Types of Mecoptera, Megaloptera, Raphidioptera, and Neuroptera (Insecta) in the collection of the California Academy of Sciences

23 Pages·1996·8.2 MB·English

by  Norman D Penny

PROCEEDINGS
OF THE
CALIFORNIA ACADEMY OF SCIENCES
Vol. 49, No. 4, pp. 127-149. April 9, 1996
TYPES OF MECOPTERA, MEGALOPTERA, RAPHIDIOPTERA,
AND NEUROPTERA (INSECTA) IN THE COLLECTION OF
THE CALIFORNIA ACADEMY OF SCIENCES
By
Norman D. Penny and Vincent F. Lee BtologfealUbacateiy/
ttfiKM^
DepartmentofEntomology, CaliforniaAcademy of '
Golden Gate Park, San Francisco, California 94118
APR 2 2 1996
Abstract: Completelabel information ispresentedforall primary typespecimens,allotypes,
and neallotypes of Mecoptera, Megaloptera, Neuroptera, and Raphidioptera housed in the
CaliforniaAcademyofSciences.Numbersofnon-allotypkparatypesaregivenwiththenames
ofstates or provinces ofthe North American species, or countries ofother nations.
Received March 28, 1995. Accepted August23, 1995.
Introduction California Academy of Sciences now contains
one of the ten largest collections of neu-
TheCaliforniaAcademyofSciencescontains ropteroid types in the world. Currently there
oneofthemajorentomologicaltypecollections are 9 holotypes, 4 allotypes, and 64 paratypes
in the world. Although a rather young collec- of Megaloptera; 7 holotypes, 3 allotypes, and
tion, it has been growing very actively since 89 paratypes ofRaphidioptera; 77 holotypes, 2
its 1915 reopening in Golden Gate Park after lectotypes, 10 allotypes, 1 neallotype, and 190
San Francisco's great earthquake and fire of paratypes of Neuroptera; and 12 holotypes, 3
1906. Since that time it has become the largest allotypes, and 33 paratypesofMecoptera in the
collectionoftypespecimensinthewesterncon- collection. Among those researchers actively
tinental North America, at present including contributing to trie neuropteroidandMecoptera
more than 14,000 primary type specimens. In type collection during the past 30 years are:
order to inform the scientific community as to PhillipA. Adams(California),HorstandUlrike
what types exist in this collection, a series of Aspock (Austria), George W. Byers (Kansas),
catalogs has been published. Previous parts of Elvin D. Evans (Michigan), Jason G. H. Londt
this series include type catalogs ofthe Orthop- (South Africa), Martin Meinander (Finland),
tera (Rentz, 1965), Thysanoptera (Arnaud and Victor J. Monserrat (Spain), Waro Nakahara
Lee, 1973), and Diptera (Arnaud, 1979). (Japan), John D. Oswald (Washington, D.C),
Thereare noprimarytypes ofMecopteraand Norman D. Penny (California), Lionel A.
Neuropterida in this collection predating the Stange (Florida), and Catherine A. Tauber
1906 earthquake. In fact, the first neuropteroid (New York).
type in the collection was described in 1936. Primary types (holotypes, lectotypes, neo-
However, through the efforts of several con- types, and syntypes) are defined as those type
temporary neuropterists (sensu lato), the specimens which are the name-bearing speci-
[127]
128 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES
Volume49,No.4
mens of the species and are used to definitely dicatedby*ratherthan the quotemarks " used
relate the name of the species to an individual to enclose label information.
with certain recognizable traits. Secondary We have tried to maintain the style of pre-
types (allotypes, neallotypes, paratypes, etc.) vious type catalogs. The information contained
are those type specimens used to delimit the on labels is transcribed as closely as possible
phenotypicvariationwithinaspecies. Although totheoriginalprint,althoughnotalllabelscon-
ofmarginal value in determining the true iden- tain neat, linearlines ofinformation, and occa-
tityofaspecies, thesesecondary typesareuse- sional placement interpretation was needed.
ful to some researchers and because we wish Each lineisdelimitedby slash marks,andeach
to maintain consistency in content within the label by quotation marks and a semicolon. The
series, these secondary types have been in- journal abbreviations in the text generally con-
cluded.Wehavelistedthenumberofparatypes, form to those used by BIOSIS, although we
aswellascountryoforigin,andinNorthAmer- prefer a more abbreviated listing for Psyche
ica the state or province of origin. Where a (Camb.).IntheLiteratureCitedsection,journal
paratype locality is found in a country deemed titles have been spelled out fully. The number
to have complex faunal relationships, such as ofparatypesindicatedis exclusive ofany spec-
Ecuador(GalapagosIslands)andIndonesia,we imen designated as allotype or neallotype.
haveprovided moreprecise information. There
is some confusion about the definition of MECOPTERA
neallotypes, as opposed to neoallotypes.
Neallotype is herein defined as an allotype Bittacidae
which was not part of the original type series,
but subsequently designated as allotype. Until Bittacus rossi Londt.
adoptionofthe 1985editionoftheInternational 1977, J. Entomol. Soc. South. Afr., vol. 40,
CodeofZoologicalNomenclature(Recommen-
no. 1, p. 94.
dation 72A), there had been no formally ac- Holotype (male), CAS Type No. 16648; "B.
cepteddefinition ofallotype,andsomeworkers CONGO: / 5 mi. S. of / Fizi, 1320m / 1-10-
preferredtodesignateallotypesfromspecimens 1958"; "E.S. Ross & / R.E. Leech / collec-
other than the original paratype series, so the tors"; "HOLOTYPE / Bittacus / rossi [MA] /
term neallotype was coined to separate these LONDT. J.G.H."
specimens from allotypes designated from the
original type series. Bittacus smithersi Londt.
Primary type specimens in the CAS collec- 1972, J. Entomol. Soc. South. Afr., vol. 35,
tion are deposited through several different ar- no. 2, pp. 328-329, figs. 31, 32, 49, 62.
rangements. Types from theUniversity ofCal- Paratypes, 2 (South Africa).
ifornia,Riverside,andOregon StateUniversity,
Corvallis, are entrusted to The Academy
Boreidae
through a permanent deposit arrangement. The
University of California, Berkeley, currently
Boreus brevicaudus Byers.
has an indefinite loan arrangement with the
1961, J. Kans. Entomol. Soc., vol. 34, no. 2,
Academy. If not otherwise indicated, the in-
cluded types are thepermanent property ofthe p.H7o4l.otype(male),CASTypeNo.9440;"Ore.,
California Academy of Sciences. All pinned
Lane Co., / 2mi. S. Eugene, / Spencer Butte
specimens and any associated parts separated
area, / east side, Nov. 27, / 1959, David R.
fAAlrcsoaomd,etahmlelybs/olidodyefshSahcvaieveenacaelsasbi/emlilrTaeyrapdCeinA/gSN"oCt.aylpi/efo.srl.niid.ae" /SmBAioltrlheo"ut;sy"p/embo(rsfesevmifacrlaeou)md;u/s"oOl/rdeG.et,roeLre"ag;ene"WH.COo.BL,yOe/rTs3Y".mPiE.
label attached. Where quoted label information
contains amale or female symbol, we have in- S. Eugene, / Spencer Butte, / Nov. 23, 1959 /
dicatedthiswith [MA] and [FE] letterandsym- J".ADL.LLOatTtYinP";E"/mBoosrseuasnd/b/relevaifcaluitdtuesr //G1e40o0r"g'e;
bol combination. Further, where quoted label
W. Byers".
information contains ditto marks, these are in-
PENNYANDLEE:TYPECOLLECTIONS 129
Remarks: Both specimens are on permanent Neopanorpa lungtausana Cheng.
depositfromOregonStateUniversity.Thisspe- 1957,Mem.Coll.Agric,Natl.TaiwanUniv.,
cies is currently placed in the genus Hespero- vol. 5, no. 1, pp. 32-33, figs. 14, 15.
boreus (see Penny, 1977:184). Holotype (female), CAS Type No. 7212;
"Kwangtung Prov., S. China / Lung-tau Shan.
Boreus elegans Carpenter. AIL 600M. / near Yiu Vill. above Tso- /kok-
1935, Psyche, vol. 42, no. 2, p. 119. wan. VI-6-47 Gressitt"; "Neopanorpalungtaus-
Holotype(male),CAS TypeNo.4529;"Van- ana n.sp. / By F. Y. Cheng"; "HOLOTYPE
couver, B. C./XII-1930"; "H.E. Hinton/Col- [FE]".
lector"; 'Type"; "Boreus elegans / Carp."
Neopanorpa mokansana Cheng.
Boreus nix, Carpenter. 1957,Mem.Coll.Agric,Natl.TaiwanUniv.,
1935, Psyche, vol. 42, no. 2, p. 114, fig. 7. vol. 5, no. 1, pp. 30-32, figs. 9-11.
Paratype, 1 (Montana). Holotype (male), CAS Type No. 7213
"MokansanChina/CheKiang Pr./IX-19-27"
Boreus notoperates Cooper. "Mrs. Dora / E. Wright / Collector"
1972, Psyche, vol. 79, no. 4, pp. 270-281, "Neopanorpa mokansana n.sp. / By F. Y.
figs. 1-11. Cheng"; "HOLOTYPE [MA]".
Paratypes, 2 (California).
Remarks: This species is currently placed in Neopanorpa tuberosa Byers.
the genus Hesperoboreus (see Penny, 1965, Pac. Insects, vol. 7, no. 4, p. 737.
1977:185). Holotype (male), CAS Type No. 8903;
"THAILAND: / Khao-Yai Nat. Pk. / 750m:
Boreuspilosus Carpenter. VTI-26-62"; "E. S. Ross / D. Cavagnaro / Col-
1935, Psyche, vol. 42, no. 2, pp. 114-115, lectors";"HOLOTYPE/Neopanorpa/tuberosa
fig. 10. / [MA] / George W. Byers".
Paratype, 1 (British Columbia). Allotype (female); same dataas holotype.
Paratypes, 24 (Thailand).
Panorpidae
Panorpa kellogi Cheng.
Neopanorpa globulifera Byers. 1957,Mem.Coll.Agric,Natl.TaiwanUniv.,
1983,Pan-Pac. Entomol. (1982), vol. 58, no. vol. 5, no. 1, pp. 29-30, figs. 5, 6.
2, pp. 92-95, figs. 1-9. Holotype (male), CAS Type No. 7214;
Holotype (male), CAS Type No. 15901; "Foochow / China"; "C.R. Kellog [sic]/ Col-
"LAOS Sam Thong / Xieng Khouang Prov / lector"; 'Tanorpa kellogi [sic] n.sp. / By F. Y.
90mi NE Vientiane / 12 May 1968 / Gary L Cheng"; "HOLOTYPE [MA]".
Peters"; "HOLOTYPE / Neopanorpa / Remarks: The original description indicated
globulifera / [MA] / George W. Byers". that this species was named for the collector
Allotype (female)(in alcohol); same data as "C.R. Kellog,"amisspellingofClaudeR.Kel-
holotype,exceptdated"28July 1968"; withad- logg. Since there was no internal evidence of
ditional label reading "specimen dissected; / the correct spelling within the article, the in-
watch for loose parts!". correct original spelling must be retained.
Paratype, 1 (Laos).
Panorpa mokansana Cheng.
Neopanorpa lichuanensis Cheng. 1957,Mem.Coll. Agric,Natl.TaiwanUniv.,
1957,Mem.Coll. Agric,Natl.TaiwanUniv., vol. 5, no. 1, p. 27, figs. 1, 2.
vol. 5, no. 1, p. 32, figs. 12, 13. Holotype (male), CAS Type No. 7215;
Holotype (female), CAS Type No. 7211; "MokansanChina/CheKiangPr./DC-19-27";
"Suisapa, 1000M./LichuanDistr./W. Hupeh, "Mrs. Dora/E. Wright / Collector"; 'Tanorpa
China/VHI-21-48";"Neopanorpalichuanensis mokansana n.sp. / By F.Y. Cheng"; "HOLO-
n.sp. / By F.Y. Cheng"; "HOLOTYPE [FE]". TYPE [MA]".
Remarks: The original description indicated
PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES
130
Volume49, No.4
that the holotype was a female, but it is in fact Remarks: This species is currently placed in
a male. Orohermes.
Panorpa thompsoni Cheng. Dysmicohermes ingens Chandler.
1957,Mem.Coll.Agric,Natl.TaiwanUniv., 1954, Pan-Pac. Entomol., vol. 30, no. 2, pp.
vol. 5, no. 1, p. 29, figs. 7, 8. 105-107, figs. 3-5.
Holotype (female), CAS Type No. 7216; Holotype (male), CAS Type No. 7198;
'Tsu Shima / Lu Chu Ids. / X-5-1910"; "JC "Miami Ranger Sta. / Mariposa Co. Cal. /
Thompson / Collector"; "Panorpa thompsoni 7/27/46"; "H. P. Chandler / No. 67 Expo. / /
n.sp. / By F.Y. Cheng"; "HOLOTYPE [FE]". /4 NE /Elv 5,000 SW"; "HOLOTYPE / Dys-
micohermes / ingens".
Panorpa wrightae Cheng. Allotype (female); samedataasholotype,ex-
1957,Mem.Coll.Agric,Natl.TaiwanUniv., cept2ndlineof2ndlabelreads"No. 61 Expo.",
vol. 5, no. 1, pp. 28-29, figs. 3, 4. andholotypelabelisreplacedbyallotypelabel.
Holotype (female), CAS Type No. 7217; Paratypes, 4 (California).
"Mokansan China / Che Kiang Pr. / Sep. 28
1927"; "Mrs. Dora / E. Wright / Collector"; Neohermes nigrinus Van Dyke.
"Panorpawrighti [sic],n.sp./ByF. Y. Cheng"; 1944, Pan-Pac. Entomol., vol. 20, no. 3,
"HOLOTYPE [FE]". p. 110.
Remarks: Although the original description Holotype (male), CAS Type No. 5430;
of this species bore a masculine ending "Richardson's/Spr., Butte/Co, Calif./ V-12-
(wrighti), the authorclearly stated that the spe- 44";"Presentedby/ECVanDyke/Collector";
cies was named for Mrs. Dora E. Wright, the "Holotype Neohermes / nigrinus / Van Dyke".
collector. Thus, the name should have a femi- Remarks: Thisspeciesiscurrentlyconsidered
nine ending, which is given here. ajunior synonym of Protochauliodes minimus
(Davis).
Panorpodidae
Protochauliodes aridus Maddux.
Brachypanorpa sacajawea Byers. 1954, Pan-Pac. Entomol., vol. 30, no. 1, pp.
70-71.
pp.1929101,-J2.1K7a,nfsi.gsE.n1t-o1m2o.l. Soc., vol. 63, no. 2, Holotype (male), CAS Type No. 7201;
Paratypes, 2 (Idaho). "Chico / R 5-10-51 / D. Maddux"; "HOLO-
TYPE PROTOCHAULIODES ARIDUS
/ / /
MADDUX"; "N.sp. /HOLOTYPE/ [MA] #20
MEGALOPTERA / CSC #1".
Protochauliodes cascadius Evans.
Corydalidae
1984, Pan-Pac. Entomol., vol. 60, no. 1, pp.
1-3, figs. 1-3.
Dysmicohermes crepusculus Chandler. Holotype(male),CAS TypeNo. 11404; "Ca.
1954, Pan-Pac. Entomol., vol. 30, no. 2, pp. 8mi W. Mill City / along Little Santiam R., /
107-108, figs. 1, 2, 6. Marion Co., Ore. / VII-26-63"; "[MA] [FE]
Holotype (male), CAS Type No. 7197; "Pyr- Protochauliodes montivagus / Sept. 1963
a"Jm.iWd.R.MaS.cS/wEalidnor/aCdoollCeoc.tor/"C;al"iHfOVLIIO-T20Y-P52E";/ CThOaCndHlAeUrL?I"O;D"EHSO/LCOATSYCPAEDI[AMA1]970//EPlRwOi-n
Dysmicohermes / crepusculus". D. Evans".
Allotype(female);"2mi s.w./MiamiRanger Allotype(female); samedataasholotype,ex-
Sta. / Mariposa Co. /7/25/1946"; "H.P. Chan- cept allotype label.
dler / No. 54 Expo. / / / 4 NE / Elv 5,000"; Paratypes, 7 (Oregon).
"ALLOTYPE / Dysmicohermes / crepus-
culus".
Paratypes, 2 (California).
PENNYANDLEE:TYPECOLLECTIONS 131
Protochauliodes montivagus Chandler. RAPHIDIOPTERA
1954, Pan-Pac. Entomol., vol. 30, no. 2, p.
Ill, figs. 9, 10. Inocellidae
Holotype (male), CAS Type No. 7199; "St.
Charles Hill / Sierra Co Cal. / July 7, 1923"; Negha meridionalis Aspock.
"Pies, by / E.H. Nast / Collector"; "HOLO- 1988, Z. Arbeitsgem. Osterr. Entomol., vol.
TYPE /Protochauliodes / montivagus". 39, nos. 3/4, pp. 107-111, figs. 1-11.
Paratypes, 2 (California). Paratype, 1 (Baja California).
Protochauliodes simplus Chandler.
Raphidiidae
1954, Pan-Pac. Entomol., vol. 30, no. 2, pp.
110-111, figs. 7, 8. Agulla banksi Carpenter.
Holotype(male),CAS TypeNo.7200; 'Tan- 1936,Proc. Am. Acad. Arts Sci., vol. 71, no.
/baJrDkPHaastch/kLeo/sCAolnlgeeclteosr";Co".H/OCLalOifT.YVPIEI-/13P-r5o-0 2,Ppa.ra1t2y0p,esp,i.22(C(failgisf.or1n1i,a).12).
tochauliodes / simplus". Remarks: This species is currently consid-
Paratypes, 3 (California). ered ajunior synonym ofAgulla modesta Car-
penter.
Sialidae
Agulla bractea Carpenter.
Sialis arvalis Ross. 1936,Proc. Am. Acad. Arts Sci., vol. 71, no.
1937, 111. Nat Hist Surv. Bull., vol. 21, 2, pp. 130-131, pi. 2 (fig. 16).
no. 3, p. 68. Paratypes, 2 (California).
Holotype (male), CAS Type No. 4647;
"MOKEL. HILL, / CAL."; "coll. by F.E. / Agulla directa Carpenter.
BlaisdeU"; "Apr."; "HOLOTYPE/Sialis [MA] 1936,Proc. Am. Acad. Arts Sci., vol. 71, no.
/ arvalis / Ross". 2, pp. 124-125, pi. 1 (fig. 3).
Paratypes, 2 (California). Holotype (male), CAS Type No. 4475; "Or-
oville Cal. / IV-17-1928"; "H H Keifer / Col-
Sialis mohri Ross. lector"; 'TYPE"; "Agulla directa Carpenter /
1937, 111. Nat. Hist Surv. Bull., vol. 21, no. det F.M. Carpenter"; "H. & U. Aspock vid.
3, p. 74. 1967". Separate pin with genitalia vial, label
Paratypes, 11 (Illinois). reads "Agulla directa / Carp.".
Paratype, 1 (California).
Sialis occidens Ross.
1937, 111. Nat. Hist Surv. Bull., vol. 21, no. Agulla nigrinotum Woglum and McGregor.
3, p. 69. 1964, Bull. South. Calif. Acad. Sci., vol. 63,
Holotype (male), CAS Type No. 4648; "Se- pt 4, pp. 201-203.
quoia Nad. /Park, Calif. / Alt 7000-9000ft."; Holotype (male), CAS Type No. 9548; "Mill
"Wolverton / VI-25-29"; "Pres. by / EC Van Potrero / Kem 7-1-63 / r. s. woglum col";
Dyke/Collector"; "HOLOTYPE/Sialis [MA] "Agulla / negrinotum [sic] / Type on HOLO-
/ occidens / Ross". TYPE"; "Agulla nigrinotum / Woglum et
Allotype(female); samedataasholotype,ex- McGregor/Holotypus [MA] /U. Aspock vid.
ceptallotypelabelsubstitutedforholotypelabel 1972.".
andadditionallabelreading"FromColl./Calif. Allotype (female); "Mill Potrero / Kem 7-
Acad. Sci." 1-63 /r. s. woglum col"; "ALLOTYPE"; "AL-
Paratypes, 6 (5 California, 1 Nevada). LOTYPE / Agulla / nigrinotum / Wog. &
McGreg."
Sialis velata Ross. Paratypes, 7 (California).
1937, 111. Nat. Hist Surv. Bull., vol. 21, no.
3, p. 71. Agulla occidentis Carpenter.
Paratypes, 27 (Michigan). 1936,Proc. Am. Acad. Arts Sci., vol. 71, no.
132 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES
Volume49,No.4
2, pp. 129-130, pi. 2 (fig. 21). /VI-10-1953"; "mediumwith genitalia/water-
Holotype (male),CAS TypeNo. 4476; "Pine soluble"; "Holotypus / Raphidia / (Agulla) /
Crest [sic], Calif. / July Johnson"; "Type"; arnaudi / U. Aspock 1973".
"Agulla occidentis Carpenter / det. F.M. Car- Allotype(female); samedataasholotype,ex-
penter"; "Agullaherbsti/Esb.-Pet/U. Aspock cept that allotype label reads: "PARATYPUS
det. 1967". Separate pin with genitaliavial, la- /(ALLOTYPUS) [FE]/R. (Agulla)/arnaudi";
bels read "Agulla herbsti: Esb.-Pet."; "part of "U. Aspock 1973".
Holotype / Agulla/ occidentis / Carpenter". Remarks: Abdomens ofthe holotype and al-
Allotype (female); "Pine Crest [sic], Calif. lotype are mounted on cards attached to the
/ Aug. 15 Johnson"; "ALLOTYPE"; "Agulla respective pins.
occidentis Carpenter / det F.M. Carpenter";
"O. S.C. TYPE". Raphidia (Agulla) barri Aspock.
Paratypes, 3 (1 California, 2 Oregon). 1973, Pol. Pismo Entomol., vol. 43, no. 2, p.
Remarks: The holotype and allotype are on 238.
permanent deposit from Oregon State Univer- Holotype (male), CAS Type No. 12698;
sity. "WestgardPass/InyoCo./CALIF.VI-11-69";
"beating / Pinyon Pine"; "W.F. Barr/ Collec-
Agulla singularis Carpenter. tor"; "Holotypus [MA] /R. (Agulla) barri /U.
1936,Proc. Am. Acad. Arts Sci., vol. 71, no. Aspock 1973". Separate pin with genitalia
2, p. 132, pi. 2 (fig. 23). vial; label reads "genital segments of / R
Holotype(male),CAS TypeNo.4477; "Ban- (Agulla) barri / Holotypus [MA] ".
ning Cal / V-29-28"; "Pres by / EC Van Dyke Paratypes, 6 (5 California, 1 Nevada).
/ Collector"; "From Coll. / Calif. Acad. Sci.";
"TYPE"; "Agulla singularis Carp. / det. F.M. Raphidia (Aliaberaphidia) tenochtitlana
Carpenter"; "H. & U. AspSck vid. 1970". Sep- Aspock and Aspock.
arate pin with genitalia capsule, labels read 1978, Z. Arbeitsgem. Osterr. Entomol., vol.
"Banning, CAL. / V-29-28 / E. C. Van Dyke / 30, nos. 1/2, p. 21.
collector"; "Dissection of / TYPE, Agulla / Holotype (male), CAS Type No. 13336;
singularis / Carpenter". "MEXICO: BAJA CALI-/FORNIA SUR: Si-
erra La / Laguna, 1770-1850m / 30-VHI-77 /
Agulla unicolor Carpenter. E.Fisher,R.Westcott/Calif.Acad. Sci. Coll.";
1936,Proc. Am. Acad. Arts Sci., vol. 71, no. "HOLOTYPUS [MA] /Raphidia (Aliaberaphi-
2, pp. 121-123, pi. 1 (fig. 1). dia) / tenochtitlana/ Aspock et Aspock 1978";
Paratypes, 60 (39 California, 8 Nevada, 10 "2". Separate pin with genital capsule label
Oregon, 3 Utah). reads "2"; "HOLOTYPUS"; "Raphidia / ten-
ochtitlana / [MA] / H. et U. Aspock det 19".
Raphidia arizonica Banks. Paratypes, 7 (Baja California Sur).
1911, Trans. Am. Entomol. Soc., vol. 37, no.
4, p. 338, pi. 11 (fig. 8).
Neallotype (female), CAS Type No. 4501; NEUROPTERA
"Mono Lk / Cal. / VM7-'17"; "Donor / C.L.
Fox / Collector"; "ALLOTYPE"; "Agulla Berothidae
arizonica (Banks) / deL F. M. Carpenter".
Remarks: Carpenter (1936:118-119) desig- Hemerobius hamulus Walker.
nated the neallotype. 1853, List of Specimens ofNeuropterous In-
sects in the Collec—tion of the British Museum.
Raphidia (Agulla) arnaudi Aspock.
Part II. (Sialidae Nemopterides), p. 278.
1973, Pol. Pismo Entomol., vol. 43, no. 2, p. Neallotype (female), CAS Type No. 5592;
234H.olotype (male), CAS Type No. 12458; "'PTraesm.pabyFl/aE/.MP.ayVa2n, D'u08ze/eV/auCol[lseicc]toDru"z;e"ea"l;-
"MEX: B. Calif., Sierra / San Pedro Martir / lotype"; "Lomamyia / hamata Walker / det
P. H. Amaud,Jr.";'Trail,La/JoyatoLaZanja F. M. Carpenter".
PENNYANDLEE:TYPECOLLECTIONS 133
Remarks: Carpenter (1940:265-266) desig- Christobal,/9mi. E.,Chiapas/MEX., VU-30-
nated the neallotype. 57"; "AT/ light"; "J. A. Chemsak/ B.J. Don-
nells / Collectors"; "Meleoma / antennensis
Lomamyia latipennis Carpenter. Tauber / det C. Tauber '68 / HOLOTYPE".
1940,Proc. Am. Acad. Arts Sci., vol. 74, no. Remarks: The holotype is on indefinite loan
7, pp. 262-263, fig. 64A-D. from the University of California, Berkeley.
Holotype (male), CAS Type No. 5590;
"Phoenix Lk. / Marin Co Cal. / VH-4-1927"; Meleomapowelli Tauber.
"H.H. Keifer/ Collector / Type"; "Lomamyia 1969, Univ. Calif. Publ. Entomol., vol. 58,
/ latipennis Carp. / det F.M. Carpenter". pp. 19-20, figs. 4, 17, pi. 2a.
Allotype (female); "Mt. Tamalpais / Marin Holotype (male), CAS Type No. 10006;
Co. Cal. / V-23-09"; "Coll. by / E.C. Van "MEX: Dgo., El / Salto, 3 mi. W. /9000' VII-
Dyke"; "From Coll. /Calif. Acad. Sci."; "allo- 23-64"; "Quercus / omissa"; "J. Powell / Col-
type"; "Lomamyia / latipennis Carp. / det lector"; "Meleoma / powelli Tauber / det. C.
F. M. Carpenter". Tauber '68 / HOLOTYPE".
Allotype (female); "MEX: Dgo., El / Salto,
Manselliberotha neuropterologorum Aspock 3 mi. W. / 9000' VH-23-64"; "J.A. Chemsak
and Aspock. & / J. Powell Black / & White lights";
1988, Z. Arbeitsgem. Osterr. Entomol., vol. "Meleoma / powelli Tauber / det. C. Tauber
40, nos. 1/2, pp. 4-7, figs. 1-21. '68 / ALLOTYPE".
Paratype, 1 (Namibia). Remarks: These types areon indefinite loan
from the University of California, Berkeley.
Chrysopidae
Nodita oenops Adams.
Leptochrysaprisca Adams and Penny. 1987, Neuropt Int, vol. 4, no. 4, pp. 290-
1992, Pan-Pac. Entomol., vol. 68, no. 3, pp. 291, figs. 2, 19-24.
219-221, figs. 10-14. Holotype (male), CAS Type No. 16027;
Holotype (female), CAS Type No. 16743; "May 2, 1976 3 /Leon, Nica / W. Barclay ";
"PERU: Dept. Amazonas/ 18km N. ofPuente "HOLOTYPE / Nodita [MA] / oenops / P.
/ Ingenio, KM 320 Alt. / 1750 m. 9-X-1964 / Adams".
P.C. Hutchison & / J.K. Wright"; "Collected Allotype (female); "4/20/86 / Leon, Nica /
on / Baccharis latifolia I #6830"; "HOLO- W. Barclay"; "ALLOTYPE / Nodita / oenops
TYPE / Leptochrysa / prisca / Adams & [FE] / P. Adams".
Penny". Gut contents in vial of glycerin on
pin; label reads "Leptochrysa/prisca / Adams Pimachrysa albicostales Adams.
- Penny / gut contents". Genitalia in vial of 1967, Bull. Mus. Comp. Zool., Harvard
glycerin on pin; label reads "Leptochrysa / Univ., vol. 135, no. 4, pp. 226-227, figs. 27,
prisca / A & P / [MA] genitalia". 28.
Holotype (male), CAS Type No. 9988;
Leucochrysa serrula Adams. "MEX: B. Calif. / 22mi. N. of/ Punta Prieta
1979, Folia Entomol. Mex., no. 41, pp. 97- / 9-XII-1958"; "H. B. Leech / Collector";
10H1,olfoitgsy.pe1-(7m.ale), CAS Type No. 15126; "3 C[bHlaRnYkSlAabe/l AwLiBthICOyeSlTloAwLEsStrip/e]A;D"APMISMA-/
mi.east/Carapan/MichMEX./VII 10 1963"; [MA] TYPE/Det P. Adams"; "13". Genitalia
"F.D.Parker/L. A. Stange/Collectors";"HO- on separate pin with label "13".
LOTYPE/Leucochrysa/serrula/[MA] Adams
/ det. P. Adams 77". Pimachrysafusca Adams.
1967, Bull. Mus. Comp. Zool., Harvard
Meleoma antennensis Tauber. Univ., vol. 135, no. 4,p. 227, figs. 1,7, 11-13,
1969, Univ. Calif. Publ. Entomol., vol. 58, 21, 24.
p. 20, figs. 4, 18, pi. 2b. Holotype (male), CAS Type No. 9989;
Holotype(male),CASTypeNo. 10005; "San "CALIF., #1 /Joshua Tree N.M., /L. Coving-
134 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES
Volume49,No.4
ton Flat"; "E.L. Sleeper / Collr."; "PIMA- capsule on pin.
CHRYSA FUSCA ADAMS HOLOTYPE
/ / /
/ Det. P. Adams"; "14". Genitalia on separate Aleuropteryx maculipennis Meinander.
pin with label "14". 1972, Acta Zool. Fenn., no. 136, p. 45, fig.
13A-D.
Pimachrysa nigra Adams. Holotype (male), CAS Type No. 11915;
1967, Bull. Mus. Comp. Zool., Harvard "CAL.: Kern Co / 19mi. NNE. Mojave / IV-
Univ., vol. 135, no. 4, pp. 228-229, figs. 6, 14-1962"; 44C. MacNeill, D. / Rentz & R.
14-16, 26, 34. Brown / C—oll—ectors"; "Genitalia CAS slide 13
Holotype (male), CAS Type No. 9990; / Wings * 38"; "Aleuropteryx / maculi-
"Gavilan / Cal. Mr. 19. 36"; "on Ceanothus / pennis n.sp. [MA] / det. Meinander 1969".
crassifolius"; "Timberlake / Coll."; "No- Slide 38 left label "Cal. Kern Co./ 19 mi NNE
thochrysa / californica / Inif. det. Banks"; Mojave / IV-14-1962 / C. MacNeill, D. Rentz
'TIMACHRYSA / NIGRA / ADAMS / HO- / & R. Brown coll. / Body pinned / genitalia
LOTYPE/Det. P. Adams"; "10". Genitaliaon slide 13 / CAS slide 38"; right label "Al-
separate pin with label "10". europteryx / maculipennis / n.sp. [MA] /
HOLOTYPUS/det. 1969/MartinMeinander".
Coniopterygidae Slide 13 left label "CAS 13 / Cal. Kern Co /
19 mi NNE Mojave / IV-14-62 / MacNeill,
Aleuropteryx cupressi Meinander. Rentz/Brown /body pinned"; right label "Al-
1974, Entomol. Scand., vol. 5, nos. 3/4, p. europteryx / maculipennis / n.sp. M/
218, fig. 1A-H. HOLOTYPUS [MA] / det. 1966-1969 /
Holotype (male), CAS Type No. 12352 (in Meinander".
alcohol); "USA. Cal. Marin Co. / 1961-05-02
emgd from/Cupressus goveniana/J. Powell"; Aleuropteryxpunctata Meinander.
"HOLOTYPUS/Aleuropteryx/cupressi; sp.n. 1974, Entomol. Scand., vol. 5, nos. 3/4, p.
[MA] / det. Meinander 1973"; "Wings slide 220, fig. 3A-H.
UCAA3". SlideUCAA3 leftlabel"USACal. Holotype (male), CAS Type No. 12193 (in
/ Marin Co / emgd 1961-05-02 /ex Cupressus alcohol); "20mi. W. Rosario, / Sin., Mex. I-
goveniana / J. Powell"; right label "Al- 30-64 / at night; E. Schlinger / black light";
HeuOrLopOteTrYyxPU/Sc/updreetssi197[3MA/]M/MeMieniannadenrd"e.r / e"u[rMoAp]terwyixn/gpsunscltiadtea5M"e;in"H[MOAL]OT2Y[PMEA]/[MAAl]-
Remarks: The holotype is on indefinite loan /det Meinander 1973"W. Slide UCR5 left label
from the University of California, Berkeley. "Mexico Sin. / 20mi Rosario / 1964-01-30
/ E. Schlinger / Slide UCR5"; right label "Al-
Aleuropteryx longipennis Meinander. europteryx/punctata/ MMeinander [MA] /HO-
1974, Entomol. Scand., vol. 5, nos. 3/4, pp. LOTYPE / det. 1973 / Meinander".
218-219, fig. 2A-D. Remarks: The holotype is on permanent de-
Holotype (male), CAS Type No. 12353; posit from the University ofCalifornia, River-
"Mex: Baja Calif. / Norte, 1 mi. N. / Meling side.
Ranch, 2100' / 111-17-1972, J. Doyen / & J.
Powell, at light"; "Wings slide UCA A4"; Aleuropteryx simillima Meinander.
"HOLOTYPUS / Aleuropteryx [MA] / longi- 1972, Acta Zool. Fenn., no. 136, pp. 46-47,
pennis sp.n. / det. Meinander 1973". Slide fig. 14A-E.
UCA A4 left label "Mexico Baja Calif/Norte Holotype (male), CAS Type No. 11916;
1 mi N / Meling Ranch / 2100' 1972-03-17 / "MEX.: Baja Calif. /Norte, San Felipe / 7-III-
J. Doyen & J. Powell"; right label "Al- 1963 / P.H. Am—au"d—, Jr."; "Genit slide CAS
europteryx / longipennis [MA] / sp.n. / det. 20 / wings 39"; "Aleuropteryx /
1973 / M Meinander". simillissimus n.sp. [MA] /det Meinander 1969
Remarks: The holotype is on indefinite loan / HOLOTYPUS". Slide 20 left label "CAS 20
from the University of California, Berkeley. / Mex: Baja Calif / Norte San Felipe / 7-III-
The body is stored in glycerin in a genitalia 1963 /P. H. Arnaud Jr/ Body pinned / Wings
PENNYANDLEE:TYPECOLLECTIONS 135
slide 39"; right label "Aleuropteryx / sim- alcohol); "Oso Flaco Lake / San Luis Obispo
illissima / n.sp. / HOLOTYPUS [MA] / det. Co. / Cal. VIJJ-5-62 / E. Schlinger"; "HOLO-
1966-1969/MMeinander". Slide 39 leftlabel TYPUS / Coniopteryx [MA] / californica
"Mexico / Baja Calif. / Norte San Felipe / Meinander/ det. Meinander 1973".
7.III.1963 / P. H. Arnaud Jr. / Body pinned /
genit slide20/CAS Slide39"; rightlabel "Al- Comopteryx (Scotoconiopteryx) chilensis
europteryx/similkssima/n.sp./HOLOTYPUS Meinander.
/ det. 1969 / Martin Meinander". 1990, Acta Zool. Fenn., no. 189, pp. 47-48,
Remarks: Thesudelabelsread"simillissima" fig. 16A-D.
or "simillisimus" rather than the published Holotype (male), CAS Type No. 16102 (in
spelling. alcohol); "Chile: Chilian / Las Trancas / 1200
/ 17-20-XH-83 / Luis Pefia"; "Coniopteryx /
Aleuropteryx unicolor Meinander. chilensis Meinander [MA] / det. Meinander
1972, Acta Zool. Fenn., no. 136, pp. 48-49, 1987 / HOLOTYPUS".
fig. 16A-J\
Holotype (male), CAS Type No. 11917; Coniopteryx (Coniopteryx) goniocera
"BoydDesertRes.Center/4 mi s.PalmDesert, Meinander.
Riv. / Co., Calif. IV-11-63"; "W.A. Steffan / 1972, Acta Zool. Fenn., no. 136, pp. 264-
Collector";"Aleuropteryx/unicolorn.sp. [MA] 265, fig. 169A^.
/ det. Meinander 1969 / wings slide 19 Body Holotype (male), CAS Type No. 11919 (in
si. 20". Slide 20 left label "CalifBoyd Desert alcohol); "S. INDIA/ 10mi. SW./Gudiyattam
/ Res. Center / 4 mi s Palm Desert / Riv. Co. / 350m IV-3-62"; "Collectors: / E. S. Ross /
IV-11-63 / W.A. Steffan / wings slide 19 / D. Q. Cavagnaro"; "genitalia / mounted on /
UCal Slide20";rightlabel"Aleuropteryx/uni- slide"; "genit slide CAS 29"; "[MA] placed in
coMlor n.sp. / [MA] / HOLOTYPUS / det. 1969 / general / collection"; "HOLOTYPE [MA] /
/ Meinander". Slide 19 with same data as Coniopteryx / goniocera n.sp. /det. Meinander
slide 20, except for slide number. 1971". Slide 29 left label "Coniopteryx /
Remarks: The holotype is on indefinite loan goniocera n.sp. / HOLOTYPUS [MA] / det.
M
from the University ofCalifornia, Berkeley. 1971 / Meinander"; right label "S. India /
10mi SW/Gudiyattam/350m IV.3.62/Ross
Coniocompsa smithersi Meinander. & Cavagnaro / CAS 29".
1972, ActaZool. Fenn., no. 136,pp. 99-101,
fig. 51A-G. Coniopteryx (Coniopteryx) mexicana
Holotype (male), CAS Type No. 11918 (in Meinander.
alcohol); "KENYA: 28 mi. NW. / Nairobi 1974, Entomol. Scand., vol. 5, nos. 3/4, pp.
6450 ft / 17-XII-1969 / M.E. Irwin & / E. S. 226-227, fig. 10A-E.
Ross"; "genitalia / & wings / mounted on / Holotype (male), CAS Type No. 12195 (in
slide"; "wings CAS slide 59, genit. 58"; "Con- alcohol); "20mi. E. Villa Union / Sin. Mexico
iocompsa/smithersi n.sp. [MA] /det Meinan- / 1-31-64 / Schlinger & Irwin / at black light";
der 1971 / HOLOWTYPUS". Slide 58 left label "HOLOTYPUS /Coniopteryx/mexicanasp.n.
"Kenya/28 mi [sic] Nairobi/6450ft. / 17- [MA] / det. Meinander 1973".
XH-1969 / M.E. Irwin & / E. S. Ross / CAS
slide 58"; right label "Coniocompsa/ smitherMsi Coniopteryx (Coniopteryx) minuta Meinander.
n.sp. / [MA] / HOLOTYPUS / det. 1971 / 1972, Acta Zool. Fenn., no. 136, pp. 259-
Meinander". Slide 59 with same data as slide 260, fig. 165A^\
58, except for slide number. Holotype (male), CAS Type No. 11920;
"Morro Bay, 8 mi E / S.L. O. Co., / Calif. V-
Coniopteryx (Coniopteryx) californica 2-62"; "Adenostoma/sp. fasciculatum"; "C. A.
Meinander. Toschi/Collector"; "genitalia UCal Slide 10";
1974, Entomol. Scand., vol. 5, nos. 3/4, pp. "HOLOTYPUS / Coniopteryx / minuta n.sp.
225-226, fig. 9A-G. [MA] / det Meinander 1971". Slide 10 left
Holotype (male), CAS Type No. 12194 (in label "USA. Calif. / S.L. O. Co. / Morro Bay
136 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES
Volume49,No.4
/2.V.1962/C. A.Toschi/UCalslide 10";right Remarks: The holotype is on indefinite loan
label "Coniopteryx / minuta n.sp. [MA] / HO- from the University of California, Berkeley.
M
LOTYPE / det. 1971 / Meinander".
Remarks: The label data "8mi E" were not Helicoconis (Helicoconis) similis Meinander.
transcribed to the slide label. The holotype is 1972, Acta Zool. Fenn., no. 136, p. 124, fig.
on indefinite loan from the University of Cal- 65A-C.
ifornia, Berkeley. Holotype (male), CAS Type No. 11923;
"CANADA: B. COL. / Top of Moyie Mtn. /
Coniopteryx {Coniopteryx) simplex elev. 6868 ft/EastKootenays/ 16-VII-1959";
Meinander. "H.B.Leech/Collector";"slidesCAS 16-17";
1974, Entomol. Scand., vol. 5, nos. 3/4, p. "Helicoconis / similis n.sp. / det. Meinander
228, fig. 11A-J7. 1969 / HOLOTYPUS". Slide 16 left label
Holotype (maWle), CAS Type No. 12354; "CAS 16 / Canada B. Col. / Top of Moyle
"PinoGrande,S /ofLakeEdson,/ElDorado [sic] Mtn/elev 6868ft. /East Kootenaya [sic]
Co. / Calif. VH-8-67"; "W.J. Turner/ Collec- / Genit. si. 17 / Body pinned"; right label
tor"; "HOLOTYPUS / Coniopteryx / simplex "Helicoconis / similis n.Msp. [MA] /
sp.n. [MA] / det. Meinander 1974". HOLOTYPUS / DET. 1969 / Meinander".
Remarks: The holotype is on indefinite loan Slide 17 has same label dataas slide 16,except
from the University of California, Berkeley. for slide number.
Body is storedin glycerin ina genitaliacapsule Remarks: Right slide labels overlie identifi-
on pin. cation labels for Helicoconis eglini Ohm.
Conwentzia californica Meinander. Heteroconis bifurcata Meinander.
1972, Acta Zool. Fenn., no. 136, pp. 297- 1990, Acta Zool. Fenn., no. 189, pp. 11-12,
298, fig. 192A-E. fig. 1A-E.
Holotype (male), CAS Type No. 11921; Holotype (male), CAS Type No. 16602 (in
"CAL. Lee Vining / Campgd. Mono Co. / W. alcohol); "PAPUA NEW GUINEA: / MadanWg
ofMonoLk./ l-VII-61 /D.C.RENTZ/COL- Province / Sapi Forest Reserve / (30km
LECTOR"; "CAS slide 56"; "HOLOTYPUS / Madang)/5*12'S, 145*30,E/ 19February 1987
Conwentzia / californica n.sp. [MA] / det. / Norman D. Penny clr"; "Heteroconis /
Meinander 1970". Slide 56 left label "WUSA bifurcata Mein [MA] / det. Meinander 1987 /
Calif/ Mono Co/Lee Vining Campgd/ of HOLOTYPUS"; "Onepairof/wingsonslide".
Mono Lk. / 1.VH.1961 / D.C. Rente / CAS Slide left label "Papua New GuineaW/ Madang
Slide 56"; rightlabel"Conwentzia/californiMca Prov./SapiForestReserve/30km Madang
n.sp. / HOLOTYPUS [MA] / det. 1970 / / 5*12'S 145*3(nE / 19 Febr 1987 /Norman D.
Meinander". Penny clr"; right label "Heteroconis/bifurcata
/ Meinander sp.n. [MA] / HOLOTYPUS".
Helicoconis (Helicoconis) californica
Meinander. Heteroconis curvata Meinander.
1972, Acta Zool. Fenn., no. 136, p. 119, fig. 1990, Acta Zool. Fenn., no. 189, pp. 12-14,
61A-E. fig. 2A-I.
Holotype (male), CAS Type No. 11922 (in Holotype (male), CAS Type No. 16603 (in
alcohol); "Calif: Mono Co. / Tom's Place, 1 alcohol); "PAPUA NEW GUINEA: / Madang
mi./SW. Vffl-10-1963"; "M.J. Tauber/C. A. Province, / Nagada Harbor / 8km N Madang
Toschi / Collectors"; "Genit UCal Slide 18"; / 5'09'S 145'48'E / 8 February 1987 /Norman
"Helicoconis / californica n.sp. [MA] / det. D. Penny"; "Heteroconis / curvata Meinander
Meinander 1969 / HOLOTYPE". Slide 18 left [MA] /det. Meinander 1987 /HOLOTYPUS";
label "USA. Calif. Mono Co. / Tom's Place - "Onepairof/wingsonslide";"genit.in micro-
1 mi SW/Vm-10-1963 / M.J. Tauber/ C. A. / vial". Slide left label "Papua New Guinea /
Toschi / UCal slide 18"; right label Madang Province / Nagada Harbor / 8km N
"[HMeAl]ico/cdoenti.s1/9c6a9li/foMrniMceai/nna.nsdpe.r/"H.OLOTYPE NMaDdaPnegnn/y"5;'0r9i'gSht l1a4b5e,l48"'HeEt/er8ocFoenbirs /1c9u8r7vat/a