Three additional new genera and two replacement names for northeastern Pacific prosobranch gastropods

3 Pages·1995·1.1 MB·English

THE NAUTILUS 108(3):80-82, 1995 Page 80
Three Additional New Genera and two Replacement Names for
Northeastern Pacific Prosobranch Gastropods
James H. McLean
Los Angeles County Museum of
Natural History
900 Exposition Blvd
Los Angeles, CA 90007 USA
ABSTRACT Remarks: Sculpture of typical speciesof Margarites and
species of other available subgenera of Margarites is
Three new genera or subgenera and two replacement names
fTorrocshpiedcaiee:sCfoisotommoanrygmasriatrees.pnroepwosseudb.geNnueswofgMeanrugsarlieJvfesl;ttayxpae; sfpoirraglr,otwotthheienxccrleumseinotns.ofTahxiealsstcruolnpgtauxriealalstcougleptthuerreexofcejput-
species Trochus costalis Gould. 1841. Calyptraeidae: Grandi- venile shells of Margarites (Costomargarites) costalis
crepidula, new subgenusof Crepidula. type species Crepidula suggests a closer resemblance to Solariclla obscure
grandis Middendorff, 1849. Turridae, Crassispirinae: Pseudo- (Couthouy, 1838) than to other species of Margarites.
taranis new genus; type species Mangelia (Tarants) strongi The species in which axial sculpture dominates the mor-
Arnold, 1903. New names for homonyms: Trochidae: Marga- phology of the early teleoconch (and may or may not be
rites (Costomargarites) baxteri, new name for Margarites expressed in the mature shell) need to be distinguished
(Pupillaria) rudis Dall, 1919, not Margarita groenlandica var. at least at the subgeneric level. For many years I have
rudis Morch, 1869 Turridae: Leucosyrinx hantori, new name been surprised that this common Arctic species has not
for Antiplanes aniycus Dall, 1919, not Leucosyrinx amycus been made the type species of a subgenus, a gap that I
Dall, 1919.
hereby
fill.
Keywords: Mollusca; Prosobranchgastropods;Trochidae;Ca-
lyptraeidae; Turridae; new species; new names Family CALYPTRAEIDAE Lamarck, 1809
Grandicrepidula McLean, new subgenus
INTRODUCTION
Type species: Crepidula grandis Middendorff, 1849.
Thispaperfollowsa—previouspaper(McLean, 1995) with Included species: Type species [Boreal Pacific]; Crepi-
the same objective to make some generic level names dula princeps Conrad, 1856 [early Miocene through
available for other work in progress. Additionally, the middle Pliocene of southern California]; C. excavate
opportunity is taken to provide replacement names for (Broderip, 1834) [Panamic],C. maculosa Conrad, 1846
two homonvms. [Western Atlantic]; C. convexa Say, 1822 [Western At-
lantic].
Diagnosis: Beakexcavated, projecting posteriorlyon right
NEW GENUS LEVEL TAXA side;septal margin nearlystraight, extendingfartherfor-
Family TROCHIDAE Rafinesque, 1815 ward on left side; single muscle scar on right side, rect-
angular or chevron-shaped.
Subfamily MARGARITINAE Stoliczka, 1868
Remarks: Hoagland (1977) reviewed the Recent and
Costomargarites McLean, new subgenus
fossil species of Crepidula. treating them in alphabetical
Type species: Trochus costalis (iould, 1841. order, while not using the available subgenera. Surpris-
ingly, none of the 14 available taxa (see Hoagland, 1977:
Included species: Type species [Arctic, North .Atlantic
399) has a type species in which the septum projects on
andNorth Pacific]and Trochusochotensis Philippi, 184(i
[Northwestern Pacific]. The complex synonymy for the the right s—iiie. It is evident that this condition is not
type species was outlined by Abbott (1974;36). The ju- ambiguous all species illustrated b\ Hoagland can be
assigned on the basis ol this character. These diagnostic
venile shell was illustrated by Pilsbry (1889, pi. 60, figs characters were mentioned and show clearly in Hoag-
27, 28).
land's drawings of septal configuration (figure 2B, C.
Diagnosis: Subgenus of Margarites in which early tc- excavate: figure 2F, C. gratidis). Moreover, the five spe-
leoconch .sculpture is axial rather than spiral. cies assigned above key out together in Hoaglaml s key
,
H. McLean, 1995 Page 81
J.
to the species (page 363, couplets 16-18). Contrary to Remarks: Margarites(Costomargarites)baxteri was fig-
Hoagland s assertion that the division of Crepidula into ured by Dall (1921:179, pi. 18, fig. 13, 14) and by Kosuge
subgenera is not warranted, I find it useful to base su- (1972, pi. 2, fig. 5), but has otherwise been ignored in
praspecificdistinctionson themorphologyoftheseptum, the literature. It will be treated by me in work in prep-
particular!} when considering fossil species. In my opin- aration asa geographicsubspecies (in theGulfof Alaska)
ion the remaining species can also be assigned to the of theboreal M. (Costomargarites)costalis (Gould, 1841).
available taxa on shell characters. The new name honors the late Rae Baxter for his efforts
at reviewing Alaskan mollusks, cut short by his untimely
Family TURRIDAE Swainson, 1840 death in 1991.
Subfamily CRASSISPIRINAE Morrison, 1966 Family TURRIDAE Swainson, 1840
Pseudotaranis McLean, new genus
Subfamily COCHLESPIRINAE Powell, 1942
Type species: Mangelia (Taranis)strongi Arnold, 1903. LeucosyrinxkantoriMcLean, new nameforAntiplanes
Included species: Type species and the more slender anujeus Dall, 1919a (p. 36, pi. 11, fig. 5), notLeucosyrinx
Antiplanes hijperia Dail, 1919a (p. 35, pi. 9, fig. 6). The amyctis Dall, 1919a (p. 5, pi. 3, fig. 7).
type species was described from the Lower Pleistocene
Remarks: Theproposalof Leucosyrinxkantori remedies
of San Pedro, California; it was also illustrated by Grant a problem of secondary homonymy that was initiated
& Gale(1931:572, pi. 26. fig. 37);Borsonia inculta Mood) when Dall (1919a)described aspeciesinAntiplanes that
s1p9e1c9ies(p.(P5s4e,itpdio.ta1r,afniigss.s2tar,on2gbi)ainsdaPs.>hnyopne\rnii.a)Tlhivee tofwfo- I18n8o9.wDcaolnlsi(d1e9r19tao)bealasotprureopmoesmedbearspoefciLeesucwoistyhrtihnexnDaalmle,
shore in moderately deep water (100-400 m).
Leucosyrinx amycus. Note that both species were de-
Diagnosis: Shell small (length to 17 nun), spire high, scribed in the same paper. Previously (McLean in Keen,
anterior canal short, whorls 6. Axial sculpture lacking, 1971:713), I placed Leucosyrinx amycus Dall, 1919 in
spiralsculptureof twocordsemergingon teleoconch and the synonymy of Aforia goodei (Dall, 1890). The new
three cords on base; anal sinus shallow, at periphery and name honors Yuri Kantor for his recent work on turri-
coinciding with uppermost cord; lip not projecting. Pro- form gastropods.
toconch paucispiral, of 1.2 low, rounded whorls. Radula
of marginal teeth attached to membrane; teeth of long, LITERATURE CITED
flat type.
Remarks: Authors prior to 1971 used Taranis Jeffreys, Abbott, R. T. 1974. AmericanSeashells. Second Edition. New
1870 for the type species, but the presence of a radula Dall,YWor.k:H.Van19N19oas.traDnedscRreiipnthioolnds,of66n3epwp.species of mollusks
precludes placement in that genus, in which the radula of the family Turritidae from the west coast of America
is lacking (see Powell, 1966:55). Earlier (McLean, 1971; and adjacent regions. Proceedings of the United States
120, fig 40), I illustrated the radula of strongi and as- National Museum 56(2288):1-86, pis. 1-24.
signed it to Antiplanes (Rectisulciis), a genus now in the Dall, W. H 1919b Description of new species of Mollusca
subfamily Cochlespirinae, according to Ta\lor et al. from the North Pacific Ocean in the collection of the
(1993). Pseudotaranis differs in having the sinus more UnitedStatesNationalMuseum. ProceedingsoftheL'nited
shallow, theliplessprotracted, theanteriorcanalshorter, SWtates National Museum 56(2295):29;3-371.
and the protoconch more compressed. The radula lacks Dall, H, 1921. Summary ofthemarineshell-bearingmol-
the vestigial rachidian tooth and the marginals are not lusks of the northwest coast of America, from San Diego,
California, to the Polar Sea, mostly contained in the col-
of the bifurcated type indicated for Antiplanes by Kan-
tor and Sysoev (1991:122). The flat morphology of the ltreacttiioonnsofofthheitUhnerittoedunSftiagteusreNdatsipoenciaels.MuUsneiutemd,SwtiatthesilNlaus--
lateral teeth agrees with the subfamily Crassispirinae, as tional Museum, Bulletin 112, 217 pp., 22 pis.
defined by Taylor et al. (1993). Thus, the radular dif- Grant,U.S,IVand H. R Gale 1931. Catalogueofthemarine
ferencesplaceAntiplanesandPseudotaranisindifferent Pliocene and Pleistocene Mollusca of California and ad-
turrid subfamilies. However, on shell characters, Pseu- jacent regions. Memoirs of the San Diego Society of Nat-
dotaranis is atypical of Crassispirinae in not having pro- ural History 1:1-1036, pls.1-32.
nounced callus developed near the anal notch. Hoagland, K. E. 1977. Systematic review of the fossil and
Recent Crepidula and discussion of evolution of the Ca-
lyptraeidae. Malacologia 16(2):353-420.
NEW NAMES FOR SPECIES LEVEL HOMONYMS Kantor, Y. I, and A. V. Sysoev. 1991. Mollusks of the genus
Antiplanes (Gastropoda: Turridae) of the northwestern
Pacific Ocean. The Nautilus 105(4):119-146
Family TROCHIDAE Rafinesque, 1815 Keen, A, M. 1971. Seashellsoftropicalwest America. Second
Subfamily MARGARITINAE Stoliczka, 1868 Edition. Stanford Universitv Press, Stanford, 1064 pp.,
22 pis.
Margatites (Costomargarites) baxteri McLean, new- Kosuge, S. 1972. Illustrations of type specimens of Molluscs
name for Margarites (Pupillaria) rudis Dall, 1919b:364, described by William Healey Dall. National Science Mu-
not Margarita groenlandica var. rudis Morch, 1869:23. seum, Tokyo, 29 pis. and captions.
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andsubgenerafromtheEasternPacific. TheVeliger 14(1): Powell, A. W. B. 1966. The molluscan families Speightiidae
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