Type specimens of recent species of Pectinidae described by Lamarck (1819), preserved in the Muséum d'Histoire Naturelle of Geneva and the Muséum National d'Histoire Naturelle of Paris

68 Pages·1994·33.4 MB·

by  H H Dijkstra

REVUE SUISSE DE ZOOLOGIE, 101 (2) : 465-532; juin 1994
Type specimens of recent species of Pectinidae described by
Lamarck (1819),
preserved in the Muséum d'Histoire Naturelle of Geneva
and the Muséum National d'Histoire Naturelle of Paris
(with 30 plates)
Henk H. DIJKSTRA
University of Amsterdam, Institute of Systematics and Populationbiology
P.O. Box 94766, 1090 GT Amsterdam, The Netherlands.
Types specimens of recent species of Pectinidae described by Lamarck
(1819).- LAMARCK listed and described 60 species of recent Pectinidae (1
Pedum species, and 59 Pecten species) in 1819. Twenty-nine were newly
described species, of which 4 are junior homonyms, 1 senior homonym, 11
junior synonyms, 1 possible junior synonym, 9 extant taxa, and 3 un-
certain. He validated 9 non-binominal species, and one manuscript name of
Péron. One type specimen (P. pellucidus) cannot be found. Nineteen
primary types are present in the MHNG, and nine in the MNHN. Most of
Lamarck's taxa were figured by BRUGUIERE (1797), DELESSERT (1841), and
CHENU (1844-1845, 1862), but seven taxa were not illustrated. Two
lectotypes previously selected by FLEMING (1951) and CLARKE (1965) are
invalid, according to ICZN art. 74 (a)(v), and twenty lectotypes are herein
selected. Five new type localities are also designated for species which
had unknown or erroneous type localities. Seventeen varieties described by
Lamarck as new are excluded from the type series, according to ICZN art.
72 (b)(i).
Key-words: Lamarck - Mollusca (Bivalvia : Pectinidae) - Type species -
Taxonomy - Nomenclature.
INTRODUCTION
In 1793, Jean-Baptiste Pierre Antoine de Monet, Chevalier de Lamarck (1744-
1829), was appointed as Professor in Zoology (Invertebrates) at the Muséum
d'Histoire Naturelle in Paris. At the same time, Lamarck possessed a private
collection of fossil and recent mollusca, which was offered in 1796 to the French
government for 33.000 livres, but this collection was not accepted. In the meantime,
Lamarck described many new molluscan taxa from his private collection and the
Manuscript accepted 10.01.1994.
466 HENK H. DIJKSTRA
collection of the MNHN, including Pectinidae, purchased or collected by French
voyages around the world, among which the famous "Voyage aux Terres Australes"
(1800-1804). In the volume 6 of his "Histoire Naturelle des Animaux sans Vertèbres"
(1819), he described the recent Pectinidae, divided into two groups, i.e. "Auricles
equal or nearly equal" and "Auricles inequal", listing 59 species based on material of
"Mus. n°." [= MNHN] and "Mon cabinet" [= his private collection, now in MHNG].
During the last years of his life, he suffered with blindness, and his daughter Rosalie
assisted him with his work, as indicated in a letter addressed to her sister Cornélie.
After Lamarck's death in 1829, his rich private collection (13.288 species) was
purchased by Prince Masséna, Duke of Rivoli, in 1831. Afterwards, in 1840, Baron
J.P.B. de Lessert (or Delessert) of Lyon acquired Lamarck's collection and joined it
with his huge and impressive own collection. After his death, his brother Baron F. de
Lessert inherited the collection, and in 1869 his widow and daughters, the baronesses
Hottinger and Bartholdi, offered it to the city of Geneva. Lamarck's library was
scattered after his death, and his private copy of the "Histoire Naturelle des Animaux
sans Vertèbres" (1815-1822), with the marginal handwritten annotations made by
Rosalie, went to Great Britain.
Fortunately, Delessert purchased this copy afterwards, and it is actually kept in
the MHNG. The annotations made by Rosalie de Lamarck are important as they
provide information on Lamarck's original collection and type series, which had been
incorporated in the meantime with the Delessert collection, together with other
material from Hwass, Prince Masséna and Solier de la Touche. In some cases, it was
difficult to recognize the specimens of Lamarck in the Delessert collection, when the
indications were not sufficient to permit to trace them; the Delessert collection may
still contain overlooked material, especially when the number of specimens in the
actual Lamarck collection is smaller than the number indicated by Rosalie. This was
already pointed out by G. MERMOD (manuscript notes), who worked for many years
(1913-1955) on Lamarck's material in the MHNG, revising the Spondylidae and
Pectinidae in 1947, but without publishing his notes.
MATERIAL STUDIED
In the volume 6 of "Histoire Naturelle des Animaux sans Vertèbres" (1819),
Lamarck treated the family Pectinidae ("Les Pectinides"), and divided this family into
seven genera: "Houlette" [= Pedum], "Lime" [= Lima], "Plagiostome" [= Plagio-
stoma], "Peigne" [= Pecten], "Plicatule" [= Plicatula], "Spondyle" [= Spondylus], and
"Podopside" [= also Spondylus]. Actually only Pedum ("Houlette") and Pecten
("Peigne") are representative genera of the Pectinidae.
Lamarck listed and described 59 recent Pecten species, and divided these
species in two groups, mainly based on the form of the auricles, and one Pedum
species; 29 of these 59 Pecten species are new taxa, mostly figured in BRUGUIERE
(1797), DELESSERT (1841), and CHENU (1844-1845, 1862). Lamarck studied material
of his own collection ("Mon cabinet")(ca. 215 specimens) and material from the
TYPE SPECIMENS DESCRIBED BY LAMARCK 467
WW)
\ ea
je U
NR Ay REN
TAX b
Jean-Baptiste Pierre Antoine de Monet, Chevalier de Lamarck
(1744-1829)
468 HENK H. DIJKSTRA
MNHN. For the Pectinidae, he used the following previous publications as references:
BONANNI (1681), LISTER (1685-1692), RUMPHIUS (1705), SEBA (1734-1765),
GUALTIERI (1742), KNORR (1757-1772), REGENFUSS (1758), PENNANT (1777), DA
CosTA (1778), D'ARGENVILLE (1780), BORN (1780), FABRICIUS (1780), CHEMNITZ
(1784), BROWNE (1789), GMELIN (1791), PoLI (1795), BRUGUIERE (1797), and MATON
(1807). It has to be noted, that he did not refer to LINNAEUS' "Systema Naturae...",
10th ed. (1758), but only to GMELIN (13th ed., 1791).
In the volume 7 of the second edition of "Histoire Naturelle des Animaux sans
Vertèbres" (1836), DESHAYES added several notes and references to Lamarck's
species.
Lamarck clearly distinguished varieties among the species described in the text
and on his labels. These varieties, for example Pecten rastellum var. [b], indicated by
a letter in parentheses (Table IV) are here excluded from the type series, according to
art. 72b,i (ICZN, 1985 : 141). However, letters in parentheses are sometimes indi-
cated without mentioning a variation. These are included in the type series or
regarded as different taxa, for example Pecten unicolor [a], and Pecten opercularis
luteus [a]. Although Lamarck mentioned the size ("Largeur' = width) in millimetres
of only one specimen, sometimes several (additional) specimens are present. For the
designation of a lectotype, these measurements, together with Rosalie's annotations in
the margin of Lamarck's own copy of the "Hist. Nat....", are of great importance.
He described 29 new taxa (actually 4 homonyms, 1 senior synonym, 11 junior
synonyms, | possible junior synonym, 9 extant taxa, and 3 uncertain taxa), of which
three are type species of nominal genera. Two lectotypes were erroneously designated
by FLEMING (1951) and CLARKE (1965) respectively (art. 74 (a)(v), ICZN, 1985: 155).
GRAU (1959) erroneously indicated a holotype for Pecten purpuratus Lamarck.
Twenty lectotypes are herein selected (Table II), and eight holotypes mentioned. The
type specimens of Pecten pellucidus Lamarck are absent, and could not be traced with
certainty in the Delessert collection (MHNG). GRAU (1959) and CLARKE Jr. (1965)
designated two type localities.
Handwritten numbers on the outer- or inner side of the shells mostly match
the numbers indicated in LAMARCK (1819). These handwritten numbers are visible
only on Lamarck's specimens and not on Delessert's specimens, nor on the pectinids
of the MNHN. Sometimes larger numbers, of uncertain origin, are present on the
shells.
As Mermod noted in a manuscript , the letters D (Delessert collection) or L
(Lamarck collection) written on the cardboards, prepared by Lunel, are not always
correct. No original label of Delessert is present.
ABBREVIATIONS
ICZN = International Code of Zoological Nomenclature
MHNG = Muséum d'Histoire Naturelle, Genéve
MNHN = Muséum National d'Histoire Naturelle, Paris
NNM = Nationaal Natuurhistorisch Museum, Leiden
TYPE SPECIMENS DESCRIBED BY LAMARCK 469
2 + na +
Ean :i : ci 3 \
si 168 + ANIMAUX 8- 2
: Ostrea plica. Lin. Gmel. n°. 14.8 RA RA set
| Rumph. Mus. 1. 44.,4ig. O. , unters 2?
h D9Argenv. Conch. t. 24. fig. C. so
, Chemn. Conch. 5. t. 62. f. 598. a, b. : vo
È Facyclop. pl. 212.f . 5? i n
[b] War. testé purpureo-nigricante.
Habite l'Occan indien. Mus. n°. Mon cabinet. Coquille oblongue-
arrondie, à stries longitudinales bien marquées dans sa moitié
supérieure. La variété [b] est rare, fort belle, large de 47 mil-
limétres. Mus. n°. La base tronquée est étroite. .
\ oad ÿ _20. Peigne glabre. Pecten glaber.
P. 1estd subæquivalvi; radiis 10 lavibus, superne dilatato-eva-
nidis, alternis minoribus ; striis longitudinalibus ad interstitia.
Ostrea glabra ? Lin. Gmcl. n°. 5o.
Bonan. recr. 2. f. 12.
Knorr. Vergn. 2. t. 10. f. 2; et 5. t. 10. f. 5, 6.
Chemn. Conch. 7. t. 67. f. 642. 643.
Encyclop. pl. 213. f. 1.
1 [h] Har. testd fulvd, immaculatd.
t
Knorr. Vergn. 5. t. 9. f. 2.
Habite la Méditerranée. Mus. n°. Mon cabinet. Espèce commune ;
dans les collections, tés-vari¢e dans ses couleurs ct scs carac-
tères, ct 4M difficile à circonscrire dans ses limites. Beaucoup de \
À > es variétés sont fort jolies. ;
: t
lyc ulle di:. Peigne sillonné. Pecten sulcatus. - DX
4 valtsa fe P. testd subequivalvi, alba, fusco maculatd; radiis 10 æquali-. -
bus, undiquè convezis , uti interstitiis longitudinaliter sulcatis. _ È >N
An Chemn. Conch. 7. t. 67. f. 641? ose Lal
1 gabe [b] Far. testé roseo tinctd ; flammulis albis transversis. - +:
- Habite dans la Méditerrance, au golfe de Tarente. Mus. n°. Mon 4
cabinet. Espèce jolie, variée, distinete de la RE ainsi
5 que celles qui suivent. Tee =
È 1 22. Peigne vierge. Pecten virgo. STIRIA ET
7 P. testd tenui, pellucid , albd, roseo pani tinctd ; radiis 1e ,
b convexiusculis interstitiisque SIDE à <ayi f 5 \ >
- Gualt. test. t. 73. fig. HP) * 0° EL EN
u .- Habite la Méditerranée; au golfe de Tarente. Mon9 cabinet. Ses
È stries longitudinales sont peu STE Largear,a 4 millimètres.
8i4 | . i Préc x via, PRE ti
CE >
Marginal handwritten annotations of Rosalie de Lamarck
470 HENK H. DIJKSTRA
WAM = Western Australian Museum, Perth
ZMA = Zodlogisch Museum Amsterdam
ZMK = Zoologisk Museum Kgbenhavn
D = Diameter: convexity of the shell
H = Height: dorsal to ventral margin of the shell
W = Width: anterior to posterior margin of the shell
Lv. = left valve
rv. = right valve
All specimens treated are pairs of matching valves, unless otherwise mentioned (1.v.
for left valve and r.v. for right valve).
THE SPECIES
Pecten medius Lamarck, 1819 (Figs. 1-3)
References:
CHEMNITZ, 1784 : 272-273, pl. 60, figs. 586, 587, 589?; LAMARCK, 1819 : 163, n° 2 (Orig.
descr.); DEFRANCE, 1825 : 239; DESHAYES, 1832 : 715-716, n° 1; DESHAYES in Lamarck, 1836 :
130, n° 2; CHENU, 1843 : 2, pl. 4, figs. 2-6; FLEMING, 1951 : 129, pl. 17, figs. 1-4 (designation
of lectotype [= error]).
Type locality: unknown.
ype mia Keita:
Specimens cited : "Mus. n°." [MNHN]
Size given : not indicated.
H W D
1. MNHN, lectotype 93 mm 111 mm 24 mm
2. MNHN, paralectotype 1 85 mm 96.5 mm 18.5 mm
3. MNHN, paralectotype 2 75 mm 85.8 mm 24.5 mm
4. MNHN, paralectotype 3 70 mm 78.1 mm 17.5 mm
Other material:
MHNG n° 1088/8 53 mm 60 mm 13 mm
with a handwritten label of Lamarck.
(= Pecten medius Lamarck; erroneous designation of lectotype, see remarks below)
MHNG n° 1088/16 59 mm 64 mm 21 mm
(= Pecten ziczac var. [b] Lamarck)
MNHN 63 mm 68.5 mm 20 mm
(= Pecten ziczac var. [b] Lamarck)
REMARKS :
LAMARCK (1819 : 163) described this taxon as a species intermediate between
Pecten maximus (Linnaeus, 1758), the former species (n° 1), and Pecten jacobaeus
(Linnaeus, 1758), the following one (n° 3), without indicating the type locality. He
referred to CHEMNITZ9 figures 586, 587 and 589 with a questionmark. CHEMNITZ
mentioned in his description that this species is a variety of P. maximus from the Red
Sea, and collected by Forsskäl (figs. 586 and 587) (ZMK). According to CHEMNITZ,
fig. 589 is a P. jacobaeus.
TYPE SPECIMENS DESCRIBED BY LAMARCK 471
The description of Lamarck was based on material from the MNHN (type
series), indicated as "Mus. n°", without registration number. However, Rosalie de
Lamarck's marginal annotations on Lamarck's personal copy of the sixth volume of
"Histoire Naturelle des Animaux sans Vertèbres", suggest that there could be also a
"type specimen" in Lamarck's collection, as she noted one specimen. It is quite
possible that this specimen was observed after the description of P. medius, because
in the original text there is no indication of "Mon cabinet=, generally mentioned by
Lamarck, when a species is described from his own collection. Thus, it cannot be
proved whether this specimen really belongs to the type series (ICZN art. 74a.v), and
therefore it must be considered as erroneously designated as lectotype by FLEMING
(9515129);
LAMARCK (1819 : 164) described Pecten ziczac var. [b] with the locality
"Nouvelle Hollande= [= Australia]. Both specimens observed are typical for the Shark
Bay area (W. Australia) and similar to P. medius Lamarck. It is possible that Péron
and his party, on board of the French corvette "Le Naturaliste", collected these
specimens in March 1803, because the ship anchored there for several days (CORNELL,
1974). Unfortunately neither accompanying labels nor notes of Péron are traced.
DESHAYES (1836 : 131) noticed that this variety is much like Pecten medius:
"Nous avons vu cette variété dans la collection du Muséum et nous avons reconnu
qu'elle n'appartenait pas au Pecten ziczac, mais au Pecten medius n° 2...."
CHENU (1843) described this variety as a distinct species: Pecten lamarckii,
from the Atlantic Ocean (error); however this name was already preoccupied by a
fossil pectinid from Malta (DEFRANCE, 1825 : 260).
The lectotype of P. medius Lamarck, herein selected, agrees to TATE's des-
cription of Pecten meridionalis (1887 : 115) from Tasmania. The first paralectotype
is also similar to the lectotype. The second paralectotype is a typical pectinid from the
South Australian area, and also described by TATE (1887 : 114) as Pecten fumatus var.
albus (emend. for alba). It is quite possible that this material was also collected by the
French expedition to Australia (1800-1804), because the corvettes anchored also in
southwestern Australia and Tasmania (CORNELL, 1974).
NOMENCLATURAL STATUS :
Cox (1929 : 203) noted that P. medius Lamarck is a homonym of Pecten
medius Bosc, 1802, but IREDALE (1939 : 365) rejected it, because Bosc did not quote
Ostrea media Gmelin, 1791. However, Bosc did not have the intention to describe
new taxa, but he only redescribed former publications, such as of GMELIN, and they
both only referred to GUALTIERI (pl. 74 figs. N, O). According to the ICZN art. 53c,
P. medius Lamarck is a junior homonym of P. medius Bosc.
CURRENT TAXONOMIC POSITION AND DISTRIBUTION :
The valid name for this taxon is Pecten fumatus Reeve, 1852. The following
names are synonyms: Pecten medius Lamarck, 1819 (preocc. by Bosc, 1802); Pecten
fuscus [Klein] Sowerby II, 1842 (preocc. by Bosc, 1802); Pecten bifidus Menke, 1843
(preocc. by MÜNSTER in GOLDFUSS, 1836); Pecten lamarckii Chenu, 1843 (preocc. by
472 HENK H. DIJKSTRA
DEFRANCE, 1825); Pecten modestus Reeve, 1852 (preocc. by Bosc, 1802); Pecten
fumatus var. albus Tate, 1887; Pecten meridionalis Tate, 1887 (preocc. by von
EICHWALD, 1865); Notovola preissiana Iredale, 1949; Pecten jacobaeus byronensis
Fleming, 1955.
WALLER (1991 : 73) mentioned three morphological or geographic clusters
from Australia: Pecten fumatus fumatus Reeve, 1852, from eastern and southeastern
Australia , P. fumatus "meridionalis" Tate, 1887 (non P. meridionalis von Eichwald,
1865), and P. modestus Reeve, 1852, from western Australia.
Recent protein electrophoresis relevant to the genetic relationships of southern
Australasian Pecten (s.s.) by WOODBURN (1989 : 226) suggests that there is only one
extant species in Australia, although somewhat different morphological features are
observed within populations from Western Australia and Tasmania.
P. fumatus lives in shallow waters on muddy sand bottom from southern
Queensland, New South Wales, and Tasmania to southwestern Australia. No records
are known from northwestern, northern or northeastern Australia. Notovola Finlay,
1926 is a junior synonym of Pecten Miiller, 1776.
Pecten bifrons Lamarck, 1819 (Figs. 4-8)
References:
LAMARCK 1819 : 164, n° 4 (Orig. descr.); DESHAYES in Lamarck 1836 : 131, n° 4; DELESSERT
1841 : pl. 15, figs. 5a-c, text; CHENU 1843 : 3, pl. 22, figs. 3, 3a-b; CHENU 1862 : 184, figs. 925-
926.
Type locality: "Les mers australes et de la Nouvelle Hollande. Péron". [Only known
from southern Australia, det. Dijkstra]
lSvapiezmraitze nata le:
Specimens cited : "Mus. n°." [MNHN], "Mon cabinet" [MHNG]
Size given : "Largeur [= width], 105 millimètres".
H W D
1. MNHN, lectotype 101 mm 106 mm 32.5 mm
2. MNHN, paralectotype1 96.5 mm 97.5 mm 28 mm
3. MNHN, paralectotype 2 82 mm 82.5 mm 25.5 mm
4. MNHN, paralectotype3 60 mm 59 mm 2 mim
5. MHNG n° 1088/14/2, paral. 98.5 mm 100.5 mm 34.5 mm
illustr. DELESSERT (1841) : pl. 15, figs. Sa-c.
illustr. CHENU (1843): pl. 22, figs. 3, 3a-b.
illustr. CHENU (1862) : figs. 925-926.
6. MHNG n° 1088/14/1, ?paral. 106 mm 110 mm 32 mm
REMARKS :
Lamarck mentioned Péron in the original text, indicating that this material
from southern Australia was collected by the French expedition (1800-1804) to
Australia.
Although two specimens of P. bifrons are in the Lamarck collection (MHNG),
having been isolated from the Delessert collection as Lamarck's specimens, only one
was mentioned by Rosalie de Lamarck, and was figured by Delessert and Chenu.
TYPE SPECIMENS DESCRIBED BY LAMARCK 473
CURRENT TAXONOMIC POSITION AND DISTRIBUTION :
P. bifrons Lamarck is the type species of Equichlamys Iredale (1929 : 162).
This species occurs in shallow water off southern New South Wales, Tasmania and
South Australia.
Pecten purpuratus Lamarck, 1819 (Figs. 9-11)
References:
LAMARCK, 1819 : 166, n° 11 (Orig. descr.); DESHAYES in Lamarck, 1836 : 134, n° 11;
DELESSERT, 1841 : pl. 16, figs. 5a-c; CHENU, 1843 : 4, pl. 14, figs. 1-9; GRAU, 1959 : 103-105,
pl. 34.
Type locality: "Les mers orientales et australes... On le dit du Japon". Type locality
designated by GRAU (1959 : 104) : Callao, Peru.
MVP material:
Specimens cited : "Mus. n°. [MNHN], Mon cabinet" [MHNG]
Size given : "Largeur, 112 millimètres".
H W D
1. MHNG n° 1088/20/2, lectotype 109 mm 114 mm - (1.v.)
illustr. DELESSERT (1841) : pl. 16, fig. Sc.
2. MHNG n° 1088/20/1, paralect. 66.5 mm 71 mm 24 mm
illustr. DELESSERT (1841) : pl. 16, figs. 5a-b.
3. MNHN, paralectotype 1 87 mm 95 mm - (r.v.)
4. MNHN, paralectotype 2 86 mm 90 mm - (r.v.)
REMARKS :
The specimens in the Lamarck collection (1 complete specimen and 1 valve)
correspond to the annotations made by Rosalie. The single left valve is ciosest to
Lamarck's original measurement (112 mm), and is herein selected as lectotype. The
paralectotypes are semi-adult specimens, and much smaller than the width mentioned
by Lamarck.
CURRENT TAXONOMIC POSITION AND DISTRIBUTION :
The species is presently placed in Argopecten Monterosato, 1889, which is a
senior synonym of Plagioctenium Dall, 1898. The geographical range is from
Colombia to Chile, and living material is observed in shallow inshore waters on rocky
and sandy bottoms.
Pecten lineolaris Lamarck, 1819 (Figs. 12-16)
References,
LAMARCK, 1819 : 166, n° 12 (Orig. descr.); DESHAYES in Lamarck, 1836 : 134, n° 12;
DELESSERT, 1841 : pl. 15, figs. 3a-b, text; CHENU, 1843 : 4, pl. 15, figs. 1, la-b.
Type locality: unknown.
IDE material
Specimens cited : "Mon cabinet" [MHNG]
Size given: 8Largeur, 26 ou 27 millimètres=.
H W D
1. MHNG n° 1088/21, holotype 25 mm 26.5 mm 13.5 mm
illustr. DELESSERT (1841) : pl. 15, figs. 3a-b.
illustr. CHENU (1844) : pl. 15, figs. 1, 1a-b.
474 HENK H. DIJKSTRA
REMARKS:
MERMOD isolated the type, a complete specimen from the Delessert collection,
and it was also mentioned by Rosalie in the margin of Lamarck's personal copy. This
holotype is illustrated by DELESSERT (1841) and CHENU (1844).
CURRENT TAXONOMIC POSITION AND DISTRIBUTION :
This species is placed in Argopecten Monterosato, 1889. Aequipecten maya-
guezensis Dall & Simpson, 1901 is a junior synonym (ABBOTT, 1974 : 445). The
geographical distribution is from SE Florida to the Caribbean region, and living
specimens are observed from littoral depths on sandy or muddy sand bottoms.
Pecten rastellum Lamarck, 1819 (Figs. 17-26)
Reitemenges:
LINNAEUS, 1758 : 698, n° 196? (sic, = 166?); LAMARCK, 1819 : 166-167, n° 14 (Orig. descr.);
DESHAYES in LAMARCK, 1836 : 135, n° 14; DELESSERT, 1841 : pl. 16, figs. la-b, 2a-b, text;
CHENU, 1844 : 5, pl. 15, figs. 2, 2a-b, 4, 4a.
Type local"Lesi mters ydu :nor d".
Lamarck's indication of locality is very confusing, as it could be either the northern Atlantic
Ocean or the northern Pacific Ocean. This species does not exist in either region. Therefore a
new type locality is here designated: Off Punta Engano, Mactan, Cebu, Philippines (ZMA
coll.).
pe material:
Specimen cited : "Mon. cab." [MHNG]
Size given : 8Largeur, 34 millimètres=.
H W D
1. MHNG n° 1088/24, holotype 32.5 mm 34.5 mm 8.5 mm
white specimen (large).
illustr. DELESSERT (1841) : pl. 16, figs. 2a-b.
illustr. CHENU (1845) : pl. 15, figs. 2, 2a-b.
Ojchrerzmiaterval:
- Pecten rastellum var. [b] Lamarck [= Pecten mirificus Reeve, 1853][MHNG n° 1088/23,
Lamarck collection] , red specimen (W= 32 mm) (figs. 22-26).
illustr. DELESSERT (1841) : pl. 16, figs. 3a-b.
illustr. CHENU (1845) : pl. 15, figs. 3, 3a.
- Pecten rastellum Lamarck [MHNG, Delessert collection]
(W= 46 mm).
illustr. DELESSERT (1841) : pl. 16, figs. 1a-b.
illustr. CHENU (1845) : pl. 15, figs. 4, 4a.
REMARKS :
In Lamarck's collection, there was one complete specimen, matching
Lamarck's measurement cited in the text, and one complete specimen of var. [b], in
agreement with Rosalie's marginal annotations. Although MERMOD treated P. ratellum
var. "beta" as a type in his manuscript cardfile (MHNG), 1.C.Z.N. (1985) article 72
(b)(i) clearly states that varieties mentioned by a letter should be excluded from the
type series. In addition, the specimens and the original labels of Lamarck were
reversed before MERMOD's registration, so that the holotype, which is the white