Two New Sympatric Species of the Genus Kusumia (Coleoptera, Trechinae) from the Southeastern Part of the Kii Peninsula, Central Japan

2005·7.4 MB·

by  Ueno, S-I

Elyt'a,Tokyo,33(1):341-352,May30,2005
TWoNewSympatricSpeciesoftheGenusKusumja(Coleoptera,
Trechinae) fromtheSoutheasternPartofthe
KiiPeninsula,CentralJapan
Shun-lobi UENo
DepartmentofZoology,NationalScienceMuseum(Nal.Hist),
3-23-1Hyakunin-cho,Shinjuku,Tokyo,169-0073Japan
and
TakaoNAITo
3-4-13Ikeda,Neyagawa-shi,Osaka,572-0039Japan
Abstract TwonewupperhypogeanspeciesofthetrechinegenusKusumlaarede-
scribedfromthesoutheasternpartoftheKiiPeninsula,CentralJapan. 0neofthembe-
longstothee1ongatagroupandisnamedK.cl,ocodilusinviewofthepeculiarlyformed
aedeaga1apica11obe1ookingliketheupperjawofacrocodile.Theotherspeciesisamem-
beroftheyos/likawatgroupandisnamedK.inspe・tata.Thetwospeciesaresympatricnear
thesouthernendoftheOhmineMountains,thoughtheydonotseemtocoexistintheex-
actlysamehabitat.
Twoyearsago,thesecondauthorofthepresentpaperunexpectedlycameacrossa
newStyg1otrechusat thesouthernmostof theOhmineMountainRangeinthesouth-
easternpartoftheKiiPeninsula,CentralJapan(UENo&NAIT0,2003a).Beforethen,
norepresentativesofthegenuswereknownintheareaseastofthedeepwidevalleyof
theTotsu-kawaRiver,whichdividesthesouthernpart of thepeninsulaintotwore-
gions. At thesame time, he foundout two corporaof anundescribed species of
Kusumlaonthesamemountainabout1,000msouthbywestofthegully inwhichwas
discoveredtheStyg1otrechus(cf.UENo&NAIT0,2003a,p 235).
sincethen,werepeatedlymadesearchesforlivingspecimensoftheKusumiaatthe
easternsjdeofChausu-yama,butoureffortswerenotrepaidsoeasilybecauseofthe
greatrarityofthespeciesconcerned. 0ntheotherhan a tenera1 female and a dead
bodyofasecondspeciesofthesamegenusweremetwithbythesecondauthoratthe
collectingsiteofslygiot,-echus,andweredeterminedasanundescribedmemberofan-
otherspecjes_group.Thus, itbecameapparentthattheeasternslopeofChauSu-yama
wasinhabitedbytwoundescribedKusumlaspeciesbelongingtotwodifferentlinea9eS,a
sjtuatjonseldommetwithinJapanesetrechinesotherthanthoseofthegenusTreChiama.
342 Shun-IchiUENoandTakaoNAIT0
Andtherefore,renewedeffortswereexerted,mostlybythesecondauthO「,toob-
tajnperfectspecimensofthetwospeciesofKusumla,andbrought forthth「ee9oOd
specjmens(1 2 )andseveraldeadbodiesofthefirstspecies,butonlyonemature
,
femaleofthesecondone.Unfortunately,theonlycolluviumknowntoharbOu「theSly-
g1otrechusandthesecondspeciesofKusumlawascompletelywashedawaybyaflood
afteroneofthetyphoonsthatattackedtheKiiPeninsulaintheautumnof2003,Sothat
wehad toabandonhope toobtainadditional specimensof thesecondspeciesof
Kusumlainthenear future,atleastatthatparticularspot.Neartheendof2004,how-
ever,amaturemaleofanupperhypogeanKusumiamostprobablyreferabletothesec-
ondspeciesinquestionwasunexpectedlytakeninagullyattheothersideofChausu-
yama,6.2kmsouthwestofthelostcolluviuminabeeline.Havingthisinvaluablespec-
imenathands,wehavedecidedtointroducethetwonewspeciesintoscience forthe
reasonofbothtaxonomicalandzoogeographical interest.Thefirstspeciesbelonging
tothee1ongatagroupwillbenamedK.crocodilusinviewoftheuniqueconformation
oftheaedeaga1apical lobe,andthesecondonebelongingtotheyoshika、,laigroupwill
benamedK.inspe1・atabecauseofitsunexpectedoccurrenceinsympatrywiththefirst.
Theabbreviationsusedhereinarethesameasthoseexplainedinpreviouspapersof
UEN0S'.
Beforegoingintofurtherdetails,wewishtoexpressourheartfeltappreciationto
Dr.YoshiakiNlsHIKAwAforhiskindhelpinfieldworks.
KusumiacrocodihlsS.UENoetNAIT0,sp nov.
(Figs.1-3)
Length:4.95-5.35mm(fromapicalmarginofclypeustoapicesofelytra).
Belongingtothee1ongatagroupandrelated toK dentataS.UENo(1999,pp.
289,295,figs6-8)fromamineaditatKaminagaiinKumanogawa-cho,butsomewhat
smalleronanaverage,withnarrowerconfigurationofprothoraxandelytra,theformer
ofwhichislesscontractedposteriadandhaslessclearlydefinedbasalpart,andthelat_
terobviouslylessampleinapicaltwo-thirds.Strikinglydifferent fromK dentataand
alltheothercongenersinthepeculiarlyformedapical lobeofaedeagusaswj1l bedo_
scribedlater.
Bodynarrow,withsmallheadandrelativelyshorthindbody;surfacewhollypu_
beSCentWiththeexceptionof fronsandsupraorbitalareas;microsculptureasjnthe
othe「Speciesofthee1ongatagroup.ColourdarkerthaninKdentata,darkbrownpar_
flatly「eddiSh,dullshinyandhardlyiridescent;palpipale;apjca1halvesofantennae,
legsandventerofhindbodymoreorlesslighterthandorsum
HeadSmall,Subquadrate,aboutaswideasoralittlewiderthanlong,wjdestata
levelbetweenSuP「aOrbitalpores,andmoregraduallynarrowedanteriadthanposterjad;
do「SumdeP「esse(i,withentirefrontalfurrowsgentlyarcuateinfrontandwidelydjver_
9entbehind;fronsandsupraorbitalareasgentlyconvex;genaegentlyconvexatthepos_
te「iO「Pa「tS,「atherdenselycoveredwith fairlylonghairs;neckwjde,wjththeanterjor
TWoNewSymPat「CISpeciesoftheGenusK1ls1ml a
343
Fi9 1 KtlSM11aC'coe '/i'SSUEN0etNAIT0,SP
nov., ,fromshinog6jnK1tayama_mura
344 Shun-lobi UENoandTakaoNAIT0
constrictionnotdeep thoughdistinct;apicalmarginoflabrumstraightat themedian
part;mandjblesratherstout;mental toothsimple;palpislender;antennae fairlylong,
reachingapicalthirdofelytrain ,apical three-seventhsofelytrain ,segment2the
shortestandsegment3thelongestasusual,segments5-7eachnearly fourtimesas
longaswide,terminalsegmentaboutaslongassegment7,obviouslylongerthanscape.
pronotumsubcordate,alittlenarroweronanaveragethaninK dentata,widestat
alevelbetweentwo-thirdsandfive-sevenths frombase,andobviouslywideratbase
than inthelatterspecies;PW/HW143-1.53(M I48),PW/PL0.91-0.98(M0.94)
[pL/pw 1.02-1.10 (M 1.07)], PW/PA t48-1.60(M I54),PW/PB145-1.51 (M
1.49);sidesnarrowlyborderedthroughout,widelyandlessstronglyarcuatein front
thaninK dentata,almoststraightatmiddle,shallowlysinuateataboutbasalfifth,and
thenslightlydivergenttowardssharphindangles,whichareproducedpostero-latera11y,
ante-basal sinuationnotsosharplymarkedas inK dentata;marginalsetaeas inK
dentata; apex usually somewhat narrower than base, PB/PA 0.99-1.07 (M 1.03)
[PA/PB0.94-1.01(M0.97)],withfrontanglesnarrowlyroundedandhardlyproduced;
basewidelyemarginatebetweenhindangles; dorsumgentlyconvex, ratherdensely
coveredwith fairly longhairs,sculpturesasinK dentata.Lateralexpansionofpro-
pleuraslightlyvisiblefromabove.
Elytraelongatedsuboval,widest at about orslightlybehindmiddle,e'vidently
widerthanandtwoandahalftimesaslongasprothorax,andalmostequallynarrowed
in frontandbehind;EW/PW157-1.67(M I 62),EL/PL2.38-2.62(M2.48),EL/EW
1.58-1.68(M I64);shouldersdistinctandobtuselytuberculate,withprehumera1bor-
derscomplete,nearlystraight,oblique,andfringedwitharowofhairs;sidesmoder-
atelyreflexedthroughout,veryslightlyemarginatebehindshoulders,then feeblyand
widelyarcuate, andconjointly roundedat apices, eachwith aslight preapica1emar-
gination;dorsumevidentlylessconvexthaninK dentata,thoughsteeplydeclivousat
thesides,apical declivitygentle; adistincttransverse foveapresent inbasalareas,de-
limitedoneachsidebyaveryobtusebasalcarinaformedbythebasalportionofinter-
val 5; striae entire, clearly impressed throughoutandambiguously crenulate, inner
striaedeepenedinbasal fovea,stria8irregularlydeepenedinapicalportion;scutellar
strioleeitherveryshortorvestigial;apicalstrictodeepbutmoreorlessirregular,usu-
allymergingintostria5attheanteriorend;intervalsmostly at,eachbearingarowof
fairlylongsuberecthairs,whichextendstoapex;apicalcarinaveryobtuse;stria3de-
voidofsetiferousdorsalpore;preapica1poreclosetoapex, thoughstillmoredistant
fromapexthan fromsuture;stria5alwayswithtwosetiferousdorsalporesaf t/7-1/5
and1/2-3/5frombase,respectively;marginalumbilicateseriesordinary foramember
of Kusumia.
Ventralsurfacepubescentexcept for lateralparts;analsetaeordinary. Legsrela-
tivelyshortandstout;protibiaegraduallydilatedtowardsapices;tarsi relativelyshort,
tarsomere1shorterthantarsomeres2-4combinedinbothmeso-andmetatarsi; in
,
twoproximalprotarsomereswidelydilated,markedlyproducedinwardsatapices,and
furnishedbeneathwithsexualadhesiveappendages.
TwoNewSympatricSpeciesoftheGenusK1ls1l,nla 345
3
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/ /
/、一一ー/
0.5mm
Figs. 2-3. MalegenitaliaofKusumlac,-ocodilusS.UENoetNAIT0,spnov.,fromShinogoinKitayama-
mura;leftlateralview(2),andapicalpartofaedeagus,dorso-apicaIview(3).
Malegenitalorgan verysmall thoughmoderatelysclerotized.Aedeagus three-
tenthsaslongaselytra,fairlyslender,hardlyarcuateatmiddlethoughventrallybentor
curved inbothbasalandapicalparts, thelatter of which is very long; basal part
stronglybentventra fairlyelongate,withsmallbasalorificewhosesidesaredeeply
emarginate;sagittalaileronverylarge,protrudingventro-proxima11y;apical lobevery
longandnarrow,nearlyparallel-sidedandnarrowlyroundedat theextremityindorsal
aspect;viewedlaterally, apical lobegently cuvedventradat thebase,straightlyand
gradually acuminatetotheextremity,whichisslightlyre?exedandprovidedwitha
minutetoothattheventralside,venter furnishedwithtworowsofrecurvedteeth,three
to fiveon thele量sideandoneor twoon therightside, the teethbeingmuchmore
spacedthaninK dentataevenintheleftrow;ventralmarginnearlystraightatmiddle
inprofile.Innersacpartiallyscalythoughdevoidofsclerotizedteeth-patches;copula-
torypiecemuchsmallerandsimpler thaninK dentata,nearlytwo-ninthsaslongas
aedeagus,withelongateapicalpartbluntat theapex,twistedat themiddle,andnar-
rowlyextendingproximadtoanaciculateproximalend.Stylesbroadinbasalpartsbut
shortandnarrowinapicalparts,leftstylelongerthantheright,eachbearingonlythree
setaeofdifferentlengthattheapex.
Typeseries. Holotype: ?, (300malt)5-VIII-2003,T. NAIT01eg. Allotype: ?,
(300m alt) 2-XI-2003, T. NAIT01eg. Paratypes: 19, (400m alt) 16-II-2004, T.
NAIT01eg;1e, (300malt)(founddead;appendagesexcept femoramostlymissing),
29-IV-2003, T. NAIT0 leg; 2 , (300m alt) (found dead; fair condition),
25-VII-2004, T. NAIT0 leg; Ie, (300m alt) (found dead; fair condition),
346 Shun-IchiUENoandTakaoNAITo
9-VIII-2004,T. NAIToleg.AlldepositedinthecollectionoftheDepartmentofZool-
ogy,NationalScienceMuseum(Nat.Hist),Tokyo.
Type locality. Shinogo, 300 and 400m in altitude, in Kitayama-mura of
WakayamaPrefecture,CentralJapan.
Notes. ThoughdoubtlessbelongingtothesamelineageasK e1ongataS.UENo
(1999,pp 289,290,figs 2-5),therelationshipbetweenthisnewspeciesandK den-
fatamaynotbesocloseasthatbetweenthelatterspeciesandK e1ongata. This canbe
inferred fromthe fact that thepresentspeciescanberecognizedonitsnarrowbody
formalone. 0n thecontrary, K dentataispractically indistinguishablefromK e1on-
gatainexternalmorphologyandalsoingeneralconfigurationofthemalegenitalia,
withtheexceptionofincrediblydifferentapical lobeoftheaedeagus. Themalegeni-
taliaofK.crocodilusarenotonlydifferent fromthoseofK dentataintheuniqueapi-
cal ichebutalsoinotherpartsoftheaedeagus.
Kusumia c1-ocodilus has so far been found from twoscrees in thesteepgully
calledtheNuka-dani,whichrunsdownnortheastwardsinadeciduousbroadleaved for-
estbetweenChausu-yama(1,181minheight)andAmedani-yama(840m inheight),
bothverysteeprockypeaksonasoutheastwardbranchridgeat thesouthernmostof
theOhmineMountainRange.ThisgullyisasmallbranchattherightsideoftheShi-
ne-goStream,and24.5kmdistanttothenorth-northeastinabeelinefromShimono-
sako-ko Adit in Kumanogawa-cho, thetype locality of K dentata. The Nuka-dani
Gully isforkedatanaltitudeof300m,andalargescreeisdepositedatthefork inthe
right-handbranch.Sixofthesevenspecimensofthetypeseriesandfragmentsofdead
bodiesweredugout fromthisscree,usuallybelow50cmindepth.Theremainingone
was found fromcolluviadepositednear thesourceof theleft-handbranch,equally
fromthedepthofabout50cm.Theywereveryagilewhenexposedtolight.
Itisworthnotingthatallthedeadbodiesofthisspeciesareheavilyinfectedwith
fungiandthatoldonesarereadily fractured. Threeofthe fourparatypesfounddead
must havedied very recently,sincetheyare ina fair condition to the tips ofap-
pendages and even their genitalia can be examined rather satisfactorily. Unusual
rarenessof fullymatureadultsandrelativeabundanceofdeadbodiesmaybedueto
serious contagion of agerm-produced disease caused by the fungi, although the
Laboulbenialesareusuallyconsideredratherharmlesstohostbeetles.
Aswasmentionedintheintroduction,thespecificnameofthisnewspeciesisthe
Latinnouncrocodilusinapposition,denotingtheuniqueconfigurationoftheaedeaga1
apicallobethatresemblestheupperjawofacrocodile.
KusumlamsperataS.UEN0etNAIT0,sp nov.
(Figs4-6)
Length5.15-5.75mm(fromapicalmarginofclypeustoapicesofelytra).
Belongingtotheyoshika、?,algroupandsimilarinmanyrespectstoKyoshikawai
1atipennisS. UENo (1999, pp 290, 313) from the vicinities ofOba-ga-mineat the
TwoNewSympatricSpeciesoftheGenusKusunlia 347
Fig. 4. Kusumla insperataS.UENoetNAIT0,sp nov., from the Kuma-daniat Shimokuzukawaof
,
Totsukawa-mura.
348 Shun-IchiUENoandTakaoNAIT0
northernpartoftheOhmineMountains,buttheprothoraxandelytraarealittlemore
elongate,thepronota11atera1sidesaremoregentlyarcuateinfrontandmoreshallowly
sinuatebehin andtheexternaldorsalseriesofelytrausuallyconsistsoftwosetifer-
ouspores.
ColourasinK y. latlpennls.HeadsimilartothatofK y. lalipennls,withanten-
naereachingthemiddleofelytraevenin Pronotumrelativelyelongate,widestat
.
about five-seventhstothree-fourths frombase;PW/HW130-1.38(M I34),PW/PL
0.83-0.85(M0.84) [PL/PW1.17-1.20(M1.18)],PW/PA t40-1.46(MI43),PW/PB
1.36-1.40(M I39);sidesnarrowlyborderedthroughout,gentlyandwidelyarcuatein
front,shallowlysinuateataboutbasalsixthwithoutdistinctlymarkedante-basalsinua-
tion, andthensubpara11e1towardshindangles;apexaboutaswideasbase,PB/PA
1.01-1.08 (M1.03) [PA/PB0.93-0.99(M0.97)]; front anglessmall thoughslightly
producedforwards;basestraightatthemedianpart,withsharphindanglesproduced
postero-latera11yatthetips;otherpronota1featuresasinK yoshikawa1.
Elytraovate,relativelynarrow,widestataboutfive-ninthsfrombases,andalmost
equallynarrowedtowardsbasesandapices;EW/PW197inallthethreespecimensof
thetypeseries,EL/PL2.46-2.59(M2.52),EL/EW147-1.55(MI52);shouldersdis-
tinct,obtuselytuberculate,withprehumera1bordersoblique,nearlystraight,andbear-
ingonly oneanteriormosthair;sidesmoderately reflexedthroughoutexcept forpre-
humeralparts,slightlyemarginatebehindhumeraltubercles,thenmoderatelyarcuate,
andconjointlyroundedatapices,eachdevoidofappreciablepreapica1emargination;
dorsummoderatelyconvex,steeplydeclivousatthesidesandinapicalarea,withadis-
tinct transversefoveainbasalareas;basalcarinamat-defined;striaeclearly impressed
onthediscbutobliteratedattheside,striae7and8nearlyobsolete,thoughthelatteris
impressednearthemiddleandapicalsetsofmarginalumbilicatepores;scutellarstri-
olevestigial;apicalstrictodistinct thoughshallowandoftenmat-definedalongapical
margin, anteriorlyjoiningoralmostjoiningstria5; intervalsslightly convexonthe
disc,completelyflatattheside,eachbearinganirregularrowofsuberecthairs,which
doesnotextendposteriorlybeyondtheleve1ofpreapica1pore;stria3devoidofdorsal
pores,preapica1porelocatedjustbeforetheleve1oftheterminusofapicalstriole,and
muchmoredistant fromapexthanfromsuture;stria5usuallywithonlytwosetiferous
dorsalporesatabout1/6and2/5-3/7frombase,respectively;intheparatype,athird
dorsalporepresentonstria5oftherightelytronat4/7frombase.
VentralsufaceasinKy. latipennis.Legsalittleslendererthaninthelatter;tarsi
not long,tarsomere1aboutas longastarsomeres2-4combinedinbothmeso- and
metatarsi.
Malegenitalorgansmallandlightlysclerotize differingfromthoseoftheprevi_
ously knownmembers of theyoshikawai group in themuch thicker aedeagus not
stronglybentataboutmiddle.Aedeagusaboutone-thirdaslongaselytra,bold,alittle
broaderthanhigh,andventrallycurvedatboththebasalandapicalparts,withthedor_
satmarginsemicircularly roundedinprofile; basal part fairlyelongate, moderately
curved ventra withsmall basal orificewhosesidesare feeblyemarginate;sagittal
TwoNewSympatricSpeciesoftheGenusKtlstl'川a 349
6
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、、_ _ -
l 1
0.5mm
Figs 5-6. Malegenitaliaof Kusumla inspe,-ataS.UENoetNAIT0,sp nov., fromthe Kuma_dani at
ShimokuzukawaofTotsukawa-mura;leftlateralview(5),andapicalpartofaedeagus,dorso_apjca1
view(6).
aileronvestigial;vieweddorsally,apicallobeshortandverybroad,widelyroundedat
theapex;viewedlaterally,apicalpartabruptlytaperedintoshortapicallobe,whjchjs
Straight,producedventro-apica11y,andminutelytuberculateat theextremity; ventral
marginalmoststraightbehindmiddleinprofile.Innersacwhollycoveredwithminute
scales,whicharepoorlysclerotizedforthemostpartbutformacompactpatchoffjne
sclerotizedspinulesjustinsideapicalorifice;nodifferentiatedcopulatorypiece.Styles
fairlylarge,leftstyleevidentlylongerandbroaderthantheright;intheholotype,rjght
stylebearsfourapicalsetae,whereastheleftbearsonlythreesetaeattheapex.
Typeseries. Holotype: (5、,Kuma-dani,23-XII-2004,T.NAIToleg.Allotype:
,
Shinogo,5-VIII-2003,T.NAIToleg.Paratype:19(tenera1),Shinogo,23-VI_2003,T.
NAIToleg.AlldepositedinthecollectionoftheDepartmentofZoology,NationalSci_
onceMuseum(Nat.Hist),Tokyo.
Typelocality. Kuma-dani,330minaltitude,atShimokuzukawaofTotsukawa_
murainNaraPrefecture,CentralJapan.
Otherlocality. Shinogo,190minaltitude,inKitayama-muraofWakayamaPre_
fecture,CentralJapan.
Furtherspecimense;x;amined 1 Takigawa,460malt.,Totsukawa-mura,Nara
,
Pref.,CentralJapan,7-XII-2004,T.NAIT01eg;1 Hashi-tani,500malt.,Asahi,To_
,
tsukawa-mura,NaraPref.,2-X-2004,T.NAIT01eg.Bothdepositedincoll.NSMT.
Notes. It isdifficult todeterminethetrueaffinityofKusumlainsperata,even
350 Shun-IchiUENoandTakaoNAIT0
thoughthenewspeciescloselyresemblesK yoshlka、?,ai latipennts. Itislocalizedin
thesouthernpartoftheOhmineMountains,whilethelatteroccursinthenorthernpart
ofthesamemountainrange,andtheinterveningcentralpartisoccupiedbyK amice-
tumS. UENo (1999, pp 290, 315, figs 22-24),at least from Misen toNehan-dake.
Judging from the instability of elytra1chaetotaxy and other minor evidences, K
yoshikawaiyoshikawaiS.UENo(1960,pp.124,130, fig 4;1963,p it2,fig.1;1999,
pp 289, 312) endemic to Mt. 0hdaigahara-zan may be themost primitive of the
knownmembersofthespecies-groupandmay havegivenrisetootherspeciesand
subspecies. Ifthisconjectureberight,K y. latipennis,K amicorumandK. lnsper,ata
mayhavebecomedifferentiatedinthecourseofsouthwarddispersaloftheancestral
trechinealongtheOhmineMountainRange,andtheancestorofK.1nspelatamayhave
becomeisolatedbythespeciationofmoredominantspecies,K amlcorum.
Aswasnotedintheintroduction, thisspecieswasdiscoverednear theoutletof
thegully called Kurozo-dani, which empties into the Shi-ne-go Stream at about
1,000mnorthbyeastofthetypelocalityofK.cl,ocodilus. 0nlytwospecimensalive
andafragmenteddeadbodyofK.1nspelataweretakenatthisplace,all foundbydig-
gingadepositofgravellysoilatadepthof30-50cm.Regrettably thishabitat was
completelyscrapedawaybyaflood,andneitherKus1,imianorStyg1otrechushasbeen
metwithsincethen.Atthebeginningofthewinter2004,however,amalespecimenof
aKusumiamostprobablybelongingtothesamespecieswasunexpectedly dugout
fromaloosedepositofgravellysoil intheKuma-daniValleynearthesouthwestern
footofChausu-yama(orrather thesouthwestern foot ofDague-yama), inanycase
6.2kmsouthwestoftheShinogo1ocality.Thecolluviumat thecollectingsitewasina
plantationofcryptomeria,verywetbeingfedbyseepingwater,andonlysupportedby
tanglingtree-roots;inotherwords,itwasbynomeansagoodhabitat forharbouring
upperhypogeantrechines.Probablyduetothisunfavourablecondition,noadditional
specimenscameoutinspiteofcarefulinvestigationmadebythesecondauthor.
TrechinebeetlesprobablyreferabletoK.insperataarealsoknown fromtwosta-
tionsontheleftsideoftheTotsu-kawaValleyatthenortheasternpartofTotsukawa-
mura. 0neofthemislocatedatthesoutheasternfootofTenjiku-yamaontheleftside
oftheTaki-gawaValley,atributaryoftheTotsu-kawaRiver,andIsl3kmdistanttothe
north-northwestinabeelinefromtheKuma-dani,thetypelocalityofthespecies.The
otheroneliesintheHashi-taniGullyintheAsahi-gawadrainage,alsobelongingtothe
Totsu-kawaRiver,andis6.8kmfurthernorth-northwestfromtheTakigawalocality.
Onlyonemalehasbeentakenattheformerstation(fromadepthofabout80cm),
andonlyonefemalefromthelatter,buttheydoubtlessbelongtothesametaxonjudg-
ing fromtheirexternalmorphology includingthestandardratios. In themale from
Takigawa(5.20mminbodylength),thestandardratiosareasfollows:PW/HW143,
pw/PL 0.90 [PL/PW 1.11],PW/PA t39,PW/PB 150,PB/PA 0.97 [PA/PB 1.03],
Ew/PW191,EL/PL2.62,EL/EW152. Inthefemale fromtheHashi-tani(5.45mm
inbody length), theyare:PW/HW144,PW/PL0.89 [PL/PW 1.12],PW/PA t44,
pw/PB 145,PB/PA 0.99 [PA/PB 1.01], EW/PW 191, EL/PL2.55, EL/EW 149.