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The winter crane flies of North America north of Mexico (Diptera: Trichoceridae) PDF

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Preview The winter crane flies of North America north of Mexico (Diptera: Trichoceridae)

PROC. ENTOMOL.SOC.WASH. 105(4).2003.pp.901-914 THE WINTER CRANE FLIES OF NORTH AMERICA NORTH OF MEXICO (DIPTERA: TRICHOCERIDAE) H.^RRY D. Pratt 104 South Almond. Simpsonville. SC 29681, U.S.A. (e-mail: [email protected]) — Abstract. Twenty-eight .species of winter crane flies in the family Trichoceridae are found in North America north of Mexico; Diazosmapratti Stary. D. siihsinuatum Alex- ander, Paracladura trichoptera (Osten Sacken), Trichocera (Metatrichoceni) coleiAlex- ander, T. (M.) garretti Alenander. T. (M.) liiteci Becher. T. (A/.) mackenzie (Dahl), T. (M.) salmaniAlexander, T. (M.)tetoncnsisAlexander, T. (M.)ursamajorMc\i\ndci\Trichoceni (Trichocera) annulata Meigen, T. (T.) arctica Lundstrom. T. (T.) aniaudi Pratt, n. sp., T. (T.) haiifft Pratt, n. sp., T. (T.) bimacula Walker, T. (T.) bituherciilata Alexander, T. (7.) borealis Lackschewit/, T. (T.) brevicornis Alexander, T. (T.) coliimbiana Alexander, T. (T.) excilis Dahl, T. (T.) faitigiaiia Alexander, T. (T.) gracilis Walker, T. (T.) hieimilis (De Geer), T. (T.) idahoensis Pratt, n. sp.. T. (T.) maculipenuis Meigen, T. (T.)pollens Alexander, T. (T.) regelationis (Linnaeus), and T. (T.) setosivena Alexander. Trichocera loiigisetosa Alexander is a new synonym of Trichocera {Trichocera) setosivena Alex- ander. Keys to genera and species, notes on distributions, and illustrations are provided. Kcx Words: Diptera, Trichoceridae. winter crane flies. North America, taxonomy Winter crane flies of the genus Tricho- collected by net. light trap. Malaise trap, cera Meigen are found on warm sunny af- and molasses trap. ternoons in fall, winter, and spring in the The following acronyms indicate the lo- contiguous United States, Canada, and cation oftypes: Alaska. Ontheotherhand,adultsofthetwo ANSP Academy of Natural Sciences, species ofDiazosma Bergroth are collected Philadelphia, PA.. USA. from June to September, the only trichocer- BMNH The Natural History Museum, idsonwinginthesummerinnorthern Unit- London, UK. ed States. Adults of Paracladura trichop- CAS California Academy of Sciences, tera (Osten Sacken) are found from August San Francisco, CA, USA. through the winter to spring in British Co- CNC Canadian National Collection, lumbia. Washington, Oregon, and Califor- Ottawa. ON, Canada. nia. Swarms of Trichocera males are often LSI Linnean Society ofLondiMi. Lon- seen dancing in the late afternoon sunlight, don, UK. sometimes thousandsofindividuals in hun- MCZ Museum of Comparative Zoolo- dreds ofswarms over many acres oflawns gy, Harvard University, Cam- and open woodlands. Biologists also have bridge, MA, USA. noticed these coldhardy insects swarming MNHN Museum National d'Histoire Na- above the snow, or specimens crawling on turelle, Paris, France. the snow when temperatures were between NMW NaturhistorischesMuseum,Wien, 0° and 10°C. Specimens in this study were Austria. 902 PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON LLSNM National MuseumofNatural His- because their types are lost. They are not tory, Smithsonian Institution, treated further here. Trichocera gracilis Washington. DC, USA. Walker (1848) was described from a single ZIL Zoological Institute, Lund, Swe- female from York Factory, Manitoba, Can- den. ada. Byers (1976) published notes and fig- ZISP Zoological Institute, Academy of ures ofthe terminal abdominal segmentsof .Sciences, St. Petersburg, Russia. thetype. I knowofnocharacterstoseparate this species from females ofother species. Taxonomy' Alexandei' published twokeys totheTri- Family Characteristics of the choceridae found in the eastern United Trichoceridae States (1919a, 1942) and one for the west- Winter crane flies in the family Tricho- ern United States (1967). He wrote a fine ceridaearesinallto medium sized (wingup summary ofthe family Trichoceridae in the to 12 miTi in Diazosma) with slender long "Manual of Nearctic Diptera" in 1981. legs similar to the true crane flies in the Dahl published many papers on winter family Tipulidae. Trichoceridae differ in crane flies. Dahl's 1966 publication on the having 3 ocelli, the V-shaped suture on the Swedish Trichoceridae includes much in- mesonotum incomplete in the middle, and formationonseveral speciesfound inNorth long hairlike antennae. Tipulidae lackocel- America, and her 1967 paperon arctic and li, have a complete V-shaped suture on the subarcticTrichoceridaeisthebasicworkon mesonotum separating the praescutum and northern species. The world catalogue of scutum, and antennae usually short. True Trichoceridae by Dahl and Alexander cranefliesareoftenmuchlargerthanwinter (1976) lists all the species known to 1976, crane flies, with wings up to 25 mm or references to the original descriptions, and more. — known locations of types of most species. Description. Head with 3 ocelli. Anten- Pratt and Pratt (1984) published keys, illus- na with 16 ""segments," scape and pedicel trations and data on ten species of eastern enlarged, flagellomeeres 3-16 elongate and North America. Pratt (1992) published a progressively more slender. Mouthparts re- key to the Trichoceridae ofNorth America. duced, mandibles absent, galea of maxilla Stary (1995) described Diazosma pratti well-preserved, palpus with 4 free seg- and wrote that Diozosnui hirtipeivie (Sieb- ments, no trace ofsensory pore (as in 5v/- ke) is a European species and that North vicola Harris and Culicoides Latreille). American records of this insect should be Thorax with V-shaped suture separating called D. sitbsiiniatiiin (Alexander). Stary praescutun and scutum incomplete in mid- and Martinovsky (1996) wrote that Tricho- dle, and no distict suture separatingscutum cerajaponicu Matsumura (1916) probably and scutellum. Wing with Sc long joining is not a North American species and that C at midlength, 4 branches of R and 3 M records of this species by Dahl and Alex- branches of reaching wing margin. A, ander (1976) and Pratt (1992) should be long. A, short and curved to wing inargin called Trichocera excilis Dahl (1967). in Trichocera and Paracladitra Brunetti Krzeminska (2001) wrote that Trichocera longer and subsinuous in Diazosma. Legs major Edwards is a European species and not markedly deciduous as in Tipulidae: that American records of this species Abdomen with 8 distinct pregenital seg- should be called Trichocera setosivena Al- ments, first segment short. Male genitalia exander. with medio-basal projections of gonocox- Trichocera scutellata Say (1824) from itesformingincompleteorcompletebridge, Minnesota and Trichocera hrumalis Fitch gonostylus single, phallosome with aedea- (1847) from New York are unrecosinized gus or sperm pump somewhat pearshaped VOLUME 105. NUMBER4 903 and parameres joined medially by thin portionofgonostylusbyondlobewithsparse membrane. Female with ovipositor formed long setae; bridge connecting gonocoxites by sclerotized, elongate, downward curved withpointedprotuberanceinmiddle(Fig. 10) (AK.OR,northwesternU.S.)(Fig. 10) .... cerci in Trichocera and Panuhuliira or tetonensisAlexander short and fleshy in Diazosnia. 7. Lobeofgonostylusatba.sal fourthofgonos- Key to the Winter Crane Flies btyrluussh;loafterlaolngl,obreesddoifshninhtahirster9glihteswteirtnhitdeenHsael (Trichoceridae) of North America (Fig. II)(NH.MA.MDwestto KS) North of Mexico salmaniAlexander - Lobeofgonostylusarisingatbaseofgonos- tylus; later lobes ofninth tergite with a few I Tarsiwithbasitarsomereveryshort,aboutone- sparse hairs; ninth sternite balloon-like with eighth as long as second tar.somere (Fig. 4): setosedorsal lobe (Fig. 12) (AK. northwest- wing with m-cucrossveinpresent(Fig. I);fe- ernCanada) niackcnzieDahl cmkaulheiiaw.itohneonsepecsipeesi.maBtCh.ecWaA.(FiOg.R,32C)A)(Pc.im.-. 8. lBaarsgaeliarrnedgudliasrtarloupnodrteidonlsoboef(Fgiogn.os1t4y)lu(sGreweint-h - Tarsi withPabrausciltaadristormaerIehclhoonpgteerrath(aOnstseencoSnadcken) Blaansde)ofgonostylus without large rouInudrealoBbeecher (Fig. 5); wing with m-cu crossvein absent 9 2. e(WFviiengnsgl.y2w-it3ot)h;wiAfn,egmlaomlnaegr,gwiistnuhb(s3Fiinsgu.poeu2rs)m,;attnhiobeticalaceusrpv.ue.rd.s 2 9. T(Fiipg.ofIIg)on(oAsKt.yluBsC,wiWtAh,laOrRge, rCoAcu)onldeeidAlleoxbeander (abDsieanztos(mFaig). 4): ovipositorshort, oval, fleshy 24 Tipofgonostyluswithverycomplexenlarge- - 5twW)oii;nntoiggpv,impwacoirustgrihivtneoArd(,sFdhiovgo.rewtin3n)tw;oastlrhioodbrnitga,{l,Tstrcpaiuupcrrehvsroeicpdnergreeasmv)eaenrntkle(yFdilgty.o 3 10. (hWmYeioTnnm,tgdNaownWridiTtghc)ionanosdfpiisRtciSunocautnsddbarraukdsahsrpoikofitrcsslienaotmuaCadeejlo(olFrviegRAr.lerb1-x.e5m-a)nder 3. Ginognso;stgyolnoucsoxwiittehsofnreeqourenmtolryeenlloabregsedo;rbsrwiedlgle- - Wcrionsgsvewiinth(oFuigt.a3)dark spot in Cell R. entirely II connectinggonocoxitescomplete, often with clearorwithadarkcloudoverr-mcrossvein protuberanceinmiddle(Figs.9-\>)d'rului- 12 cera{Meuirrichocerti)) 4 11. Gonostylus simple; 9th sternite deeply con- Gonostylussimple,orwithbasal swelling or caveinmiddleofposteriormarginandwith- protuberance: gonocoxites not markedly en- outsetaeinmiddle(Fig. 17)(easternU.S.and larged; bridge connecting gonocoxites com- Canada) hinniciilaWalker plete or incomplete, without protuberance in - Gonostylus with small pointed tubercle; 9th middle (Figs. 16-30) iTrichaccra {Tricho- sternite with two low bulges near middle of cera)) 10 posterior margin and setae along posterior 4. Distal portion ofgonostylus cylindrical, not margin (Fig. 18) (BC toQC) enlarged or strongly expanded (Figs. 9-12) /ntiriilif)cniii.\Meigen 5 12. Abdomen distinctly annulate, Icrgites ob- - Distal portion ofgonostylusexpanded, com- scurely yellowish with posterior margins plex,oftensomewhatclub-shaped(Figs. 13- brownish (CA.OR,NFD, NY.CT. MD,VA) 1.5) 8 (Fig. 19) aniHilala Meigen .5. Lobe on basal portion of dististylus short, Abdomen entirely dark, or dark above and aboutas longaswidthofgonostylusatpoint pale beneath with two color meeting essen- ofattachment(Figs.9-10) 6 tiallyinastraightlinelaterally 13 - Lobeonbasalportionofdististyluslong,two 13. Wing with distinct cloud overr-m crossvein ormoretimesaslongaswidthofgonostylus (eastern U.S. and Canada) (Fig. 20) .... atpointofattachment(Figs. I1-12) 7 rciiclalioiiis(L.) 6. Lobeongonostyluswithblunttip.distalpart - Wing clear sometimes with slight cloud in ofgonostylus beyond lobe with mesal face stigmalare. i.e..Cell R, 14 den.selysetwithshondarksetae;bridgecon- 14. Scape of antenna and thorax pale orange necting gonocoxites without pointed protu- brown; abdomen with dorsum dark, \enter beranceinmiddle(AK.BC,CAacrossnorth- pale, the two colors meeting laterally essen- ernU.S)(Fig.9) fiorreniAlexander tially inastraightline(CA,WA, BC) .... - Lobe on gonostylus with pointed lip; distal anuiiitli. n. sp. PROCEEDINGSOFTHE ENTOMOLOGICALSOCIETYOFWASHINGTON I Poroclodura tnchoptcro 6 Paracloduro tricho^ttra 2 Oioxotma tubtinuatum 7 Oiozotmo tubsinuotum A| CiiAg' 3 Trichoctro bimocMlo ^ 4 Poroclodura trlchoptcro 8 Triehocoro fottigioro TARSAL SEGMENT I V' ^ 1 9 Trichoeoro bimoculo Figs. l-.S. L Wing.I'unicUuhiniirichoplem. San Franci.sco,CA.2,Wing.Diiizosniasiibsiniiaimn. Halifax Gorge.VT3.Wing, Trichocerahiinaciila. Atlanta.GA.4.Tarsus.P. liichopleni. SanFrancisco.CA..^.Tarsus. T. biinaciila, Atlanta.GA. 6. Malegenitalia. P. Iriclioplem, San Francisco.CA. 7. Malegenitalia.D. siihsiini- atum, HalifaxGorge,VT.8.Malegenitalia,T.fauigiona, Atlanta.GA. VOLUME 105, NUMBER4 905 Figs.9-16. Malegenitaliadetails.9,Trichoceragarretti,Atlanta,GA. 10,T.tcmnensii.WatcrstonNational Park. AB.Canada. II, 7". saliiiaiu. Amherst, MA. 12, T. mackenzie. Masseure.Sweden. 13, 7'. Iideci. Moravia, l-'i. 7". ursiiiiuijin: redrawnfrom Dalil (1967). IS. 7". hicinalis. Atlanta,GA. PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON Scape ofantenna, thorax and abdomen dark scoopedorconcave, with4to8 setaealong L^i posteriormargin betweenpostero-Iateralcor- Malewith completebridgejoininggonocox- ners(Fig. 26)(Canada,AK) excilisDahl ites(Figs. 16,21-23) 16 9th sternite with posterior margin slightly Male with incomplete bridge joining gono- concave or straight usually with 10 or more coxites(Figs.24-27 19 setaebetweenpostero-Iateralcorners .... 25 GonostyUissimple (Fig. 21):wingwith long Setae along posterior margin of9th sternite trichia. those on distal portionofwing veins almostallofsamesize(easternU.S.andCan- about as longasbasal portionofR, ada) re^ehilionis(L.) setosivenaAlexander Setaealongposteriormarginof9thsterniteof Gonoslyliis with basal protuberanceortuber- more than one size (Fig. 27) (AK, northern cle (Figs. 16. 22-2.^) wingwithouttrichiaon Canada) borealisLackschewitz wingveins 17 Male9thsternitewithposteriormargindeep- Malewith9thsterniteusuallywithanobtuse ly concave, without setae in middle portion pointinmiddleofposteriormargin(Fig. 16); (Fig. 17) (eastern U.S. and Canada) .... first flagellarsegmentofantennaelongate. 3 hiinaciilaWalker to4 times as long as basal width Male 9th sternite shallowly concave or luemalis(DeGeer) straightalongposteriormargin,withsetaeall Male9th steniite willioutapoint in middleof along posterior margin between postero-Iat- posterior margin; first flagellarsegmentofan- eralcorners(Figs.20,29-30) 27 tennaaboutaslongasfollowingsegments . . IS Male9thsternitewith 10ormoresetaealong Male9thsternitewithfew setaenearmiddle posteriormarginbetweenpostero-Iateralcor- of posterior margin (AK, northern Canada) ners (easternU.S. andCanada) (Fig.22) arclicaLundstriim regelationis(L.) Male9thsternitewithmanysetaenearmiddle Male9thsternitewith6to8setaealongpos- ofposteriormargin (CA,OR)(Fig. 23).... teriormarginbetween postero-Iateralcorners pallensAlexander 28 Halves of incomplete bridge slender, arising Aedeaguswithshortbasalapodemeandonly froinmid-lengthofgonocoxites(Figs.34-35); slight trace oflongitudinal groove (Fig. 29) parameresaboutaslongasaedeagus 20 (eastern LI.S. andCanada) Halvesofincomplete bridge broader, arising brevicoinisAlexander near base ofgonocoxites (Figs. 24-27, 29- Aedeaguswithlongerbasalapodemeandlon- 30); paramereslongerthanaedeagus 21 gitudinal groove(Fig. 30)(western U.S.and Gonostylus stout, tip rounded(Fig. 34); par- Canada) columhianaAlexander Gaomneor.esstywliutshslbernodaedr,ttiipp(pIoDi)nte.d.(Fiidga.ho3e5n)s:isp.ar-n. sp. bMaaslael lgoobneoscofxoirtmeisngvedirsytinscttoutb,ridignef;lagteodnoswtiyt-h eGaNromHnet)orh.eassntyowlniuteshwatpiotbihans2taeltdutbtheiirprcsdle(h(sFiA,itBguo).bne2er.4c)a.utl.(abAtahKasa,enAffMlliaeA.rxg.a-n.ndsepr. ct(lfieFuezsirmegcasd.ullsied;3gu1hcwst)tilp,yte(hsrCbumrOtbao,eetarqhgdMuieetaEcnel,aedt1ooN0vdHoisssi,ptudeab,rlVelmTqywa,,uitathilQhtesCctila)inolpnlgrdeoin.uagsnmctdelheetrdeto;ro- Gonostylus with one ba.sal tubercle, a setig- Diazosmasuhsinuatmn(Alexander) erous protuberance, or simple (Figs. 25-27, Male gonocoxites comparatively slender, 29-30) 22 without apparent bridge; gonostylus slender aGomneorsetsy.lsutsongwliythanbgalseadl,twuibtehrcslpeur(Fbieg.yo2n5)d,mpiadr-- bmeafloerewiatphexterignittoem1o0rleonogrerletshsanobcteursceust;ips:pefre-- dleofaedeagus(Fig. 8);bothsexeswithlast matheca spherical with sclerotized duct less s(eCgTmesnotutohftpoalGpAu,s wnoetstcotnostMrSic)ted in middle t(hOaRn.hUaTl)f spermathecal diaDmieatezro)sm(aFipgr.att3i6)Stary fattigianaAlexander Gonostylus with basal setigerous protuber- anceorswelling,orsimple(Figs.26—27,29- Species 30): parameres gently curved, somewhat scimitar-shaped; last segment ofpalpus con- Diazosina Pratti Stary strictedinmiddle 23 (Fig, 36) Gonostylus with basal setigerous protuber- anceorswelling(Figs.26-27) 24 Gonostylussimple(Figs.29-30) 26 Diazosmapratti Stary 1995: 210-213. Iig.s. 9th sternite with posterior margin deeply 5-8. Holotype 6. Cowly Canyon. Cache VOLUME 105. NUMBER4 W-W gonostylut IJ ] cr 9th sTernite 17 T bimocula 21 T. sotosivena gonocoxit* nj^^j^llUi^ incomplet bridge-r ^ 18 T. maculipennis rriu_iu 19 T. onnulato 23 T. pollens 20 T. rtgelotionfs 24 T. bituberculoto Blay.nFtifi/i.,s^aAiBIm7.u-2lC4aa.wn.adMaAr.alli2en3g.gteoTnn.i.tfaaVlliAlae.nd,e2.t0a.iSlasTn..ta1r7e.CgreuTl/ra,iticCohono.ic.sc.rCaAIt.hhma2cu4au.,nlTN.aY..h,An2t,1lh.acnrTtca„.sUc„GiaoA.s.ivAcImmNh,.e.r/:sBtmo'ayceMurlA,OpcRnn,2s2BeT.ararIscllaunad PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON gonottylut gonocoxit* 9,h ,,,,„.^. 25 T. fottlgiona 26 T txcriit 31 D-subtinuotum spcrmathtcov 28 R trichoptvr* 32 P. trichopttro Fig.s. 25-32. 25-31, Malegenitaliadetails. 25. Trichocenifatuaiaiia. Atlanta. GA. 26. T. excilis, redrawn fromStaryandMartinov.sky(1996),Dawson,YT,Canada.27,T.bnrealis.Akiavik.NT.Canada.28.Pantcladura nichoplcm. SanFrancisco,CA.29.T.brevicornis. Atlanta.GA.30,T.coliiinhiana. Pullman.WA.31.Diazosina siihsimuiiiiiii. HalifaxGorge,VT. 32.Femaleabdomen. P. iriclio;>icra. MarinCo..CA. Co., UT in collection J. Stary. Faculty of Diazosma siib.siniiatiim Alexander Science. Olomouc. Czech Reputblic. (Figs. 2. 7. 31) Geographic range, -UT and OR (Stary Tr'uliovcni [Diazosma) .siibsitniatuu) Alex- 1995). ander 1916: 124-125, tig. 10. Hoiotype VOLUME 105, NUMBER4 33 T. ornoudi tr) 36 D. prottJ Figs. 33-36. Dctiiilsofmale genitalis. 33. Trulmcenicinuiiuli. Inverness. CA. 34. 7". iilcilimnsis, Moscow Ml.. ID. 35. T. iKiiifti. Banff, AB.Canada. Mx DuKoMuapr,iiri. CacheCo.. LIT. PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON 6. Hall Valley. Platte Canyon, CO. Car- Trichocera (Metatrichocera) inacl<enzie cass and slide in USNM. (Dahl) Geographic range.—BC to CA across (Fig. 12) southern Canada and northern U.S. east to Metatrichocera maclienzie Dahl 1967: 60— NY and VT (Alexander 1967, Pratt and 63, figs. 11-15, 56-60. Holotype 6, Pratt 1984). Mackenzie Delta, NWT.Canada inCNC. — Geographic range. AK, northern Cana- Piircukuliira trichoptera (Osten Sacken) da, northern Sweden. (Dahl and Alexander (Figs. 1, 6, 28, 32) 1976). Trichocera trichoptera Osten Sacken 1877: 204. Holotype 9,LagunitasCreek,Marin Tricliocera (Metatrichocera) sahnani Co., CA in MCZ Alexander (Fig. 11) Geographic range.—BC, WA. OR, CA Tricliocera sahnani Alexander 1927: 72. (Alexander 1967) Holotype 6,Amherst,MA,slideonly,no Trichocera (Metatrichocera) colei carcass in USNM. Alexander Geographic range.—MA, MD, KS, NH (Fig. 13) (Pratt and Pratt 1984). Trichocera coleiAlexander 1919: 162. Ho- lsloitdyepeinSU.SFNorMe.st Grove, OR, carcass and Trichocera (MeAtlaetxraincdhoecrera) tetonensis (Fig. 10) Geographic range.—AK, BC, WA, OR, CA (Alexander 1967). Trificghs.oce1,ra2.teHtoolnoetnyspiseA6l.exHaindddeern1F9a4l5l:s,3s9t8n,. Triclioccra (Metatrichocera) garretti 3 WY in USNM, carcass only, no slide. Alexander Trichocera hyaloptera Alexander 1949: (Fig. 9) 274, fig. 3. Holotype c?, Peavine Ridge, Yamhill Co.. OR. carcass only, no slide Trichocera garretti Alexander 1927: 71- in USNM. Synonymy by Dahl and Al- 7sl2i.deHoolnolty,ypneoSc.arMcaarsyssviinllUe.SNBCM,Canada, exander (1976). — Trichocera alexanderi Dahl 1967:59, figs. Geographic range. AK, OR. northern 7-10. Holotype <5, Mt. Robson,BC,Can- Canada, northernU.S. (DahlandAlexander ada in CNC. Synonymy by Dahl and Al- 1976). exander (1976). Trichocera Metatrichocera nrsainajor Geographic range.—AK to CA east ( Alexander ) across southern Canada and northern U.S. (Fig. 15) to NH and GA (Pratt and Pratt 1984). Trichocera iirsamajorAlexander 1959: 58. Holotype d. Bear Lake, NWT, Canada, Trichocera (Metatrichocera) liitea Becher slide only in USNM (Fig. 14) — Trichocera liitea Becher 1886: 64. Holo- Geographic range. AK, northern Cana- type (J, Jan Mayen in NMW. da (Dahl and Alexander 1976). Geographic range.—Jan Mayen, Spitz- Trichocera {Trichocera) aninilata Meigen bergen. Bear Island, Greenland, Iceland, (Fig. 19) Fennoscandia, northwestern Russia, Alps TrichoceraammlataMeigen 1818:215. Ho- (Dahl and Alexander 1976). lotype d, Austria in MNHN.

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