Two new genera and four new species of fossilweevils (Coleoptera: Curculionoidea) in Baltic amber

2016·2.4 MB·

by  Andrei Legalov

©EntomologicaFennica.31August2016
Two new genera and four new species of fossil weevils
(Coleoptera: Curculionoidea) in Baltic amber
AndreiA.Legalov
Legalov,A.A.2016:Twonewgeneraandfournewspeciesoffossilweevils(Co-
leoptera:Curculionoidea)inBalticamber.—Entomol.Fennica27:57–69.
Twonewgenera,Palaeodexipeusgen.n.(typespecies:Palaeodexipeuskirej-
tshukisp.n.)andSuccinalophusgen.n.(typespecies:S.attenboroughisp.n.),
andfournewspecies,Oxycraspeduspoinarisp.n.,Palaeodexipeuskirejtshuki
sp.n.,Dorytomuselectrinussp.n.andSuccinalophusattenboroughisp.n.,are
describedandillustrated.ThesearethefirstrecordsofthetribesOxycraspediniin
thefamilyBelidae,andStrombosceriniandTropiphoriniinCurculionidaeinthe
Balticamber.
A.A.Legalov,InstituteofSystematicsandEcologyofAnimals,SiberianBranch,
Russian Academy of Sciences,FrunzeStreet,11,Novosibirsk 630091,Russia;
E-mail:[email protected]
Received25September2015,accepted28October2015
1.Introduction Germany,aswellasintheBalticandDominican
amber(Legalov2015).
Curculioinoidbeetlesarewellrepresentedinthe In this paper, two new genera and four new
fossil record (Bouchard et al. 2011, Legalov speciesfromtheBalticamberaredescribed.
2015),probablyasaresultoftheirsolidintegu-
mentandabilityofmanygroupstofly.Fossilrep-
resentatives of wingless extant weevils are also 2.Materialsandmethods
knownprimarilyfromamber.Theoldestweevils
arediscoveredinlayersofthemiddle–lateJuras- TheBalticamberminesarelocatedalongtheBal-
sicofKaratau,andtheyrepresent57speciesfrom tic Sea coast at the Amber quarry of Yantarny
3 families (Gratshev & Legalov 2011, Legalov nearKaliningradintheKaliningradOblast(Rus-
2011, 2013b, Gratshev & Legalov 2014). The sia). The amber fromthis deposit was probably
number of the fossil specimens and described produced by the tree pine Pinus succinifera
speciesrapidlyincreasesthankstocurrentstud- (Goeppert)Conwentz(Schubert1961).TheBal-
ies,andespeciallyintheperiodbetweentheearly tic amber from this Prussian Formation dates
Cretaceous and the Neogene (Legalov 2012d). backtotheupperEocene(Grigjalisetal.1971),
The superfamily Curculionoidea is the major although sometimes it is considered older, even
groupofColeopterarepresentedintheCenozoic the Lowermost Eocene (Weitschat & Wichard
sedimentary deposit that exhibits extraordinary 2010).
fossils with exceptional preservation such as ThetypespecimensarestoredintheInstitute
GreenRiverandFlorissantbedsinUSA,Aix-en- ofSystematicsandEcologyofAnimals,theSibe-
Provance in France and Rott and Oeningen in rianBranchoftheRussianAcademyofSciences
58 Legalov (cid:127) ENTOMOL.FENNICAVol.27
(Novosibirsk, Russia). Comparative material is large, rounded, convex, 1.1 times longer than
depositedintheInstituteofSystematicsandEco- wide;vertexweaklyflattened,punctate;temples
logy of Animals, Museum für Tierkunde, short,0.8timesaslongaseye,punctate;antennae
Senckenberg Naturhistorischen Sammlungen inserted at base of rostrum ventrally, non-
Dresden(Germany)andtheZoologicalInstitute geniculate,quitelong,reachingfirstthirdofpro-
oftheRussianAcademyofSciences. notum; scape quite long, 2.4 times longer than
Descriptionsandphotographsweretakenus- wide;desmomeresalmostconical;1stdesmomere
ing the Zeiss Stemi 2000-C dissecting stereo- 1.6timeslongerthanwide,0.6timesaslongand
microscope. 0.9 times as wide as scape; 2nd desmomere 2.2
Thesystematicsofthestudiedtaxaaregiven timeslongerthanwide,1.2timesaslongand0.8
by Legalov (2009, 2015) and Gratshev and times as wide as 1stdesmomere; 3rd desmomere
Legalov (2014) with corrections by Zherikhin 1.8timeslongerthanwide,0.7timesaslongand
andEgorov(1991),Kuschel(1995a)andAnder- 0.9 timesaswideas2nd desmomere;4th desmo-
son (2002). The terminology of the names of mereequalto3rddesmomere;5thdesmomere1.4
body parts were used here after Klimaszewski timeslongerthanwide,0.9timesaslongand1.1
andWatt(1997)andAnderson(2002)withsome times as wide as 4th desmomere; 6th desmomere
corrections. equalto5thdesmomere;7thdesmomere1.3times
longerthanwide,1.1timesaslongand1.2times
aswideas6thdesmomere;clubnoncompact,0.6
3.Systematics times as long as flagellum; 1st club article 1.3
timeslongerthanwide,1.1timesaslongand1.2
3.1.DescriptionofOxycraspeduspoinari timesaswideas7thdesmomere;2ndand3rdclubar-
sp.n. ticlesfused,2.1timeslongerthanwide,2.6times
aslongand1.6timesaswideas1stclubarticle,
BelidaeSchoenherr,1826 weaklyacuminate.
OxycoryninaeSchoenherr,1840 Pronotum. Pronotum transverse, 1.4 times
OxycraspediniMarvaldi&Oberprieler,2006 longer than apical width, 0.9 times longer than
OxycraspedusKuschel,1955 wideinmiddleandatbase;disknarrowedatapex
andatbase,denselyandquitecoarselypunctate.
Oxycraspeduspoinarisp.n.(Fig.1) Mesonotum.Scutellumsmall,rectangular.
Typematerial.Holotype:no.BA2015/2,female. Elytra. Elytra weakly convex, slightly elon-
Diagnosis.ThisnewspeciesissimilartoO. gate,2.3timesaslongaspronotum,1.6timeslon-
cribricollis(Blanchard,1851)fromChileandAr- gerthanwideatbase,1.4timeslongerthanwide
gentina(Ferreretal.2007,Legalov2009)butdif- in middle, 2.1 times longer than wide in apical
fersintheprotibiaewithoutafringeoflongsetae, quarter; greatest width behind middle; humeri
abodyblack,amoreconvexformandwiderpro- flattened;puncturedstriaequiteregularanddis-
notumandelytra. tinct;scutellarstriolepresent;puncturesrounded,
Description. Female. Body length (without dense; intervals somewhat flat, narrow, slightly
rostrum)3.7mm;rostrumlength1.1mm. widerthanpunctures;epipleurondistinct.
Body black, naked, partially appearing sil- Thorax. Prosternum punctate, with lateral
veryshinyduetothepresenceofcavitiesbetween serrulate carina; precoxal part of prosternum
specimenandinternalsurfaceofitsimpression. weakly elongate, 1.2 times as long as procoxa
Head.Head0.4timesaslongasrostrum;ros- length; procoxal cavities round, separated;
trumlong,narrow,almostequalinlengthtopro- postcoxalpartofprosternumshort,0.5timesas
notum,5.2timeslongerthanwideatapexandin long as procoxa length; mesocoxal cavities
middle, 3.4 times longer than wide at base, rounded,separated;metasternumshort,1.9times
weakly convex, finely and densely punctate; as long as metacoxa length, convex, punctate;
foreheadwide,1.2timesaswideasrostrumbase metepisternumnarrow,denselypunctate.
width, flattened, densely punctate, without me- Abdomen. Abdomen weakly convex;
dianstriae,withouttuberclesoneitherside;eyes ventrites free; 1st ventrite 1.2 times as long as
ENTOMOL.FENNICAVol.27 (cid:127) NewfossilweeviltaxainBalticamber 59
Fig.1.Habitusof
Oxycraspedus
poinarisp.n.(holo-
type).–a.Dorsal
view.–b.Ventral
view.–c.Lateral
(right)view.Scale
bar=1mm.
metacoxallength;2ndventrite0.9timesaslongas ical;tibiaealmoststraight,weaklyflattened,wid-
1st ventrite; 3rd ventrite 0.9 times as long as 2nd ened at apices, with costate dorsal margin, with
ventrite; 4th ventrite 0.7 times as long as 3rd weak apical setose fringe, without mucro;
ventrite; 5th ventrite 1.5 times as long as 4th protibiaepossessingawideapicalgrooveclothed
ventrite. infine,densevestitureontheinnerfaceopposite
Legs.Legslong;femoraweaklyclavateand the tarsal articulation, without fringe of long
flattened, without teeth; profemur length/width setae, length/width ratio 5.1; mesotibia length/
ratio 2.9; mesofemur length/width ratio 2.9; widthratio4.0;metatibialength/widthratio5.1;
metafemurlength/widthratio2.8;trochantercon- tarsilong,withthicklighterectsetaedorsally;1st
60 Legalov (cid:127) ENTOMOL.FENNICAVol.27
tarsomeretriangular,shorterthanthesecondand 3.2.DescriptionofPalaeodexipeusgen.n.
thirdtarsomerescombined;2ndand3rdtarsomeres andanewspecies
widely bilobed; 5th tarsomere elongate; claws
large,connected,withoutteeth;protarsi:1sttarso- CurculionidaeLatreille,1802
mere 0.8 times longer than wide; 2nd tarsomere DryophthorinaeSchoenherr,1825
0.6timeslongerthanwide,1.6timesaslongas StrombosceriniLacordaire,1866
and2.0timesaswideas1sttarsomere;3rdtarso-
mere1.1timeslongerthanwide,2.1timesaslong Palaeodexipeusgen.n.
asand1.1timesaswideas2ndtarsomere;5thtarso- Typespecies:Palaeodexipeuskirejtshukisp.n.
mere3.6timeslongerthanwide,1.1timesaslong Diagnosis. This new genus is similar to the
as and 0.3 times as wide as 3rd tarsomere; genus Dexipeus Pascoe, 1885 from Japan, Java
mesotarsi: 1st tarsomere 0.7 times longer than andSumatra(Alonso-Zarazaga&Lyal1999)but
wide; 2nd tarsomere 0.7 times longer than wide, differs by the unusual carinate elytral intervals,
1.8timesaslongand1.9timesaswideas1sttarso- antennalclubweaklyobliquelytruncateandthe
mere; 3rd tarsomere 0.9 times longer than wide, rostrumnotwidenedattheantennalattachment.
1.6timesaslongand1.2timesaswideas2ndtar- From the Oriental genus Orthosinus Motschul-
somere;5thtarsomere2.7timeslongerthanwide, sky, 1863 and the genus Stenommatomorphus
1.1timesaslongand0.4timesaswideas3rdtar- Nazarenko, 2009 from Eocene Rovno amber it
somere;metatarsi:1sttarsomere0.8timeslonger differsbytheeyebeingmuchwiderthanhighand
than wide; 2nd tarsomere 0.8 times longer than thecarinateelytralintervals.Theobliquelytrun-
wide,1.8timesaslongand2.0timesaswideas1st cateantennalclubofthenewgenusseparatesit
tarsomere; 3rd tarsomere 1.1 times longer than from the Oriental genus Nephius Pascoe, 1885
wide,1.6timesaslongand1.1timesaswideas and the genus Tasactes Faust, 1894 from
2ndtarsomere;5thtarsomere2.5timeslongerthan Myanmar (Alonso-Zarazaga & Lyal 1999) that
wide,1.1timesaslongand0.5timesaswideas3rd have an antenna1 club with a subconical apex.
tarsomere. Thenewgenusisclearlydistinguishedfromother
Type locality and type strata. Amber mines genera of Stromboscerini (Besuchetiella Osella,
locatedalongtheBalticSeacoastandYantarny 1974, Dryophthoroides Roelofs, 1879, Para-
AmberquarrynearKaliningrad,Kaliningradre- synommatusVoss,1956,StromboscerusSchoen-
gion,Russia,upperEocene,PrussianFormation. herr, 1838, Synommatoides Morimoto, 1978,
Etymology.Thenameisdedicatedtomydear SynommatusWollaston,1873,Tetrasynommatus
colleague, Dr. George O. Poinar, Jr. (Corvallis, Morimoto, 1985) by the antennae with a 6-arti-
USA),whohasmadeagreatcontributiontothe cledflagellum.
studyofamberfauna. Etymology. The name is formed from the
Remarks. The new species belongs to the Greek“(cid:1)(cid:2)(cid:3)(cid:1)(cid:4)(cid:1)”(ancient)andthegenericname
familyBelidaebasedontheprotibiaepossessing Dexipeus.
a wide apical groove clothed in fine, dense
vestitureontheinnerfaceoppositethetarsalar- Palaeodexipeus kirejtshuki gen. n. et sp. n.
ticulation,thefirsttofifthventritesfreeandnon- (Figs.2–3)
geniculateantennae.Thetransverseandbilobed Typematerial.Holotype:no.BA2015/3,male.
secondtarsomere,antennaeinsertedatthebaseof Description.Male.Bodylength(withoutros-
the rostrum, the distinct club, first tarsomere trum)3.1mm;rostrumlength0.7mm.
shorterthanthesecondandthirdtarsomerescom- Body black, naked, appearing silvery shiny
bined,suggestplacementinthesubfamilyOxy- duetothepresenceofcavitiesbetweenspecimen
coryninae. Theseparated procoxal cavities sug- andinternalsurfaceofitsimpression.
gest that the new species belongs to the Head. Head 0.4 times as long as rostrum;
monotypicaltribeOxycraspediniwiththegenus mouthpartswithprementumwithdrawnintooral
Oxycraspedus. cavity;rostrumlong,quitenarrow,3.2timeslon-
ger than width at apex, 3.3 times longer than
width in middle, 3.0 times longer than width at
ENTOMOL.FENNICAVol.27 (cid:127) NewfossilweeviltaxainBalticamber 61
Fig.2.Lateral(left)viewofPalaeodexipeuskirejtshuki
sp.n.(holotype).Scalebar=1mm.
base, 0.9 times as long as pronotum, weakly
curved,finelypunctate;antennalscrobesdistinct;
forehead narrow, 0.8 times as wide as rostrum
base width, flattened, punctate, without median
striae;eyesquitesmall,flat,0.6timeslongerthan
wide, separated ventrally; vertex weakly flat-
tened,sparselypunctate;templesquitelong,1.7
timesaslongaseye,punctate;antennaeinserted Fig.3.FrontalviewofPalaeodexipeuskirejtshuki
infirstthirdofrostrumventrally,geniculate,quite sp.n.(holotype).Scalebar=1mm.
short,reachingfirstthirdofpronotum;scape3.3
times longer than wide; flagellum with 6 wide-
conical desmomeres; 1st desmomere 1.1 times timeslongerthanwideinapicalquarter;greatest
longerthanwide,0.3timesaslongand0.8times width in middle; humeri flattened; punctured
aswideasscape;2nddesmomere0.8timeslonger striaeregularanddistinct;puncturesroundedand
thanwide,0.7timesaslongandequalinwidthto deep;intervalsconvex,narrow,equalinwideto
1st desmomere; 3rd desmomere 0.4 times longer striawidth;oddintervalsascarinae.
thanwide,0.6timesaslongandequalinwidthto Thorax.Prosternumlargelypunctate,without
2nd desmomere; 4th desmomere 0.4 times longer postocular lobes; precoxal part of prosternum
thanwide,equalinlengthand1.1.timesaswide weakly elongate, 1.1 times as long as procoxa
as 3rd desmomere; 5th desmomere equal to 4th length; procoxal cavities round, connected;
desmomere;6thdesmomere0.3timeslongerthan postcoxalpartofprosternum0.7timesaslongas
wide,equalinlengthand1.2timesaswideas5th procoxalength;mesocoxalcavitiesrounded,sep-
desmomere; club compact and asymmetrical, arated; metasternum long, 3.3 times as long as
withelongated1starticleandfusedremainingar- metacoxa length, weakly convex, largely punc-
ticles,1.3timeslongerthanwide,almostequalto tate;metepisternumconcealedbyelytra.
funicle, weakly obliquely truncate, with tomen- Abdomen.Abdomenweaklyconvex;ventri-
toseapicalsurfaceflat. tesorientedinoneplane;1stand2ndventriteselon-
Pronotum. Pronotum 1.5 times longer than gateand3rdand4thventritesshort;1stventrite2.2
apicalwidth,1.2timeslongerthanwidthinmid- timesaslongasmetacoxallength;2ndventrite0.7
dleand1.1longerthanwidthatbase;diskweakly timesaslongas1stventrite;3rdventrite0.3times
narrowedatapexandatbase,denselypunctate. as long as 2nd ventrite; 4th ventrite equal to 3rd
Mesonotum. Scutellumsmall, narrow, trian- ventrite; 5th ventrite 2.4 times as long as 4th
gular. ventrite;pygidiumsulcate.
Elytra.Elytraconvex,elongate,2.1timesas Legs. Legs long; femora weakly clavate,
longaspronotum,1.7timeslongerthanwidthat without teeth; profemur length/width ratio 2.6;
base, 1.5 times longer than wide in middle, 2.3 mesofemur length/width ratio 3.5; metafemur
62 Legalov (cid:127) ENTOMOL.FENNICAVol.27
length/width ratio 4.3; trochanter conical; tibiae otherarticlesfused.Theconnectedprocoxalcav-
almost straight, flattened, with long uncus and ities, metepisternum concealed by the elytra,
two apical bunches of setae; protibiae length/ sulcate pygidium, antennal club weakly
widthratio4.4;metatibialength/widthratio5.3; obliquelytruncate,flattomentoseapicalsurface,
tarsilong,withthicklighterectsetaedorsally;1st andeyesseparatedventrally,suggestplacement
tarsomere conical; 2nd and 3rd tarsomeres not inthetribeStromboscerinifromtheRovnoam-
bilobed; 5th tarsomere elongate, with large free ber,MadagascarandOrientalregion.
claws without teeth, widely separated by lobes;
protarsi:1sttarsomereequalinlengthandwidth;
2nd tarsomere 0.9 times longer than wide, 0.9 3.3.DescriptionofDorytomuselectrinus
timesaslongandequalinwidthto1sttarsomere; sp.n.
3rd tarsomere 1.3 times longer than wide, 1.3
timesaslongand0.9timesaswideas2ndtarsome- CurculioninaeLatreille,1802
re;5thtarsomere3.0timeslongerthanwide,1.8 EllesciniC.G.Thomson,1859
timesaslongand0.8timesaswideas3rdtarsome- DorytomusGermar,1817
re;mesotarsi:1sttarsomere0.9timeslongerthan
wide; 2nd tarsomere equal in length and width, Dorytomuselectrinussp.n.(Figs.4–5)
equalinlengthand0.9timesaswideas1sttarso- Typematerial.Holotype:no.BA2015/5,female.
mere; 3rd tarsomere 1.3 times longer than wide, Diagnosis.ThisnewspeciesissimilartoD.
1.3timesaslongandequalinwidthto2ndtarso- luridus (Mannerheim, 1853) from North and
mere; 5th tarsomere 4.5 times longer than wide, WestofNorthAmerica(O’Brien1970)butdif-
1.8timesaslongand0.5timesaswideas3rdtar- fers by the antennae inserted beyond middle of
somere;metatarsi:1sttarsomere1.1timeslonger rostrum, mesosternal process convex, smaller
than wide; 2nd tarsomere equal in length and bodysize,nakedblackbody,largereyesandwide
width,0.8timesaslongand0.9timesaswideas elytra. It is distinguished from D. edoughensis
1sttarsomere;3rdtarsomere1.3timeslongerthan Desbrochers, 1875 from Europe and Siberia
wide,1.3timesaslongandequalinwidthto2nd (Dieckmann1986)bythenakedblackbody,wide
tarsomere; 5th tarsomere 4.5 times longer than elytra, antennae inserted beyond middle of ros-
wide,1.8timesaslongand0.5timesaswideas3rd trum,andwiderandshorterrostrum.
tarsomere. Description. Female. Body length (without
Type locality and type strata. Amber mines rostrum)2.4mm;rostrumlength0.8mm.
locatedalongtheBalticSeacoastandYantarny Body black,naked,withoutscalesandsetae
AmberquarrynearKaliningrad,Kaliningradre- appearing silvery shiny due to the presence of
gion,Russia,upperEocene,PrussianFormation. cavitiesbetweenspecimenandinternalsurfaceof
Etymology.ThenameisdedicatedtoDr.Al- itsimpression.
exanderG.Kirejtshuk(SaintPetersburg,Russia Head.Head0.4timesaslongasrostrum;ros-
andParis,France),forhisnumerousandinforma- trumlong,narrow,8.1timeslongerthanwideat
tivestudiesonamberbeetles. apex, 7.1 times longer than wide in middle, 5.2
Remarks.Thetibiaewithuncusandtwoapi- timeslongerthanwideatbase,1.3timesaslong
cal clusters of setae, ventrites oriented in one aspronotum,weaklycurved,finelypunctatedor-
plane, elongate first and second ventrites and sally,denselypunctatelaterally;antennalscrobes
shortthirdandfourthventrites,distinctantennal distinct, with the dorsal margin directed toward
scrobesandgeniculateantennaewithacompact ventralmarginofeye;foreheadnarrow,narrower
clubindicatethatthenewspeciesbelongstothe than rostrumbase width, flattened, finely punc-
familyCurculionidae.Thenewspeciesisplaced tate, without median striae; eyes large, flat, 0.7
inthesubfamilyDryophthorinaebasedonthe5th timeslongerthanwide,separatedventrally,dis-
tarsomere having claws widely separated by placed downwards; vertex weakly flattened,
lobes, mouthparts with prementum withdrawn rarelypunctate;templesshort;antennaeinserted
intooralcavity,antennalclubasymmetricaland beyond middle of rostrum laterally, geniculate,
the first article of the club elongated with the quite long, reaching basal third of pronotum;
ENTOMOL.FENNICAVol.27 (cid:127) NewfossilweeviltaxainBalticamber 63
Fig.5.Lateral(right)viewofDorytomuselectrinus
Fig.4.Lateral(left)viewofDorytomuselectrinus sp.n.(holotype).Scalebar=1mm.
sp.n.(holotype).Scalebar=1mm.
scape8.0timeslongerthanwidth,distinctlynot outpostocularlobes;precoxalpartofprosternum
reaching eye; flagellum with 7 conical desmo- fairlylong,0.8timesaslongasprocoxalength,
meres; 1st desmomere 2.3 times longer than without impression limited on each side of the
width,0.3timesaslongandalmostequalinwide longitudinal carinae; procoxal cavities round,
to scape; 2nd desmomere 1.5 times longer than connected; postcoxal part of prosternum short,
wide,0.4timesaslongand0.7timesaswideas1st 0.2 times as long as procoxa length; mesocoxal
desmomere; 3rd–5th desmomeres subequal to 2nd cavitiesrounded,separated;mesosternalprocess
desmomere;6thdesmomere1.4timeslongerthan convex; metasternum quite long, 1.2 times as
wide,1.2timesaslongand1.3timesaswideas5th longasmetacoxalength,weakly convex,rarely
desmomere;7thdesmomere1.1timeslongerthan largely punctate; metepisternum narrow, 5.7
wide,1.3timesaslongand1.6timesaswideas6th timeslongerthanwide,withrowoflargepoints.
desmomere;clubcompact,1.7timeslongerthan Abdomen. Abdomen weakly convex, flat-
wide, 0.4 times as long as funicle, 1.8 times as tenedinmiddle;ventritesorientedinoneplane;
wideas7thdesmomere. 1stand2ndventritesquiteelongateand3rdand4th
Pronotum. Pronotum 1.3 times longer than ventritesquiteshort;1stventrite0.8timesaslong
apicalwidth,0.9timeslongerthanwidthinmid- asmetacoxallength;2ndventrite1.1timesaslong
dleand0.8longerthanwidthatbase;diskweakly as1stventrite;3rdventrite0.7timesaslongas2nd
narrowedatapexandatbase,denselyandfinely ventrite;4thventrite0.9timesaslongas3rdven-
punctate;distancebetweenpointslargerthandia- trite;5thventrite2.0timesaslongas4thventrite.
meterofpoints. Legs.Legslong;femoraweaklyclavate,with
Mesonotum.Scutellumsmall,trapezoidal. teeth; profemur length/width ratio 2.4;
Elytra. Elytra convex, weakly elongate, 2.5 mesofemur length/width ratio 2.7; metafemur
timesaslongaspronotum,1.8timeslongerthan length/width ratio 2.9; trochanter conical; tibiae
wideatbase,1.5timeslongerthanwideinmid- almost straight, flattened, with small uncus dis-
dle,1.9timeslongerthanwideinapicalquarter; placed by inner apical angle and two apical
greatest width in middle; humeri weakly flat- bunchesofsetae;protibiaelength/widthratio4.7;
tened;puncturedstriaeregularanddistinct;punc- mesotibia length/width ratio 6.0; metatibia
turesroundedanddeep;intervalsweaklyconvex, length/widthratio5.7;tarsilong,withthicklight
wide,2.0–4.0timesaswideasstriawidth,very erectsetaedorsally;1sttarsomereconical;2ndand
finelypunctate;9thstriamergeswith10thstriaat 3rdtarsomeresnotbilobed;5thtarsomereelongate,
thelevelofmetacoxae;lateralmarginsofelytra withlargefreeclawswithoutteeth,widelysepa-
not arcuately produced laterally at the overlap- ratedbylobes;mesotarsi:1sttarsomere1.2times
pingpointoftheapexofmetepisternumandfilled longer than wide; 2nd tarsomere equal in length
withsetaeinternally andwidth,0.8timesaslongandequalinwidthto
Thorax. Prosternum densely punctate, with- 1st tarsomere; 3rd tarsomere equal in length and
64 Legalov (cid:127) ENTOMOL.FENNICAVol.27
width,1.4timesaslongand1.4timesaswideas terseparatesitfromallothergeneraofthistribe.
2ndtarsomere;5thtarsomere2.7timeslongerthan Therearethreeextantgenerawithasmallbody
wide,1.2timesaslongand0.4timesaswideas3rd size (2.8–4.6 mm). The new genus differs from
tarsomere;metatarsi:1sttarsomere1.7timeslon- BeringianVitaviyusKissinger,1974andLepido-
gerthanwide;2ndtarsomere1.3timeslongerthan phorus W. Kirby, 1837 from North America
wide,equalinlengthand1.3timesaswideas1st (O’Brien & Wibmer 1982) and Chukchi Penin-
tarsomere; 3rd tarsomere equal in length and sula(Korotyaev1980)bymandiblesbeingwith
width,1.4timesaslongand1.8timesaswideas distinctprominentscarsattheattachmentofthe
2ndtarsomere;5thtarsomere3.7timeslongerthan deciduousprocesses,thescapereachingthedistal
wide,1.6timesaslongand0.4timesaswideas3rd marginoftheeyeandtheprosternumbeingwith-
tarsomere. out an ocular lobe. From Limalophus Scudder,
Type locality and type strata. Amber mines 1893fromthemiddleEoceneofUSA,itisdistin-
locatedalongtheBalticSeacoastandYantarny guishedbythelongerrostrumandpronotum.The
AmberquarrynearKaliningrad,Kaliningradre- newgenusisverysimilartotheEuropeangenus
gion,Russia,upperEocene,PrussianFormation. TropiphorusSchoenherr,1842butdiffersbythe
Etymology.ThenameisformedfromtheLa- freetarsalclaws,narrowerbodyandlongerpro-
tinwordfor“amber”–“electri”. notum.
Remarks.Thetibiaewithuncusandtwoapi- Etymology.ThenameisformedfromtheLa-
cal clusters of setae, ventrites oriented in one tinwordfor“Amber”–“succinum”andthege-
plane, elongate first and second ventrites and nericnameAlophus.
shortthirdandfourthventrites,distinctantennal
scrobesandgeniculateantennaewithacompact Scuccinalophusattenboroughigen.n.etsp.n.
clubindicatethatthenewspeciesbelongstothe (Fig.6)
familyCurculionidae.Thenewspeciesisplaced Typematerial.Holotype:no.BA2015/1,male.
inthesubfamilyCurculioninaebasedonthetib- Description.Male.Bodylength(withoutros-
iaewithsmalluncusdisplacedbyinnerapicalan- trum)3.1mm;rostrumlength0.5mm.
gleand9thstriamergingwith10thstriaatthelevel Body black, with very dense short, wide,
ofmetacoxae.Theantennalscrobesdirectedto- grey-brown contiguous and semierect setiform
wardtheeye,precoxalportionoftheprosternum scales, appearing silvery shiny due to the pres-
elongated, metafemora weakly clavate, not ence of cavities between specimen and internal
reachingtheapexoftheabdomen,flagellum7-ar- surfaceofitsimpression.
ticled, claws free, procoxal cavities connected Head.Rostrumquiteshort,wide,0.8timesas
andthetibiaewithuncusplacesthenewspecies longaspronotum,2.0timeslongerthanwidein
inthetribeEllescini.Thefemorawithteethand middle, weakly curved, densely punctate, with
tarsal claws without teeth indicate that the new twosharptransversecarinae,pterygiawelldevel-
speciesbelongstothegenusDorytomus. oped; scrobes distinct, visible dorsally, directed
to upper edge of eyes; forehead wide, convex,
denselypunctate,withoutstriae;eyessmall,0.6
3.4.DescriptionofScuccinalophusgen.n. timeslongerthanwide,barelyconvex,displaced
andanewspecies downwards; vertex weakly flattened, densely
punctate;templesveryshort;mandibleswithdis-
EntiminaeSchoenherr,1823 tinctprominentscarsofattachmentofdeciduous
TropiphoriniMarseul,1863 processes;geniculateantennaeinsertedatapical
thirdofrostrumlaterally;antennaelong,reaching
Scuccinalophusgen.n. basal third of pronotum, with rare semierect
Type species: Scuccinalophus attenboroughi setae;scape7.5timeslongerthanwide,1.1times
sp.n. as long as flagellum; desmomeres conical; 1st
Diagnosis. This new genus is characterized desmomere2.0timeslongerthanwide,0.2times
by eyes that are strongly displaced downwards as long and 0.8 times as wide as scape; 2nd
andasmallbodysize(3.1mm).Thefirstcharac- desmomere1.1timeslongerthanwide,0.5times
ENTOMOL.FENNICAVol.27 (cid:127) NewfossilweeviltaxainBalticamber 65
Fig.6.Lateral(right)
viewofSuccinalophus
attenboroughisp.n.
(holotype).Scalebar
=1mm.
aslongand0.9timesaswideas1stdesmomere;3rd joined; postcoxal part of prosternum short, 0.5
desmomere1.1timeslongerthanwide,0.9times timesaslongasprocoxalength;mesocoxalcavi-
aslongand0.9timesaswideas2nddesmomere; ties rounded, narrowly separated; metasternum
4th desmomere equal to 3rd desmomere; 5th long,1.7timesaslongasmetacoxalength,con-
desmomereequalto2nddesmomere,1.1timesas vex, punctate; metepisternum narrow, 8.4 times
longand1.1timesaswideas4thdesmomere;6th longerthanwide.
desmomere1.3timeslongerthanwide,1.3times Abdomen.Abdomenflattened;ventritesori-
aslongand1.1timesaswideas5thdesmomere; entedinoneplane;1stand2ndventriteselongate,
7th desmomere 1.3 times longer than wide, 1.1 subequalinlength;1stventrite1.5timesaslongas
timesaslongand1.1timesaslongas6thdesmo- metacoxa; 3rd and 4th ventrites short, equal in
mere;clubcompact,1.3timeslongerthanwide, length; 3rd ventrite 0.4 times as long as 2nd ven-
0.5timesaslongasflagellum,weaklyacuminate. trite;5thventrite1.6timesaslongas4thventrite.
Pronotum. Pronotum bell-shaped, 1.3 times Legs.Legslong;femoraandtibiaewithdense
longer than apical width, 1.0 times longer than broad scales and rarely semierect setae; femora
wideinmiddleand1.2longerthanwideatbase, weakly clavate, without teeth; profemur
0.8timesaswideaselytralbase;greatestwidthin length/width ratio 3.8; mesofemur length/width
middle; disk narrowed at apex and at base, ratio 4.0; metafemur length/width ratio 3.6;
denselypunctate,withoutocularlobes. trochanterconical;tibiaealmoststraight,weakly
Elytra.Elytraweaklyelongateanddistinctly flattened,widenedatapices,withmucroandapi-
convex, 2.1 times longer than wide at base, 2.0 caldarksetosefringe,withoutuncus;metatibial
timeslongerthanwideinmiddle,2.5timeslonger corbels open; protibia length/width ratio 4.3;
thanwideinapicalquarter,2.3timesaslongas mesotibia length/width ratio 4.5; metatibia
pronotum;greatestwidthbehindmiddle;humeri length/widthratio4.6;tarsilong,withthicklight
completely flattened; punctured striae regular erectsetaedorsally;1stand2ndtarsomerestrian-
and distinct;punctureselongate-oval, smalland gular; 3rd tarsomere bilobed; 5th tarsomere elon-
dense;intervalsflat,wide,6.0–7.0timesaswide gate;clawslarge,free,withoutteeth;protarsi:1st
asstriae,withmiddlerowofsetiformsemierect tarsomere1.3timeslonger thanwide;2ndtarso-
scales. mere0.9timeslongerthanwide,0.6timesaslong
Thorax. Prosternum densely punctate; pre- and0.9timesaswideas1sttarsomere;3rdtarso-
coxal part of prosternum elongate, 1.1 times as mereequalinlengthandwidth,1.5timesaslong
longasprocoxalength;procoxalcavitiesround, and1.3timesaswideas2ndtarsomere;5thtarso-
66 Legalov (cid:127) ENTOMOL.FENNICAVol.27
mere2.8timeslongerthanwide,1.6timesaslong wasdescribedfromtheMioceneDominicanam-
and0.6timesaswideas3rdtarsomere;mesotarsi: ber (Poinar & Legalov 2014b). Several extinct
1sttarsomere1.2timeslongerthanwide;2ndtarso- speciesofthetribeMetrioxenini,widespreadin
mere0.8timeslongerthanwide,0.7timesaslong theOrientalregion,areknowfromtheBalticam-
and1.1timesaswideas1sttarsomere;3rdtarso- ber (Legalov 2012a, 2012b, 2013a). The tribe
mereequalinlengthandwidth,1.6timesaslong Oxycraspedini with the genus Oxycraspedus
and1.2timesaswideas2ndtarsomere;5thtarso- fromChileandArgentinadiffersfromtheSouth
mere3.5timeslongerthanwide,1.8timesaslong American tribe Oxycorynini by the separated
and0.5timesaswideas3rdtarsomere;metatarsi: procoxal cavities. The specimens of Oxycra-
2ndtarsomere0.6timesaslongaswideas1sttarso- spedini can be identified as representatives of
mere;3rdtarsomere1.6timesaslongas2ndtarso- Metrioxeninibytheheadbehindeyesbeingwith-
mere;5thtarsomere1.5timesaslongas3rdtarso- out a groove and the pronotum being without
mere. discal carinae. Three extant species of Oxycra-
Type locality and type strata. Amber mines spedusfromChileandArgentinadevelopinfe-
locatedalongtheBalticSeacoastandYantarny male strobili of Araucaria araucana (Kuschel
AmberquarrynearKaliningrad,Kaliningradre- 1995b,2001,Ferreretal.2007).TheBalticam-
gion,Russia,upperEocene,PrussianFormation. berspeciesdevelopedpossiblyalsoinAraucaria.
Etymology.ThenameisdedicatedtoSirDa- Twenty fossil species of the subfamily
vidF.Attenborough(London,UK),forhisgreat Dryophthorinaeareknown(Legalov2015).The
contributionofpopularizingnaturalhistory. firstrecordsofthissubfamilyarebasedonage-
Remarks.Thenewgenusandspeciesbelong nusofthetribeStromboceriniintheRovnoam-
tothefamilyCurculionidaebasedonthefollow- ber (Nazarenko & Perkovsky 2009) and undes-
ing characters: ventrites oriented in one plane, cribedspeciesofthetribeOrthognathini(Schlee
elongatefirstandsecondventritesandshortthird 1990).Dryophthorinaearefoundintheterminal
andfourthventrites,distinctantennalscrobesand EoceneoftheFlorissantbeds(Scudder1893)and
geniculate antennae with a compact club. The intheMioceneDominicanamber(Davis&Engel
mandibleswithprominentscarsofattachmentof 2006, 2009; Poinar & Legalov 2014a, 2015b).
deciduousprocesses,rostrumquiteshortandtib- Palaeodexipeuskirejtshukigen.n.etsp.n.isthe
iae without uncus, suggest placement in the first record of the tribe Strombocerini, and the
subfamilyEntiminae.Theeyesdisplaceddown- first described representative of the subfamily
wards,rostrum2.0timeslongerthanwide,tarsal DryophthorinaeintheBalticamber.
claws free and lateral antennal scrobes indicate The oldest Curculioninae from the tribe
that the new genus belongs to the tribe Tropi- CurculioniniaredescribedfromthePaleoceneof
phorini. France(Piton1940),becausetheearlierrepresen-
tativedescribedinthissubfamily (Poinar2009)
belongs to another family (Legalov & Poinar
4.Discussion 2015).Membersofthissubfamily areabundant
in the terminal Eocene of Florissant (Scudder
Representing the family Belidae, the subfamily 1893; Legalov 2015) and Miocene Dominican
OxycoryninaedatesbacktotheearlyCretaceous amber(Poinar&Legalov2015a).Speciesofthe
ofSpain(Gratshev&Legalov2014).Tentribes generaSuccinostyphlusKuska,1996,Dorytomus
(Afrocorynini,Aglycyderini,Allocorynini,Allo- Germar,1817andPachytychiusJekel,1861from
xycorynini, Archicorynini, Distenorrhinoidini, the tribe Ellesini have been described from the
Metrioxenini, Oxycorynini, Oxycraspedini and Balticamber(Kuska1996,Legalovinpress).
Palaeorhopalotriini) are known (Legalov 2009, The tribe Tropiphorini is distributed world-
2013a; Anderson & Marvaldi 2013). The tribes wide. The first finds are in the middle Eocene
Distenorrhinoidini (Berriasian–Barremian of Green River (Scudder 1893). Some species are
Montsec)andPalaeorhopalotriini(upperEocene describedfromtheterminalEoceneofFlorissant
of Ale`s-Monteils) are extinct (Legalov 2009, (Scudder 1893; Wickham 1911) and Oligocene
2013a). One fossil representative of the genus of France (Zherikhin 1992). Scuccinalophus