwoirtehs.2Spfeirelmdast;heAcLaSe lawtietrahlt2oamtarjioumr. aCrmipbuellllautme l1O0..ii..-1799.5i'i,i.Gl9B95M,(S4J.21H1a5s)e;npSu,scsham(eS42da1ta1,4):7.6ii,i.s-a15m.ev.dla9t9a5,, spigots, posterior slightly smaller; about 1 (S42II6); 39, <J, Millaa Millaa Falls, 17°28', 145°36\ piriform spigots. PMS with large anterior spigot 800m, litter, 17.V.1995, GBM (S42I22); S, 21km S (minorampullate),alateralandaposteriorspigot Atherton, 17°27\ 145°28\ 1040-llOOm, sieved litter, (cylindricals), 4-5 paracribellar (one shaft per 5.xi.l983,DY,GT(S42127); 9,samedata(S42128). base) and some acinifonn spigots. PLS with a DIAGNOSIS. Tegulum with mid-proventral large anterior spigot and 9 smaller spigots of2 sizes. Femalesranged in lengthfrom 2.9 -3.2. protrusion (cf. W. major); malepalpal tibia with Male. CL 1.7, CW 1.4, AL 1.8, AW 1.3. largespur-likeproximalapophysis(cf. W.major). Eeribellate. Eyes similarto female. Legs 1=423 DESCRIPTION. Female. CL 1.4, CW 1.0, AL (Table 1). Notation ofspines. Femora: I, DUO, 1.6, AW 1.1. Ratio ofAME: ALE: PME: PLE is P002, R001; II, DUO, P001, R001; III, DUO, 4: 9: 9: 9. Legs 1423, 1 5.3; II4.6; III 4.0; IV5.2. P001, R00I; IV, DUO, R001. Patellae: I, D001; Notationofspinessimilarto W.major. Epigynum II, D001; 111, D10I, IV, D101. Tibiae: I, P101, (Fig. 1F-I) Spinnerets: similar to W. major. Females varied in length from 3.0-3.6. TABLE 1. Palp and leg measurements of 9 (c?) Male. CL 1.6, CW 1.3, AL 1.6, AW 1.1. Colour- Wabuamajor sp. nov. ation and eyessimilarto W. major. Legs 1=423, 6.7; II 5.2; III 4.9; IV 6.7. Male palp (Figs 1J-K, Femur Patella Tibia Mela- Tarsus Total 2D): tegulum with a ventral protrusion tarsus supportingtheconductorandembolus;tibiawith Palp 0.7 0.2 0.5 - 0.8 2.2(2.0) anterior ventral and retrolateral apophyses and (0.7) (0.2) (0.3) (0.8) largeproximal spur. Males varied in length from Legl (11..74) (00..54) (11..83) (11..72) (01..08) (56.-17) 2.9-3.2. 11 (11..15) (00..54) (01..39) (11..30) (00..77) (45..13) DISTRIBUTION. (Fig. 10B)Fromrainforestsat 750-1100m in the Hugh Nelson Range and 1.0 0.3 0.8 1.0 0.6 3.7 111 (1.4) (0.5) (1.1) (1.2) (0.7) (4.9) adjacentparts ofthe Atherton Tableland. This is IV 1.3 0.4 1.2 1.3 0.7 4.9 the northernmost species of Wabua. It overlaps (1.7) (0.6) (1.7) (1.8) (0.9) (6.7) partly with the range ofW. major. 132 MEMOIRS OF THE QUEENSLAND MUSEUM FIG. 1. A-E, Wabua major sp. nov; A, 8 palp (ventral); B-E, 2 epigynum (ventral, ventral cleared, dorsal, lateral).F-K, Wabuahypipameesp.nov;F-I, 9 epigynum(ventral,ventralcleared,dorsal,lateral);J,K, 8 palp (ventral,prolateral). Wabua kirrama Davies sp. nov. (Figs 2E, 3A-E, 10B) DESCRIPTION. Female. CL 1.4, CW 1.0, AL AW MATERIAL. HOLOTYPE: 2, Kirrama Ra, Mt Smoko 1.5, 1.0. Colourandpattern similarto other turnoff,NQld, 18°12'S, 145°46'E,600m,PF, 10.xii.1986- species. Legs 1423, 1 5.0; II 3.9; III 3.6; IV 4.8. 11.i.1987,GBM,GT,SH(QMS42172).PARATYPES:N Epigynum (Fig. 3B-E). Femalesvariedinlength Qld: 2, 8,samedataasholotype(S42132); 82,Kirrama from 2.9-3.1. R10a..x(ini.r1Y9u8c6,caGbiBnMe,Ck)GT18°(1S04\2113415)°;45'2,,70B0omu,ldseirevCekd,litvteira, Male. CL 1.4, CW 1.1, AL 1.3, AW 1.1. Legs TGuBllMy,, D1Y7°,5G0\T(14S54°25147'0,);63502m,,2s8ie,vsedamleitdteart,a2(7S.3X9.1291853),; 21E4,233,A1):6.v2;erIyI4s.m9a;lIlIIi4n.t4u;rIneVd6.RT1.AM.alMaelpeaslpva(rFiiegsd 225c.?v,iiD.-o30w.iniie.y19C9k2,RRdR,,PaP.l&merEs.tLoanwlNePs,s,1M7°.3S6\ha1w45(°S24462\42P)F;, little in length 2.7-2.8. M8., sSahmaewl(oSc2al4i5t9y,2)3.0.x.l991-24.vii.I992, P. Lawless, RR, DISTRIBUTION.(Fig. 10B)Occursat400-700m in rainforests of the Kirrama and Walter Hill DIAGNOSIS. Embolus with small fluted flange Ranges.Itoccursmostlytothesouthoftherangeof subdistally; palpal tibia with small blunt distal W. major but overlaps with it in the Walter Hill RTAonly (cf. W. majorand W. hypipamee). Range. NEWSPrDERGENUS FROM THE WETTROPICS 133 FIG.2.A-C, Wabuamajorsp,new.;A,epigynum(ventral);B,C, 8 palp,B,embolicregion,C.tibialapophysis.D, Wabuahypipameesp.nov..protrusionoftcgulum,embolusandconductor.E, Wabuakiiramasp.nov.(Boulder C'k) £ palp(ventral). F, Wabuaseaviewsp. nov. 8 palp(ventral|. Wabua seaview Davies sp. nov. samedataasholotype(S42133);39,Mt Fox Rd, 18°50", (Figs 2F, 3F-K, 10B) I45°50\ 600m, sieved litter, 15.xii.l986, GBM, GT (S42I34);<J,9,samelocality,2.i.1987,SH(S42135);3<3, MATERIAL. HOLOTYPE: 9, Seaview Range, Mt Fox 9. Wallaman Falls RdJunction, I8°39\ I45°52*, 650m, RdNQld. I8°38'S, I45°54'E,600m, 15.xii.l986,GBM. PF. S-12.ii.1996, GBM (S42I36); cJ, same data, flight GT, SH (QM S42171). PARATYPES: N Qld: 9, 3J, intercepl trap(S42139); 9,WallamanFallsRd.500m. PF, 134 MEMOIRS OF THE QUEENSLAND MUSEUM \-f. Wabua ktttanta sp. nov.; A. •' palp (ventral). B-E, eptgynura (ventral, ventral cleared, d it • lateral)- P-K; Wabwiseavisw sp. nov.; II. epigvnum (ventral, ventral cleared, dorsal, lateral); ,I,K. f palp (ventral, rein. si] I4.xii I986-2.U987, GBM. '.II. Ml (S42137); . Male. CL 1.7, CW 1.4, AL 1.6, AW 1.3. Legs GWaBlMlamanFallsRd, 18 !8' l45°Sr.6S0m.litter,5.ii.l996, 1=423. 1. 7 0; 11 5.6; 111 5.0; IV 7.0. i palp(FigS (S42I38). 1'f, 3J,K): retrolateral edge ofembolus reflexed. DIAGNOSIS. The distancebetween the anterioi I'alpal tibia with 3 relrovcnlral setae; RTA small, margin ofthe atrium and the anteriorloopofthe blunt. Males varied in length from 3.0-3.4. insemination ducts is almost twice the length of DISTRIBUTION, (fig LOB) Restneted to the (he atrium (cf. W\ major and other species). Seaviev, Range where i( occurs in rainforests at Keeled retrolateral edge ofembolus: small blunt 500-o50m. K A.Palpaltibiawith3discreteretrolateralsetae. I DESCARWIPTION. Female. CL 1.8, CW 1.4, AL Wab(uFiagsel4lAio-tE.D.a5vAi,csIs0pB.]nov. 1.9, 1.4 Colouration and pattern \ir,nl. o6.t9h.erEsppiecgivens.umLe(gTsig1.42331,-1I).7.0f;eImIa5l.e6s; LvITar5i.e0d; IiVn VII9DV3t0'r..SR,IAII4(,..°5I7i'Or-L.O1T1Y5P0fm:.296.,iiMil.1t9i9lilf.crtGsBuMmm,iti,kIN(QQldM, length from 3,3-3.8. S17863). PARATYPES: N Old: :»<.k data as i NEW SPIDERGENUS FROM im WETTROPlCS FIG.4. A I Wabaaeltioi sp. nov.; A. ,', palp(ventral); B-E.epiRynum (ventral, ventral cleared,dorsal. i -k. Wablui eiaigeliosp. nov>; I -F eplgyrtuiii (ventral, ventral cleared, 6 rah: .i.K. i palp (ventral, ii*cr»llrfrotllanlle.-r>«a-'l!)I.t Iiolot57v'p,e,D1t000(mS,422154v52)7;.iyt,.lM9t9)E,lliGoBt,MN,ortIhXOr(,$4i21"4i6H|);. (bFriogade4rA)th:anplromnegn,tr1:a1l1.7,keReTlAopnoienmtbedolwuist,h etidbgiea samedata(SI7876); S. Ml Elliotsummit, 1150m, incurved,Malesvaried inlengthbetween3,2 sP3i-a5maexnidilMo.lci1an9lk8iE-6tli,ylc.ic(opPJttFBtMrIi.a\-pP2sG.6,T2,iUoip.iSi.p1Hie9i9r.(1-,Sl42N.2\o.v1r.6Glt69rh)9a.1hC.akDm,C(S(I4St24H21JI6O45m7).);;2PI". rDaIiSnTfoRrIeBsUteTdIOsNu,mm(iFitg,(11U0H0)0-RIe1s5t0rincit)edotfot'hbee isolated Mt Elliot, SWofTownsville. DIAGNOSIS. Distance between the anterior margin ofthe atrium and the anterior loop ofthe Wanna enngefla DaVies sp. nov. insemination duels is less than halfthe length of (Figs4F-K, SB-F, IQB) theatriumicfl H seavtew), Longprovcniralkeelon embolus;RTApointeddigitilurrn ief iimerspecies): 3 longdiscrete ventral setae on palpal tibia. MATERIA] . hoi OTYPE; I ungella (schoolho DESCARWIPTION. Femufc. CI 1.5, CW IJ, \L m1i2d.-8c.a19s8i6OJld,Gal2l1o°nI.0*RSR, (IQ4M8°2S442E1,9Ir)ainPfAorReAstT,YPliEtnSer:, 18, 1,3. Carapace pattern oT dark lines ,,„ eastQld; 1,6 - ,sarrjedataasholorype(S42155); 6, radiating from fovea; abdominal pattern similar locality, PF. II-I4.S.I986,.1. Gallon, RR (S42192); lo other species. Eyes, notation l4 leg spines mc lata(.S42154);49 samelocality,unda l2.ii.Pwo. iS'wis). lalrymple Rd. similarto W. major. Legs 142", 17.1; it5.7; III Fui,-eilaNP2! 02', l48°36',PfJ,9,xi.l99l-29.viJ.I992,P. d5.i1st;aInVce6.b9.etEwpeiegnynanutmer(iFoirg.lo4Bo-pI-o'.f):ilrairsgeeinaitnraiiuimo;n I awlcss,KR.M.Shau (S24808):6\samedata(S24Sil I. duct and anterior margin of atrium is less than DIAGNOSIS. Epigynum wi(h large atrium, halflength ofatrium. Females varied in length anterior loop of insemination dticl just above rram.12-3.8. anterior margin ofatrium (cf. IF seavlew)', tibia I . CW 1.4. AL I 6. AW 1 !. i.ecs with 4 long rctroventral setae forming comb (cf. I -423. I 8.2; II 6.7: III 6.1; IV 8.2. Male palp W. eilint); without proventral keel on embolus. 136 MEMOIRS OF THE QUEENSLAND MUSEUM FIG 5. A. Wabua elliot sp. nov., 6 palp (ventral). B-F, Wabua eimgella sp. nov.; B.C. 2; B, ALS (1.), C, epigynum; D-F, 6 palp, conductorembolus(prolateral, rctrolateral,ventral). NEW SPIDER GENUS FROM THE WETTROPICS 137 FIG. 6. A-E, Wahua crediion sp< nov.: A, $ palp (ventral); B-E, epigynum (ventral, ventral cleared, dorsal, lateralI. F-K, tytibutiabenlccii sp. no\.:_ F-I. epigynum (ventral, ventral cleared, dorsal, lateral); J, K., 8 palp (ventral,retrolateral). DESCRIPTION. Female. CL 1.9. CW 1.4, AL DISTRIBUTION. (Fig. 10B) Occurs in rain- 2.0. AW 1.3. Legs 1423. I 8.1: II 6.4; III 5.8; TV forestsofthenorthernpartoftheEungellaportion 7.9. Epigynum (Figs 4F-I, 5C): broad atrium, ofthe Clarke Range, westofMackay. loopofinseminationductclosetoanteriormargin ofatrium. Spinnerets: ALS with 2 majorampul- Wabua crediron Davies sp. nov (Figs 6A-E, 7A-D, I0A) late spigots, about 2(1 piriforms (Fig. 5B). PMS with large anterior spigot (minor ampullate), a MATERIAL. HOLOTYPE: V. Crediton. mid-east Qld large posterior and a large median spigot 2rn'S,148°34'E.920m.litter,14-21.iv.1975.R.Ki.hout, (cylindrical) 3-4 paracribellarspigotsandsome VFD (QM S42195). PARATYPBS: Mid-east Qld: 2<J, aciniformspigots.Short(10setac)proximalcala- samedataasholotyne(S42157); 23 S.samedata, with mistrum. Females varied in length from 3.3-3.9 2eg6g,saBcrforkoemnweRb.sEiunnbgiaeslelsaofKdPe.ad2pLa1l0m".fron1d4s8°(3S04'2.15F6T,): in length. IO.xi.l991-29.viLI992. P, Lawless. RR. M. Shaw Male. CL 1.9, CW 1.5, AL 2.0, AW 1.3. Leas (S24815): i,samedata(S24033J. 1423, 1 8.3; II6.5:III 2.0; IVS.1. Malepalp(Figs DIAGNOSIS. Thedistancebetween theanterior 4.I.K, 5D-F): embolus with smooth curve from margin ofthe atrium and theanteriorloop ofthe origintoprolateraledge;embolic apophysiswith insemination duct is xT atrial length (cf. W. sea- a larger (bulbous) and smaller fold, embolus view). Male retroventral tibial setae forming pointed; tibia with 4 very long ventral setae comb (cf. II: elliol). Curve of embolus with forming a comb; RTA broad. Males varied in marked widening at prolateral edge (el If length from 3.4-?.9. eungella). 138 MEMOIRS OF THE QUEENSLAND MUSEUM FIG 7. A-D, IVabua crediton sp. nov.; A, epigymim; B-D, 5 palp conductor/embolus, (antero-retrolaleral, ventral, prolateral). E-G, Wabuaaberdeensp.nov.; K.F. 5 palp(ventral, prolateral);G, RTAandcomboflong setae.

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