TYPE LOCALITIES OF VASCULAR PLANTS FIRST DESCRIBED FROM OHIO

16 Pages·2003·9 MB·English

by  Ronald L Stuckey

OF VASCULAR PLANTS
TYPE LOCALITIES
FROM
OHIO
DESCRIBED
FIRST
James
LStuckey
Ronald S.Pringle
Herbarium Royal Botanical Gardens
OH
Columbus, 43212
Type
localiti(
name
and
subsequently were used as the basis for the scientific description of
when was new Throughout
any organism described as to science. the
first
it
numbers
history of classical and descriptive biology countless of organisms,
new
or taxa, have been named and described. Along with each taxon, usually
These
but not always, recorded original source location, or type locality
is its
type localities are deserving of permanent record, because they are the special
from
which must return he obtain living material
places to a biologist if is to
A may
descendents of the original populations. biologist need to verify the iden-
m
where population
and description of the taxon the area the original oc-
tity
and developmental and/or acquire
curred, obtain material for genetic studies,
and With expansion taxonomic
populational life-history information. the of
broader realms biosystematics and population analysis that
studies into the of
much
has come prominence withm the past half century, more impor-
into
tance now accorded the type localities than in eariier times. These classic
is
are significant biogeographical reference points at the international, na-
sites
and They should be spared from permanent destruction, such
tional, state levels.
dam
on stream
as the flooding of a river valley by construction of a a or river,
highway
the building of a housing or industrial complex, the construction of a
may
or the tilling of the land for agricultural purposes. Type localities therefore
evaluating areas or as potential nature sanctuaries, as
serve as criteria for sites
by Stuckey
discussed (1994).
from United
Information on the type localities of plants first described the
documented by Although the informa-
States inadequately as a subject itself.
is
and
recorded and scattered throughout the descriptive historical botani-
tion
is
and
has not been brought together into accessible usable data
cal literature,
it
banks. Taxonomic monographs regularly include information on the type lo-
names applied the taxa discussed in those papers, but they deal
calities for to
only with small groups of species, and the organization is taxonomic rather
A
than according the geographic origin of the specimens. project to record
to
t\ pc localities for species within a genus was presented in a catalogue of type
m
spccmicns names
lor the large genus Carex (Cyperaceae) (Shetler
et 1973).
al.
an nidcx and from which
nicludccl to the states, provinces, countries the speci-
It
mens obtamed
were Ijsts of the type specimens deposited therein have been
many
compiled
lor herbaria, but these references specimens according
list to
the repositories rather than according geographic
to their origin. "Botanical
gazetteers" have been compiled for a few states, but do not deal specifically with
type localities. At the state level, we are aware of pubhshed lists of type locali-
New
ties for only four states, viz. Illinois (Jones 1952), Iowa (Lammers 1985),
A
Mexico and West
(Standley 1910), Virginia (Core 1936). few smaller cover
lists
m
either a region or a specific county the United States. Below are listed over 60
from which
type localities for Ohio, a total of 197 vascular plants were de-
first
scribed from the state Stuckey and Wentz (1974) have previously catalogued
the type specimens from these locahties that were deposited in the herbarium
many
That new
the precise location of the original sources of of these taxa
described rom Ohio is not given is not surprising. Many of the taxa were named
I
m
when
the nineteenth century, precision as to location was of concern,
little
since no into national Code Botanical Nomenclature and em-
existed
of httle
phasis was placed on the source location of new biological entities. In those
was
considered newly
da\'s sufficient to record the location of a described
It
rom
plant such broadly defined areas as the State of Ohio, the shore of Lake
I
M
F.I le, he lami River valley, or the Ohio River The last locality, of course, means
t
rom
along the banks of the Ohio River or from vicinity. The most notable
its
I
example more
ol this situation the 112 or taxa described between 1808 and
is
1840 from Ohio by Constantine Samuel who
Rafinesque, the source
listed lo-
calities as "Ohio," "Lake Lrie," or "Ohio River." Rafmesque's 17 vascular plants
named
from
the shore of Lake Erie^ and nine noted as being from the Ohio River
arc included in the list below, although some of Rafinesque's plants may have
m
current usage and two upon which names many
currently used
are based, the
taxa descri bed b\' Rafinesque merely as from "Ohio" are not listed here, but they
can be determined Irom Index Rafinesquianum Andre
(Merrill 1949). Michaux's
the ru'cr where these plants were obtained are not known. Michaux known
to
is
have conducted some
botanical exploration near Ohio
the River in the vicinity
m
now
Kentucky, and what southern Illinois, as well as near the
of Louisville, is
Portsmouth, Ohio. Some specimens recorded by Michaux as hav-
present of
site
from
banks
ing been obtained from the "Ohio River" or its are, therefore, likely
outside present-day Ohio.
localities
In other situations only the nearest town was mentioned, with no specific
or habitats given for the plant. Some sites are sufficiently well described so
sites
Among
best-known type
that their locations can be determined today. the lo-
Pond
Ohio from which vascular plants have been described are Beaver
in
calities
Adams County, Cedar Bog (Cedar Swamp) in Champaign County, Cedar Point
in
along
and other locations near Sandusky in Erie County, the limestone chffs
Columbus and Franklin County,
the Scioto River north of other sites in
Dayton Greene County,
Hoffman's (Wright Brothers Prairie) east of in
Prairie
and
Rocky Fork Creek Highland County, Ofer Hollow in Jackson County,
in
Cranberry Island in Buckeye Lake in Licking County. Significant type locali-
development
have disappeared through agricultural or industrial in-
that
ties
Madison
Darby County,
clude the Oxford Prairie in Erie County, the Plains in
Montgomery
Van Dayton County.
and Cleve's Prairie near in
shown on
below
The general locations of the type localities listed are a
map Ohio and two additional maps show portions of the state where
of
(Fig.
1),
The
many type locaUties are concentrated in small areas (Figs. 2, 3). localities
^,
" ^
^
Mia
mostly
concentrated in the
are .
and Lake northern Ohio.
and adjacent counties in central Ohio, near Erie in
lin
These locations reflect the pioneer botanical exploration in Ohio, first in the
mid
Miami River valley in the early 1800s, in central Ohio mostly in the 1800s,
and The more unusual habi
and northern Ohio primarily about 1900 later
in
wet and limestone These habitat
types represented are fens, prairies, cliffs.
tat
Ohio where most de
are not well represented in the states to the east of of the
understandable,
botany had been written before the early 1800s.
scriptive It is
had not previously
therefore, that plants characteristic of these habitats that
been named and described were described from places in Ohio.
first
mentioned which were arranged alpha-
above,
Unlike the other state
lists
Ohio organized by beginning with the
by locality,
betically taxon, this list for is
Lake
Ohio, followed by sections of the state including the shore of
largest unit,
the Miami River valley, and the Ohio River valley, and then alphabetically
Erie,
under each
by Those taxa with more specific locality data are listed
county.
when was
county. two or more localities in Ohio were cited a taxon first de-
If
&
such
example with Valeriana Torr A.Gray, localities
ciliata all
scribed, as for
same
names based on specimens from the type locality are
are All of the
listed.
An names by
hsted under that locality, in alphabetical order. index to of taxa,
genus only, begins on page 1691.
names currently accepte as the
Double denote plar that are d
aster isks
t
names no
denote th are
the respective ta xa. Single ast risks t
m
longer
in use, or at least are not accepted the standard
references consulted
m
the present study, but which have been superseded by
currently accepted
names
based upon
directly them. Single asterisks are also used the names
for of
laxa originally described now
as species that are considered be
to interspecific
hybrids. In these cases, the present-day name generally includes
the original
meaning
component name
epithet, the adjectival of the nomenclatural
in a
combination was
that published The
type remain
later. localities the same.
When
name
the based on a specimen or specimens obtained Ohio no
in longer
is
name
in use for a taxon, the currently accepted given brackets when
in pos-
is
sible. Currently accepted names and taxonomic synonymy have been
deter-
FRANKLIN
Boundaries
Mar 1851-1990
25
Map
Fig. 2. of Franklin County, Ohio, sliowing b;
Meacham
mined from and and
primarily Kartesz Kartesz those
(1994), (1999),
volumes of the Flora North America North Mexico that had been published
of of
synonymy
some names
at the time of this writing. For old not listed even in in
these works, the modern equivalents have been determined from older floras,
such as those by Britton and Brown (1913), Fernald (1950), and Gleason (1952),
and from monographs, notably those by Boivin (1944) on Thalictrum, Davis
et
(1967-1970) on Ruhus, Fernald (1905) on Erwphorum, Heiser (1969) on
et
al. al.
Mackenzie on and Palmer on
Helianthus, (1931-1935) Carex, (1956, 1961)
When
name
Crataegus. no current status or taxonomic equivalent given for a
is
may now
published at the rank of variety, be inferred that varieties are gener-
it
^
WE
STERN LAKE
..,.
T'~"-#
--
X
.•r
"r"
^
53(r
>.
J
r^^TI
^. AWACO
?
r^~~'
-^
WOOD
CO
/ ^
\
•
1
T
I
SANDUS
? \ IP r'-y/'-r
tbea ned
thai -ibed£
C
species.
names
published the rank of form (Latin /ormci, abbreviation because
at
"f."),
names at this rank are not usually included in modern floras. In most cases, the
original epithet applied the form, transferred necessary the currently
to to
if
name
which would
accepted for the species, indicated here, be correct for
is
anyone wishing names
to use for these individual variations.
In some cases has not been possible to equate an older name with any
it
When
currently accepted taxonomic equivalent. the original description de-
is
may
ficient in detail and the original specimens no longer exist, not be cer-
it
tain which currently recognized taxon an old name, based on plants occur-
to
was
ring in Ohio, applied, although taxonomic research continues to resolve
some
of these long-persistent questions. This situation an especially frequent
is
named
problem with by The
plants Rafinesque. best single source for their in-
terpretation Merrills (1949) Index Rajinesquianum. Other pubhcations that
is
are especially significant with regard to plants originally described from Ohio
named
specimens include the author's studies of the plants and described
first
Thomas
by Daniel Drake (Stuckey 1969) and Nuttall (Stuckey 1966, and
1967),
his studies of the plants described by C.S. Kunth and E.G. von Steudel from speci-
mens Miami
obtained the River valley by Joseph Frank (Stuckey
in 1974).
1
AND
THH PLANTS TH TYPE LOCALITIES
EIR
OHIO '^Euphorbia commutata Engelm. ex A.Gra
Manual, ed. 389. 1856.
New 2., p.
**Acalypha rhomboidea 1836
Raf., 1:45.
Fl.
Glumac.
Fimbristylis frankii Steud., Syn. 2:1
Pi.
[sometimes included Acalypha virginica L,
in
auwmnalis Roem.
1855
[Fimbristylis (L)
rhomboidea
as var. (Raf.) Cooperr.].
Schult.].
Med.
Aesculus ohioensis Riddell, W.J. Phys. Sci.
*Hedysarum pauciflorum Gen. Amer.
Nutt., N.
f
8;360. 1835. Reprint 34. 1835 [Aesculus
p.
2:]09.]8]8 [Desmodium pauciflorum {HiiV
glabraWm.l
DC.].
Cakileamericana Nutl,Gen.H.Amer.P\.2:62.]8]8
?in.&Torr. g/abresceni
f.
'nr.4(20):588.1909.
var.pBoottJil.Carexp
ambigua
h
Scutellaria
subvar.grao7//moBoott,III.Carex p.38,pl.'
1858
[Carex gracilescens Steud.].
fig. 2.
*Carexmon/7eTucl<.,Enum.Metln.Caric.p.20.18
Dewey
"Carextenera var.sufaerecfa Olney,Caric
Bor.-Amer. 3:1871 [Carex suberecta (Oint
dArbor.e
Decouv.
Precis
Raf.,
;
'*Eleocharis
Glumac.2:
ca/vo Torr
LAKE ERIE,SHOREOF (Probably Sandusky,
at
County)
Erie
Amer
Aesculus muricata Raf, Alsogr 68. 838
p.
1
(Engelm
n
Glumac.
1
854
[fro,
1
OHIO
RIVER
Asplenium angustifollum Mic
2:265. 1803 [Diplaziu
(Spreng.) M.Broun].
f/iyrus;ncL/;w;sRaf, Atlantic 151 1832, not
J
1
DIscovium
gracileRaf. ,i.Phys.C
Roth 1787 nor Willd 1802 nor Rchb 1832
beliafalcataRaf,Newf\2]8
1836
[1837] [Lo-
swa't^son"]"'
nceraenem/iRaf.NewFI
18 19 1836[1838]
3
f^^^,^^
f,
ULonicerahirsuta
Ea\ion]
notWall 183
Bot 134 1840[7?;//;umf/
p
s
Ann. 1820
Raf., Nat. 1:13,
1
hwein
although prob
o, .... s. (1822)
pianTa^mTJ mZ^Trl^^^^^^^
,
t,.o„„niL
equivalent]
Illy
l\
"gme/Zni"
[P/c
VALLEY; MIAMI COUNTRY
Podalyria coeru
II
^'^'^^"
Convolvulus miaanthus R\6de\\,\N Med Phys
i ^
Roem
Sci 8 502 835 Reprint p 70 835 not
1 1
&Schult 1819norWilldexSpreng 824
[/po
1
**Potentillaparadoxa
Nutt. in Torr.& A.Gray, FI.N.
Carex flacadula Steud Syn Gluriiac 2 199
PI
,
^^^^^
^^^^- '^^^- ^'^°' "°^ ^^^^^ ^7^-
"^^
855 [Carex rosea Schkuhr ex ^
Willd.]. ^ ^
1
Scure//ara rad/cofa Raf, Atlantic 1:16.1832.
J.
**Carexgraa7escens Steud.,Syn.PI.GIumac.2:226,
**Spermococeg/abra
Michx., FI.Bor.-Amer.
1:82.
1855 1849
^03, not Roxb. 820 nor Sesse & Mog. 888,
Carex steudelii Kunth, Enum. Pl.2:480. 837 [Carex ^ 1 1
1
^mer.
R^'- ^'^og^- 838
^'''^ ^"'^'^ 45. [Tilia
jamesii Schwein.]. 1
^'^^''"^"^"'^
L.].
Eragrostis cognata Steud., Syn. Glumac. :273.
PI.
1
.
,
Moldenke, Castanea 1944 [Polygonum
9:41.
bRd V
'
fS
th 2 th
t 1
sag turn
itta L.].
of Ohio Route 348:**Silphium terebinthina-
Trifoliumpratense Lf.lilacinum Moldenke,Boissiera
7:4-5. 1943; see also Castanea 9:49. 1944.
Rhodora 53:133-135.1951.
COUNTY
BUTLER
Boissiera 7:5. 1943; see also Castanea 9:64^
03. Old near Hamilton: Amaranthus
prairie
944
[l/ernon/ag;gonfeo (Walter) Trel.subsp.
1
Med.
altissimus Riddell, W.J. Phys. Sci. 8:367.
gigantea].
1835. Reprint 41. 1835 [Amaranthus
p. 08. Roscoe:Stac/iys glabra Riddell, W.J. Med. Phys.
836
9:580. 836. Reprint [Stachys
Sci. 6.
p.
1 1 1
04. Small prairie half a south of Middletown:
rT^ile tenuifoliaWMd.].
Euphorbia Med.
herronii Riddell, W.J. Phys.
Sci.
CUYAHOGA COUNTY
8:58. 1835. Reprint 32. 1835 [Euphorbia
p.
SocBot, France
65:49. 191
Bull. 8.
CHAMPAIGN COUNTY
Hieraaum
10. Parma: ohioense Gand., Soc.
Bull.
Swamp
05. Cedar [now Cedar Bog], Urbana:
Amer
&
Valeriana citiata Torr. A.Gray, Fl. N. Rocky deep gorge about one-fourth
River,
1 1
.
&
2:49. 1841 [Valeriana edulis Nutt. ex Torr.
mile from Lake Efle:Cerastiumarvense Lvar.
&
A.Gray varciliata A.Gray) Cronquist],
(Torr. webb/7 Jennings,Ohio Naturalist 9:441 909;
1
.
[Cerastium arvense subsp. velutinum (Raf)
06. Near Springfield: "Valeriana ciliata Torr. & L.
Ugbor.var.i/e/uf/num.]
Nutt. ex Torr. & A.Gray var. o7/afo Cronquist]. DELAWARE COUNTY
COSHOCTON COUNTY 2. Delaware: Crataegus ellipticifolia Sarg., Ar-
1 J.
compocfo
nold Arbor. 194. 1922 [Crataegus
3:
07. North Appalachian Experimental Watershed,
Sarg.].
Aralia spinosa L f subinermis Moldenke, Casta- COUNTY
ERIE
Daucus Moldenke, Castanea
carota fischen
L. f. quarry by glacial grooves, N portion of
9:55.1944,
Kelleys Island, Lake Erie: **Juncus xstuckeyi
Daucus carota goodmanii Moldenke, Casta-
L. f M.Reinking, Brittonia 33:1 75. 976 [= Juncus
1
nea "goodmon/."
9:55. 944,
1
Moldenke, Castanea
Lobelia inflata L. f albiflora Ceda° *Helianthus luxunans E.Watson,
4. Point:
1
9:65. 944. Pap. Michigan Acad. 9:464. 1929
1 Sci.
Lycopodium
flabelliforme (Fernald) Blanch.f c/ufe; [Helianthus xluxunans E.Watson; pro sp.;= H.
Moldenke, Castanea 1944.
9:32.
gigonteus x H.grosseserratus M.Martens].
L.
[Diphasiastrumdigitatum (A.Braun) Holub],
Huron mouth: Phalangium
5. River at esculen-
1
Morus alba nigrobacca Moldenke, Castanea
f tum Gen. Amer. 1818
L. Nutt., N. 1:219.
PI.
9:51.1944.
[Cflmoss/asc/7/o/de5 Cory].
(Raf.)
iracunda Beadle]. *Fedia umbilicata SulL, Amer. Sci. 42:50. 1842
J.
Oxford Solidago moseleyi Fernald, [Valerianelb umbilicata Alph.Wood],
Prairie:
(Sull.)
.
Rhodora 1908 [Euthamia gymno- Man.
10:93. */-/e//onr/ius/(e//ermar)/7 Britton, FI.N. States
spermoldes Greeneioflen included mSollda- 994. ]90], "kellermani"; see also W.A.
p.
go,as S.gymnospermo/des (Greene) Fernald], Kellerman, Ohio Naturalist 2:179-181. 1902
=
'Greene, Amer. Midi. Natural- [Hellanthus xkellermanii Britton, pro
sp.;
294, ]9]2[ThalictrumdasycarpumF\s~ Hellanthus sallcifollus A.Dietr. x H.
marada
8. Sandusky Crataegus AsheJ.EIisha *Rudbeckla sullivantii C.L.Boynton & Beadle,
Mitchell Sci Soc 20 53 1904 [Crataegus Biltmore Bot. Stud. 1:15. 1901 [Rudbeckia
&
iracunda Beadle] fulglda Alton var, sullivantii (C.L.Boynton
AsheJ
^rataegw, fenera Elisha Mitchell Soc. Beadle) Cronquist],
Sci.
20 52-53 1904 [Crataegus macrosperma 24. Columbus,cat-tailswamp,three miles west:
'Coreopsis cfcco/deo Torr, & A.Gray, N. Amer.
'\,h.'l Fl.
2^339.J842[e/densd,5co/dea(To;r.&A.Gray)
^^''''l?;'ls''^'t?'™^^^^^
25. Columbus, rocky limestone banks of the
COUNTY
FAIRFIELD
20. Sugar Grove: Crataegus polybracteata Ashe,
J.EIisha Mitchell Sci.Soc.l 6:79. 1899;see also
1861.
Palmer, Arnold Arbor. 6:57. 1925
E.J.
J.
*Lonlcera glaucescens Rydb. dasygyno Rehder,
var.
[Crataegus
Intricata iar^ge].
21. Clearport;T12N, 9W,sec. miles east
R.I 10, 1.5
[Lonlcera dioica dasygyna (Rehder)
var.
L.
Highway
69, east of the covered bridge over
ton/cera flora Simsvar.|3Torr.&A,Gray,FI,N.Amer.
2:6. 1841 [Lonlcera reticulata Raf.].
48-224^983
Mortlii Casnanea |=Ty" fop
fer/s
Lonicera parviflora Lam. p? sullivantii
C [var.] [sic]
ja,n.s's.vn.5 Shaver x tenu,s (Michx.)
Alph.Wood, Class-book
Bot., ed. 298.
2, p.
component
1847. Based on the Ohio of
Lonicera parviflora var, p Hook, sensu Torr. &
22. Bexley: Triphora trlarUhophora (Sw.) Rydb. var.
Camp, Rhodora
schaflneri 42:55. 940.
type determined
1 claturally distinct; locality
Columbus:
23. *Msc/ep/as5u//;ranf/7Engelm. ex
from Torrey and' Gray [Lonicera reticulata
A.Gray, Manual, ed. 1,366.1848.
Raf.].
7^
Amer. 84
A.Gray, N. 2:1 38. [Aster borealls sl
Fl. 1 1 ohio^emis^R\dde\\. W.J.^Med^Phys. 8:49
'
(Torr.& A.Gray) Prov.;sometime5 segregated
835. Reprint 57. 835 [sometimes segre-
p.
1 1
from Symphyotrlchum
Aster as boreale Torr. gated from Solidago Oligoneuron
as
& A.Gray) A.Love &D.Ldve].
o/i;oeme(Riddell)G.N.Jone5],
*Carex conjuncta Boott, Carex 3:122, 392,
III. pi. 27. Dublin, calcareous ravine and steep decliv-
1862.
on the Scioto Prenanthes proteo-
ity River:
Carex Boott A.Gray, Amer.
sullivantii in Sci.
J. Med.
p/iy//aRiddell,W.J. Phys,Sci,8:490. 1835.
42:29. 842 [Carex xsullivantii Boott, pro
sp.;
1 Reprint 50. 835 [Prenanthes alba
p. L.].
1
C
= Carex hirtifolla Mack, x gracillima
**M//umn/va/eRiddell,W.J.Med,Phys.Sci. 8:525
Schwein.].
1835. Reprint 93, 1835.
p.
3foegusofi;oen5/sSarg„ J, Arnold Arbor. 3:1 83. 28. Between and Glenmary: Crataegus
Flint
1922 [Crataegus arborea Beadle],
fran/<//nens/sSarg.,J,Arnold Arbor,4:100,192^