The winter crane flies of North America north of Mexico (Diptera: Trichoceridae)

14 Pages·2003·3.7 MB·

by  Harry D Pratt

PROC. ENTOMOL.SOC.WASH.
105(4).2003.pp.901-914
THE WINTER CRANE FLIES OF NORTH AMERICA NORTH OF MEXICO
(DIPTERA: TRICHOCERIDAE)
H.^RRY D. Pratt
104 South Almond. Simpsonville. SC 29681, U.S.A. (e-mail: [email protected])
—
Abstract. Twenty-eight .species of winter crane flies in the family Trichoceridae are
found in North America north of Mexico; Diazosmapratti Stary. D. siihsinuatum Alex-
ander, Paracladura trichoptera (Osten Sacken), Trichocera (Metatrichoceni) coleiAlex-
ander, T. (M.) garretti Alenander. T. (M.) liiteci Becher. T. (A/.) mackenzie (Dahl), T. (M.)
salmaniAlexander, T. (M.)tetoncnsisAlexander, T. (M.)ursamajorMc\i\ndci\Trichoceni
(Trichocera) annulata Meigen, T. (T.) arctica Lundstrom. T. (T.) aniaudi Pratt, n. sp.,
T. (T.) haiifft Pratt, n. sp., T. (T.) bimacula Walker, T. (T.) bituherciilata Alexander, T.
(7.) borealis Lackschewit/, T. (T.) brevicornis Alexander, T. (T.) coliimbiana Alexander,
T. (T.) excilis Dahl, T. (T.) faitigiaiia Alexander, T. (T.) gracilis Walker, T. (T.) hieimilis
(De Geer), T. (T.) idahoensis Pratt, n. sp.. T. (T.) maculipenuis Meigen, T. (T.)pollens
Alexander, T. (T.) regelationis (Linnaeus), and T. (T.) setosivena Alexander. Trichocera
loiigisetosa Alexander is a new synonym of Trichocera {Trichocera) setosivena Alex-
ander. Keys to genera and species, notes on distributions, and illustrations are provided.
Kcx Words: Diptera, Trichoceridae. winter crane flies. North America, taxonomy
Winter crane flies of the genus Tricho- collected by net. light trap. Malaise trap,
cera Meigen are found on warm sunny af- and molasses trap.
ternoons in fall, winter, and spring in the The following acronyms indicate the lo-
contiguous United States, Canada, and cation oftypes:
Alaska. Ontheotherhand,adultsofthetwo ANSP Academy of Natural Sciences,
species ofDiazosma Bergroth are collected Philadelphia, PA.. USA.
from June to September, the only trichocer- BMNH The Natural History Museum,
idsonwinginthesummerinnorthern Unit- London, UK.
ed States. Adults of Paracladura trichop- CAS California Academy of Sciences,
tera (Osten Sacken) are found from August San Francisco, CA, USA.
through the winter to spring in British Co- CNC Canadian National Collection,
lumbia. Washington, Oregon, and Califor- Ottawa. ON, Canada.
nia. Swarms of Trichocera males are often LSI Linnean Society ofLondiMi. Lon-
seen dancing in the late afternoon sunlight, don, UK.
sometimes thousandsofindividuals in hun- MCZ Museum of Comparative Zoolo-
dreds ofswarms over many acres oflawns gy, Harvard University, Cam-
and open woodlands. Biologists also have bridge, MA, USA.
noticed these coldhardy insects swarming MNHN Museum National d'Histoire Na-
above the snow, or specimens crawling on turelle, Paris, France.
the snow when temperatures were between NMW NaturhistorischesMuseum,Wien,
0° and 10°C. Specimens in this study were Austria.
902 PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
LLSNM National MuseumofNatural His- because their types are lost. They are not
tory, Smithsonian Institution, treated further here. Trichocera gracilis
Washington. DC, USA. Walker (1848) was described from a single
ZIL Zoological Institute, Lund, Swe- female from York Factory, Manitoba, Can-
den. ada. Byers (1976) published notes and fig-
ZISP Zoological Institute, Academy of ures ofthe terminal abdominal segmentsof
.Sciences, St. Petersburg, Russia. thetype. I knowofnocharacterstoseparate
this species from females ofother species.
Taxonomy'
Alexandei' published twokeys totheTri- Family Characteristics of the
choceridae found in the eastern United Trichoceridae
States (1919a, 1942) and one for the west- Winter crane flies in the family Tricho-
ern United States (1967). He wrote a fine ceridaearesinallto medium sized (wingup
summary ofthe family Trichoceridae in the to 12 miTi in Diazosma) with slender long
"Manual of Nearctic Diptera" in 1981. legs similar to the true crane flies in the
Dahl published many papers on winter family Tipulidae. Trichoceridae differ in
crane flies. Dahl's 1966 publication on the having 3 ocelli, the V-shaped suture on the
Swedish Trichoceridae includes much in- mesonotum incomplete in the middle, and
formationonseveral speciesfound inNorth long hairlike antennae. Tipulidae lackocel-
America, and her 1967 paperon arctic and li, have a complete V-shaped suture on the
subarcticTrichoceridaeisthebasicworkon mesonotum separating the praescutum and
northern species. The world catalogue of scutum, and antennae usually short. True
Trichoceridae by Dahl and Alexander cranefliesareoftenmuchlargerthanwinter
(1976) lists all the species known to 1976, crane flies, with wings up to 25 mm or
references to the original descriptions, and more. —
known locations of types of most species. Description. Head with 3 ocelli. Anten-
Pratt and Pratt (1984) published keys, illus- na with 16 ""segments," scape and pedicel
trations and data on ten species of eastern enlarged, flagellomeeres 3-16 elongate and
North America. Pratt (1992) published a progressively more slender. Mouthparts re-
key to the Trichoceridae ofNorth America. duced, mandibles absent, galea of maxilla
Stary (1995) described Diazosma pratti well-preserved, palpus with 4 free seg-
and wrote that Diozosnui hirtipeivie (Sieb- ments, no trace ofsensory pore (as in 5v/-
ke) is a European species and that North vicola Harris and Culicoides Latreille).
American records of this insect should be Thorax with V-shaped suture separating
called D. sitbsiiniatiiin (Alexander). Stary praescutun and scutum incomplete in mid-
and Martinovsky (1996) wrote that Tricho- dle, and no distict suture separatingscutum
cerajaponicu Matsumura (1916) probably and scutellum. Wing with Sc long joining
is not a North American species and that C at midlength, 4 branches of R and 3
M
records of this species by Dahl and Alex- branches of reaching wing margin. A,
ander (1976) and Pratt (1992) should be long. A, short and curved to wing inargin
called Trichocera excilis Dahl (1967). in Trichocera and Paracladitra Brunetti
Krzeminska (2001) wrote that Trichocera longer and subsinuous in Diazosma. Legs
major Edwards is a European species and not markedly deciduous as in Tipulidae:
that American records of this species Abdomen with 8 distinct pregenital seg-
should be called Trichocera setosivena Al- ments, first segment short. Male genitalia
exander. with medio-basal projections of gonocox-
Trichocera scutellata Say (1824) from itesformingincompleteorcompletebridge,
Minnesota and Trichocera hrumalis Fitch gonostylus single, phallosome with aedea-
(1847) from New York are unrecosinized gus or sperm pump somewhat pearshaped
VOLUME 105. NUMBER4 903
and parameres joined medially by thin portionofgonostylusbyondlobewithsparse
membrane. Female with ovipositor formed long setae; bridge connecting gonocoxites
by sclerotized, elongate, downward curved withpointedprotuberanceinmiddle(Fig. 10)
(AK.OR,northwesternU.S.)(Fig. 10) ....
cerci in Trichocera and Panuhuliira or tetonensisAlexander
short and fleshy in Diazosnia. 7. Lobeofgonostylusatba.sal fourthofgonos-
Key to the Winter Crane Flies btyrluussh;loafterlaolngl,obreesddoifshninhtahirster9glihteswteirtnhitdeenHsael
(Trichoceridae) of North America (Fig. II)(NH.MA.MDwestto KS)
North of Mexico salmaniAlexander
- Lobeofgonostylusarisingatbaseofgonos-
tylus; later lobes ofninth tergite with a few
I Tarsiwithbasitarsomereveryshort,aboutone- sparse hairs; ninth sternite balloon-like with
eighth as long as second tar.somere (Fig. 4): setosedorsal lobe (Fig. 12) (AK. northwest-
wing with m-cucrossveinpresent(Fig. I);fe- ernCanada) niackcnzieDahl
cmkaulheiiaw.itohneonsepecsipeesi.maBtCh.ecWaA.(FiOg.R,32C)A)(Pc.im.-. 8. lBaarsgaeliarrnedgudliasrtarloupnodrteidonlsoboef(Fgiogn.os1t4y)lu(sGreweint-h
- Tarsi withPabrausciltaadristormaerIehclhoonpgteerrath(aOnstseencoSnadcken) Blaansde)ofgonostylus without large rouInudrealoBbeecher
(Fig. 5); wing with m-cu crossvein absent 9
2. e(WFviiengnsgl.y2w-it3ot)h;wiAfn,egmlaomlnaegr,gwiistnuhb(s3Fiinsgu.poeu2rs)m,;attnhiobeticalaceusrpv.ue.rd.s 2 9. T(Fiipg.ofIIg)on(oAsKt.yluBsC,wiWtAh,laOrRge, rCoAcu)onldeeidAlleoxbeander
(abDsieanztos(mFaig). 4): ovipositorshort, oval, fleshy 24 Tipofgonostyluswithverycomplexenlarge-
- 5twW)oii;nntoiggpv,impwacoirustgrihivtneoArd(,sFdhiovgo.rewtin3n)tw;oastlrhioodbrnitga,{l,Tstrcpaiuupcrrehvsroeicpdnergreeasmv)eaenrntkle(yFdilgty.o 3 10. (hWmYeioTnnm,tgdNaownWridiTtghc)ionanosdfpiisRtciSunocautnsddbarraukdsahsrpoikofitrcsslienaotmuaCadeejlo(olFrviegRAr.lerb1-x.e5m-a)nder
3. Ginognso;stgyolnoucsoxwiittehsofnreeqourenmtolryeenlloabregsedo;rbsrwiedlgle- - Wcrionsgsvewiinth(oFuigt.a3)dark spot in Cell R. entirely II
connectinggonocoxitescomplete, often with clearorwithadarkcloudoverr-mcrossvein
protuberanceinmiddle(Figs.9-\>)d'rului- 12
cera{Meuirrichocerti)) 4 11. Gonostylus simple; 9th sternite deeply con-
Gonostylussimple,orwithbasal swelling or caveinmiddleofposteriormarginandwith-
protuberance: gonocoxites not markedly en- outsetaeinmiddle(Fig. 17)(easternU.S.and
larged; bridge connecting gonocoxites com- Canada) hinniciilaWalker
plete or incomplete, without protuberance in - Gonostylus with small pointed tubercle; 9th
middle (Figs. 16-30) iTrichaccra {Tricho- sternite with two low bulges near middle of
cera)) 10 posterior margin and setae along posterior
4. Distal portion ofgonostylus cylindrical, not margin (Fig. 18) (BC toQC)
enlarged or strongly expanded (Figs. 9-12) /ntiriilif)cniii.\Meigen
5 12. Abdomen distinctly annulate, Icrgites ob-
- Distal portion ofgonostylusexpanded, com- scurely yellowish with posterior margins
plex,oftensomewhatclub-shaped(Figs. 13- brownish (CA.OR,NFD, NY.CT. MD,VA)
1.5) 8 (Fig. 19) aniHilala Meigen
.5. Lobe on basal portion of dististylus short, Abdomen entirely dark, or dark above and
aboutas longaswidthofgonostylusatpoint pale beneath with two color meeting essen-
ofattachment(Figs.9-10) 6 tiallyinastraightlinelaterally 13
- Lobeonbasalportionofdististyluslong,two 13. Wing with distinct cloud overr-m crossvein
ormoretimesaslongaswidthofgonostylus (eastern U.S. and Canada) (Fig. 20) ....
atpointofattachment(Figs. I1-12) 7 rciiclalioiiis(L.)
6. Lobeongonostyluswithblunttip.distalpart - Wing clear sometimes with slight cloud in
ofgonostylus beyond lobe with mesal face stigmalare. i.e..Cell R, 14
den.selysetwithshondarksetae;bridgecon- 14. Scape of antenna and thorax pale orange
necting gonocoxites without pointed protu- brown; abdomen with dorsum dark, \enter
beranceinmiddle(AK.BC,CAacrossnorth- pale, the two colors meeting laterally essen-
ernU.S)(Fig.9) fiorreniAlexander tially inastraightline(CA,WA, BC) ....
- Lobe on gonostylus with pointed lip; distal anuiiitli. n. sp.
PROCEEDINGSOFTHE ENTOMOLOGICALSOCIETYOFWASHINGTON
I Poroclodura tnchoptcro
6 Paracloduro tricho^ttra
2 Oioxotma tubtinuatum
7 Oiozotmo tubsinuotum
A| CiiAg'
3 Trichoctro bimocMlo
^
4 Poroclodura trlchoptcro
8 Triehocoro fottigioro
TARSAL SEGMENT
I
V' ^ 1
9 Trichoeoro bimoculo
Figs. l-.S. L Wing.I'unicUuhiniirichoplem. San Franci.sco,CA.2,Wing.Diiizosniasiibsiniiaimn. Halifax
Gorge.VT3.Wing, Trichocerahiinaciila. Atlanta.GA.4.Tarsus.P. liichopleni. SanFrancisco.CA..^.Tarsus.
T. biinaciila, Atlanta.GA. 6. Malegenitalia. P. Iriclioplem, San Francisco.CA. 7. Malegenitalia.D. siihsiini-
atum, HalifaxGorge,VT.8.Malegenitalia,T.fauigiona, Atlanta.GA.
VOLUME 105, NUMBER4 905
Figs.9-16. Malegenitaliadetails.9,Trichoceragarretti,Atlanta,GA. 10,T.tcmnensii.WatcrstonNational
Park. AB.Canada. II, 7". saliiiaiu. Amherst, MA. 12, T. mackenzie. Masseure.Sweden. 13, 7'. Iideci. Moravia,
l-'i. 7". ursiiiiuijin: redrawnfrom Dalil (1967). IS. 7". hicinalis. Atlanta,GA.
PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
Scape ofantenna, thorax and abdomen dark scoopedorconcave, with4to8 setaealong
L^i posteriormargin betweenpostero-Iateralcor-
Malewith completebridgejoininggonocox- ners(Fig. 26)(Canada,AK) excilisDahl
ites(Figs. 16,21-23) 16 9th sternite with posterior margin slightly
Male with incomplete bridge joining gono- concave or straight usually with 10 or more
coxites(Figs.24-27 19 setaebetweenpostero-Iateralcorners .... 25
GonostyUissimple (Fig. 21):wingwith long Setae along posterior margin of9th sternite
trichia. those on distal portionofwing veins almostallofsamesize(easternU.S.andCan-
about as longasbasal portionofR, ada) re^ehilionis(L.)
setosivenaAlexander Setaealongposteriormarginof9thsterniteof
Gonoslyliis with basal protuberanceortuber- more than one size (Fig. 27) (AK, northern
cle (Figs. 16. 22-2.^) wingwithouttrichiaon Canada) borealisLackschewitz
wingveins 17 Male9thsternitewithposteriormargindeep-
Malewith9thsterniteusuallywithanobtuse ly concave, without setae in middle portion
pointinmiddleofposteriormargin(Fig. 16); (Fig. 17) (eastern U.S. and Canada) ....
first flagellarsegmentofantennaelongate. 3 hiinaciilaWalker
to4 times as long as basal width Male 9th sternite shallowly concave or
luemalis(DeGeer) straightalongposteriormargin,withsetaeall
Male9th steniite willioutapoint in middleof along posterior margin between postero-Iat-
posterior margin; first flagellarsegmentofan- eralcorners(Figs.20,29-30) 27
tennaaboutaslongasfollowingsegments . . IS Male9thsternitewith 10ormoresetaealong
Male9thsternitewithfew setaenearmiddle posteriormarginbetweenpostero-Iateralcor-
of posterior margin (AK, northern Canada) ners (easternU.S. andCanada)
(Fig.22) arclicaLundstriim regelationis(L.)
Male9thsternitewithmanysetaenearmiddle Male9thsternitewith6to8setaealongpos-
ofposteriormargin (CA,OR)(Fig. 23).... teriormarginbetween postero-Iateralcorners
pallensAlexander 28
Halves of incomplete bridge slender, arising Aedeaguswithshortbasalapodemeandonly
froinmid-lengthofgonocoxites(Figs.34-35); slight trace oflongitudinal groove (Fig. 29)
parameresaboutaslongasaedeagus 20 (eastern LI.S. andCanada)
Halvesofincomplete bridge broader, arising brevicoinisAlexander
near base ofgonocoxites (Figs. 24-27, 29- Aedeaguswithlongerbasalapodemeandlon-
30); paramereslongerthanaedeagus 21 gitudinal groove(Fig. 30)(western U.S.and
Gonostylus stout, tip rounded(Fig. 34); par- Canada) columhianaAlexander
Gaomneor.esstywliutshslbernodaedr,ttiipp(pIoDi)nte.d.(Fiidga.ho3e5n)s:isp.ar-n. sp. bMaaslael lgoobneoscofxoirtmeisngvedirsytinscttoutb,ridignef;lagteodnoswtiyt-h
eGaNromHnet)orh.eassntyowlniuteshwatpiotbihans2taeltdutbtheiirprcsdle(h(sFiA,itBguo).bne2er.4c)a.utl.(abAtahKasa,enAffMlliaeA.rxg.a-n.ndsepr. ct(lfieFuezsirmegcasd.ullsied;3gu1hcwst)tilp,yte(hsrCbumrOtbao,eetarqhgdMuieetaEcnel,aedt1ooN0vdHoisssi,ptudeab,rlVelmTqywa,,uitathilQhtesCctila)inolpnlgrdeoin.uagsnmctdelheetrdeto;ro-
Gonostylus with one ba.sal tubercle, a setig- Diazosmasuhsinuatmn(Alexander)
erous protuberance, or simple (Figs. 25-27, Male gonocoxites comparatively slender,
29-30) 22
without apparent bridge; gonostylus slender
aGomneorsetsy.lsutsongwliythanbgalseadl,twuibtehrcslpeur(Fbieg.yo2n5)d,mpiadr-- bmeafloerewiatphexterignittoem1o0rleonogrerletshsanobcteursceust;ips:pefre--
dleofaedeagus(Fig. 8);bothsexeswithlast matheca spherical with sclerotized duct less
s(eCgTmesnotutohftpoalGpAu,s wnoetstcotnostMrSic)ted in middle t(hOaRn.hUaTl)f spermathecal diaDmieatezro)sm(aFipgr.att3i6)Stary
fattigianaAlexander
Gonostylus with basal setigerous protuber-
anceorswelling,orsimple(Figs.26—27,29- Species
30): parameres gently curved, somewhat
scimitar-shaped; last segment ofpalpus con- Diazosina Pratti Stary
strictedinmiddle 23 (Fig, 36)
Gonostylus with basal setigerous protuber-
anceorswelling(Figs.26-27) 24
Gonostylussimple(Figs.29-30) 26 Diazosmapratti Stary 1995: 210-213. Iig.s.
9th sternite with posterior margin deeply 5-8. Holotype 6. Cowly Canyon. Cache
VOLUME 105. NUMBER4
W-W
gonostylut
IJ ]
cr 9th sTernite
17 T bimocula 21 T. sotosivena
gonocoxit*
nj^^j^llUi^
incomplet
bridge-r
^ 18 T. maculipennis
rriu_iu
19 T. onnulato 23 T. pollens
20 T. rtgelotionfs 24 T. bituberculoto
Blay.nFtifi/i.,s^aAiBIm7.u-2lC4aa.wn.adMaAr.alli2en3g.gteoTnn.i.tfaaVlliAlae.nd,e2.t0a.iSlasTn..ta1r7e.CgreuTl/ra,iticCohono.ic.sc.rCaAIt.hhma2cu4au.,nlTN.aY..h,An2t,1lh.acnrTtca„.sUc„GiaoA.s.ivAcImmNh,.e.r/:sBtmo'ayceMurlA,OpcRnn,2s2BeT.ararIscllaunad
PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
gonottylut
gonocoxit* 9,h ,,,,„.^.
25 T. fottlgiona
26 T txcriit
31 D-subtinuotum
spcrmathtcov
28 R trichoptvr*
32 P. trichopttro
Fig.s. 25-32. 25-31, Malegenitaliadetails. 25. Trichocenifatuaiaiia. Atlanta. GA. 26. T. excilis, redrawn
fromStaryandMartinov.sky(1996),Dawson,YT,Canada.27,T.bnrealis.Akiavik.NT.Canada.28.Pantcladura
nichoplcm. SanFrancisco,CA.29.T.brevicornis. Atlanta.GA.30,T.coliiinhiana. Pullman.WA.31.Diazosina
siihsimuiiiiiii. HalifaxGorge,VT. 32.Femaleabdomen. P. iriclio;>icra. MarinCo..CA.
Co., UT in collection J. Stary. Faculty of Diazosma siib.siniiatiim Alexander
Science. Olomouc. Czech Reputblic. (Figs. 2. 7. 31)
Geographic range, -UT and OR (Stary Tr'uliovcni [Diazosma) .siibsitniatuu) Alex-
1995). ander 1916: 124-125, tig. 10. Hoiotype
VOLUME 105, NUMBER4
33 T. ornoudi
tr)
36 D. prottJ
Figs. 33-36. Dctiiilsofmale genitalis. 33. Trulmcenicinuiiuli. Inverness. CA. 34. 7". iilcilimnsis, Moscow
Ml.. ID. 35. T. iKiiifti. Banff, AB.Canada. Mx DuKoMuapr,iiri. CacheCo.. LIT.
PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
6. Hall Valley. Platte Canyon, CO. Car- Trichocera (Metatrichocera) inacl<enzie
cass and slide in USNM. (Dahl)
Geographic range.—BC to CA across (Fig. 12)
southern Canada and northern U.S. east to Metatrichocera maclienzie Dahl 1967: 60—
NY and VT (Alexander 1967, Pratt and 63, figs. 11-15, 56-60. Holotype 6,
Pratt 1984). Mackenzie Delta, NWT.Canada inCNC.
—
Geographic range. AK, northern Cana-
Piircukuliira trichoptera (Osten Sacken)
da, northern Sweden. (Dahl and Alexander
(Figs. 1, 6, 28, 32)
1976).
Trichocera trichoptera Osten Sacken 1877:
204. Holotype 9,LagunitasCreek,Marin Tricliocera (Metatrichocera) sahnani
Co., CA in MCZ Alexander
(Fig. 11)
Geographic range.—BC, WA. OR, CA
Tricliocera sahnani Alexander 1927: 72.
(Alexander 1967)
Holotype 6,Amherst,MA,slideonly,no
Trichocera (Metatrichocera) colei carcass in USNM.
Alexander Geographic range.—MA, MD, KS, NH
(Fig. 13) (Pratt and Pratt 1984).
Trichocera coleiAlexander 1919: 162. Ho-
lsloitdyepeinSU.SFNorMe.st Grove, OR, carcass and Trichocera (MeAtlaetxraincdhoecrera) tetonensis
(Fig. 10)
Geographic range.—AK, BC, WA, OR,
CA (Alexander 1967). Trificghs.oce1,ra2.teHtoolnoetnyspiseA6l.exHaindddeern1F9a4l5l:s,3s9t8n,.
Triclioccra (Metatrichocera) garretti 3 WY in USNM, carcass only, no slide.
Alexander Trichocera hyaloptera Alexander 1949:
(Fig. 9) 274, fig. 3. Holotype c?, Peavine Ridge,
Yamhill Co.. OR. carcass only, no slide
Trichocera garretti Alexander 1927: 71- in USNM. Synonymy by Dahl and Al-
7sl2i.deHoolnolty,ypneoSc.arMcaarsyssviinllUe.SNBCM,Canada, exander (1976). —
Trichocera alexanderi Dahl 1967:59, figs. Geographic range. AK, OR. northern
7-10. Holotype <5, Mt. Robson,BC,Can- Canada, northernU.S. (DahlandAlexander
ada in CNC. Synonymy by Dahl and Al- 1976).
exander (1976).
Trichocera Metatrichocera nrsainajor
Geographic range.—AK to CA east ( Alexander )
across southern Canada and northern U.S. (Fig. 15)
to NH and GA (Pratt and Pratt 1984). Trichocera iirsamajorAlexander 1959: 58.
Holotype d. Bear Lake, NWT, Canada,
Trichocera (Metatrichocera) liitea Becher slide only in USNM
(Fig. 14) —
Trichocera liitea Becher 1886: 64. Holo- Geographic range. AK, northern Cana-
type (J, Jan Mayen in NMW. da (Dahl and Alexander 1976).
Geographic range.—Jan Mayen, Spitz- Trichocera {Trichocera) aninilata Meigen
bergen. Bear Island, Greenland, Iceland, (Fig. 19)
Fennoscandia, northwestern Russia, Alps TrichoceraammlataMeigen 1818:215. Ho-
(Dahl and Alexander 1976). lotype d, Austria in MNHN.