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The vascular plants collected by Mark Catesby in South Carolina: Combining the Sloane and Oxford herbaria PDF

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McMillan, P.D. and A.H. Blackwell. 2013. The vascular plants collected by Mark Catesby in South Carolina: Combining the Sloane and Oxford herbaria. Phytoneuron 2013-73: 1-32. Published 27 September 2013. ISSN 2153 733X MARK THE VASCULAR PLANTS COLLECTED BY CATESBY SOUTH CAROLINA: IN COMBINING THE SLOANE AND OXFORD HERBARIA McMillan Patrick D. .1,1 I _| i. nihil. I I. II ' in. I ll,. I ir ii urn. i,l Clem son University Clemson, South Carolina 29634 Amy Hackney Blackwell Department of Biology Furman University Greenville. South Carolina 29613 and South Carolina Botanical Garden m >4 I. i, nil, n In, . i ABSTRACT We Mirk provide a list of all vascular plant specimens collected in the Carolinas by Catesby that are housed in the historic herbaria at Oxford University and the Sloane Herbarium. The identifications along with notes on the significance of selected specimens are presented. This papa- continues our work with Catesby's collections that we began with his specimens in the Sloane Herbarium at the Natural History Museum, London. The availabilit y of high-quality digital images published on the Oxford Herbarium's website has facilitated our examination of these specimens. The collections themselves shed light on the nature of the flora of the Carolinas before European settlement, including the native ranges of several problematic taxa. The presence of a number of taxa known to be introduced to the Americas indicates that these introductions must have occurred prior to the 1720s. KEY WORDS: Catesby, Sloane Herbarium, herbarium, historic botany, ecology, South Carolina, imaging digital Mark Catesby, born in England in 1682 or 1683, devoted most of his adult life to studying the He natural history of southeastern North America and the Caribbean. lived in South Carolina from 1722 to 1725, followed by several months in the Bahamas before he returned home to England. He spent those years traveling in the wilderness, collecting plant specimens, taking copious notes and making sketches. This material became the basis of the publication that made his scholarly reputation, the Natural History of Carolina, Florida, and the Bahama Islands, published between 1729 and 1747. Catesby sent his dried plant specimens to his sponsors in England, including Sir Hans Sloane, William Sherard, and Charles Dubois. Sloane's specimens became part of the original collections of the Natural History Museum in London and are today housed in the Sloane Herbarium, in volumes now H.S. 212 and H.S. 232. Sherard's specimens are in the Sherard and Dubois Herbaria at the University of Oxford. In November 2011, McMillan and Hackney Blackwell traveled to London to digitally We photograph Catesby's Sloane materials. have published these online as part of our Botanica Caroliniana project and published a of determinations of these specimens along with notes on list their ecological, taxonomic, and historical significance (McMillan et al 2013). McMillan and Blackwell: Catesby plants, Sloane and Oxford 2 ; Adding the Oxford collection to the Sloane materials was the logical next step. The Sloane and Oxford materials comprise the full set of Catesby's primary source vascular plant materials from the Carolinas, or nearly so. (James Reveal has mentioned the existence of a box of Catesby specimens on top of a shelf in the Sloane Herbarium; entirely possible that other dried plants it is exist in unknown locations.) Stephen Harris, curator of the Oxford herbaria, has digitally photographed the entire Catesby collection housed at Oxford and published them online. 1 McMillan and Hackney Blackwell have examined of these images and made determinations of almost of all all them. As with our Sloane project, the availability of digital images made quick and easy for us to it examine the specimens at our leisure in South Carolina. The Oxford specimens are well-preserved A and not fragmentary. number of taxa appear in the Oxford materials that were not in the Sloane collections. Combining the lists from the Sloane and Oxford creates a full set of Catesby's work and facilitates even more insights into the ecology of South Carolina before extensive European settlement. Methodology Identification To identify the specimens, McMillan and Hackney Blackwell first did a search on the Oxford BRAHMS database for all Carolina specimens collected by Mark Catesby ("The Collections of Mark Catesby in Oxford," 2013). As we did with the Sloane collections, we set up two laptops side by side, on which we accessed multiple pieces of information: the digital images of Catesby's PDF collections; the of Weakley's Flora of the Carolinas (2012); an online version of Catesby's USDA's Natural History, other websites such as Plants Database; and a database program (Bento we from Filemaker) in which collected and organized our data. The excellent condition of the specimens facilitated the process of identification. Unlike the Sloane specimens, which are bound in books, the Oxford collections are preserved on individual flat sheets of paper. This allowed for considerably flatter images with less distortion than was unavoidable with the Sloane materials. In addition, the Sloane herbaria pages typically contain multiple specimens of different species. The Oxford materials, on the other hand, for the most part contain one specimen per page. This made identifying individual specimens by folio much more straightforward; each specimen associated with either a unique accession number or a barcode is number. Like the Sloane materials, the Oxford specimens contain handwritten texts in addition to We plant specimens- did not take the time to transcribe these notes, many of which are written by hand in Latin. The notes have been photographed individually and are published on the Oxford website along with the plant images. Results In the Oxford collections, we made determinations to the species level for 744 specimens known or suspected to have been collected in Carolina. All determinations were made by Patrick D. McMillan in collaboration with Hackney Blackwell. We are extremely grateful to Stephen Harris for his help with our work on the Sloane specimens and for his hospitality during Hackney Blackwell' s visit to Oxford in May 2012, as well as for his valuable work in photographing and posting online the full Catesby collections from Oxford. J McMillan and Blackwell: Catesby plants, Sloane and Oxford 3 I we we In this table, have combined the Oxford specimens with the determinations already We made and published in Phytoneuron for the specimens from the Sloane Herbarium. did this in the interest of publishing in one place a more complete set of Catesby's collections from the Carolinas. All Oxford specimens are identified by either a barcode number (a string of digits beginning with 000) or a Sherard Herbarium identifying number (beginning with "Sher-"). These were the most obvious unique identifiers available on the online database. The Sloane specimens are identified by volume, either H.S. 212 or H.S. 232, and folio page. Images of the Sloane specimens are available on the Botanica Caroliniana website (Botanica Caroliniana, 2013). Basal Angiosperms W 0M A Ari"stolochiaceae canadense L. 0008743 Aristolochiaceae canadense L. H.S. 212f.58b Aristolochiaceae Endodeca serpentaria (L.) Raf. Sher-2041 Aristolochiaceae Endodeca serpentaria (L.) Raf. H.S. 232 122 f. Calycanthaceae Calycanthus floridus L. H.S. 212 16 f. Lauraceae Litsea aestivalis (L.) Fernald a oof Lauraceae Persea borbonia (L.) Sprengel Sher-0821 6R Persea borbonia (L.) Sprengel 0009 5 70 H Lauraceae Persea borbonia (L.) Sprengel S 232 f 50 Lauraceae Persea palustris (Raf.) Sargent Sher-0821-2 Lauraceae Persea palustris (Ratines que) Sargent H.S. 212 f.l Magnoliaceae Liriodendron tulipifera L. 00087309R Magnoliaceae Liriodendron tulipifera L. H.S. 212f.80 Magnoh Magnoliaceae a acuminata 00087184S (L.) L. Magnoh Magnoliaceae a grandiflora L. Sher- 11 16 Magnoliaceae Magnoha grandiflora L. 00087190P Nymphaeaceae Nymphaea odorata Aiton H.S. 212f.23 Nymphaeaceae Nymphaea odorata Aiton Sher-1089 Nymphaeaceae Nymphaea odorata Aiton 00087301 Nymphaeaceae Nymphaea odorata Aiton 00087305N Nymphaeaceae Nympha ea odorata Aiton 000873060 Nymphaeaceae Nymphaea odorata Aiton Sher-1089-2 Nymphaeaceae Nymphaea odorata Aiton H.S. 232f.84 Saururaceae Saururu, H.S. 232f.82 Monocots Amaryllidaceae Allium cuthbertii Small Sher-0704 Amaryllida ceae Allium cuthbertii Small H.S. 212f.36 Amaryllidaceae H.S. 232f.61 Anacardiaceae Rhus copallinum L. 00087446T A & Arecaceae Sabal palmetto (Walter) Loddiges ex J.H. Sher- 1501 J. Schultes A & Arecaceae Sabal palmetto (Walter) Loddiges ex J.H. Sher-1501-2 J. Schultes, Cannaceae Cannaflaccida Salisbury 00095775X Commelinaceae Commelina erecta L. H.S. 212f.57 Commelinaceae Commelina erecta L. H.S. 212 f.6 Commelinaceae Commelina 00087192R virginica L. McMillan and Blackwell: Catesby plants, Sloane and Oxford 4 I Commelinaceae Cuthbertia rosea (\ enteral) Small SIki -' •' '-1 Cyperaceae carexsp. Cyperaceae 00087555U Cyperaceae 00087571S H yperaceae Carex glaucescens S 212f 44 Elliott H yperaceae ^arex g aucescens .10 S 232 f 139 H yperaceae arex g aucescens S 232 f 139 10 eraceae arex ongH c ™z'e 00087500K >'P 00087567X yperaceae yperaceae yperus ec inatus 00 Sher-0090 .) ( H 212f yperaceae S 86 H yperaceae yperus ec inatus 00 S 232 f 30 .) ( yperaceae Sher-0083 V yperaceae 0008758 IT ' : . ,",Y", Cyperaceae Cyperus erythrorhizos Muhlenberg Sher-0080 Cyperaceae Cyperus polystachyos Rottboll SIki -' " 'X-l yperaceae yperus Sher-0086 sp. Cyperaceae 1. '. 1 . • 1 ' 1 Cyperaceae Cyrilla racemiflora L. 00095682U Cyperaceae Dulichium arundinaceum (L.) Written 00087578Z uw yperaceae lumarun inaceum( on Sher-0069-1 .) ri H S 232 f 61 yperaceae Fuiyenabreviseta (Coville) Coville H.S. 212 44 f. yperaceae uirena revi.seta (ovie)J ovi e 00087574V H uirena squarrosa c S 232 f 139 tuirena squarrosa 000875731 Mfcdsi. -f uirena squarrosa c Sher-1694 . Cyperaceae Rhynchospora caduca kjhott Sher-0099 m yperaceae C SP°ra 0008748 8Z Rh" H H yperaceae y"ch°°sP°ra coIoraa ()L-°>i pPf^iLff* S 212f 45 m ' C SP°ra CO 00087583V ? Rh" h pAff* h° f L ! ' Sher-0074 Cyperaceae Rhynchospora distans (MictoL) Vanl. Sher-0071 H Cyperaceae Rhynchospora Jascicularis (Michx ahl S 232 f 139 ) Cyperaceae Rhynchospora glomerata Vahl 00087492U (L.) Cyperaceae Rhynchospora glomerata vahl 00087559Y (L.) Cyperaceae Rhynchospora glomerata Vahl (L.) Cyperaceae Rhynchospora inundata (Oakes) Fernald 00087582U Cyperaceae Rhynchospora microcephala (Britton) Britton ex Sher-0072 Cyperaceae Ryhncnospora microcephala (Britton) Britton ex 00087575W Cyperaceae Ryhnchospora microcephala (Britton) Britton ex 00087577Y ma" ^ Cyperaceae Scirpus cyperinus (L.) Kunth H.S. 212 1.87 Dioscoreaceae Dioscorea villosa L. Sher-1374-2 Dioscoreaceae Dioscorea villosa L. 00087220J Dioscoreaceae Dioscorea villosa L. 00087224N Dioscoreaceae Dioscorea villosa L. H.S. 212 17 f. Eriocaulaceae Eriocaulon decangulare L. Sher-0196 Eriocaulaceae Eriocaulon decangulare L. 00087204L McMillan and Blackwell: Catesby plants, SI oane and Oxford her 5 Eriocaulaceae Eriocaulon decangulare L. 00087579- 212L41 Eriocaulaceae Eriocaulon decangulare L. H.S. Eriocaulaceae Eriocaulon decangulare L. H.S. 212 42 f. Eriocaulaceae Eriocaulon decangulare L. H.S. 232 133 f. Haemodoraceae Lachnanthes caroliniana (Lam.) Dandy Sher-sn-i Haemodoraceae Luchnanth.es caroliniana (Lam.) Dandy H.S. 232 110 f. Heloniadaceae Chamaelirium luteum (L.) A. Gray Sher-0763 2R Juncaceae Juncus biflorus Elliott 0008746 Juncaceae Juncus biflorus Elliott Sher-0078 Juncaceae Juncus scirpoides Lam. 00087466V Juncaceae Juncus scirpoides Lam. 00087569Z Juncaceae Juncus scirpoides Lam. H.S. 212 43 f. Liliaceae Lilium catesbaei Walter Sher-0708 232L68 Liliaceae Lilium catesbaei Walter H.S. Liliaceae Medeola virginiana L. Sher-0759 Medeola 232L48 Liliaceae virginiana L. H.S. Melanthiaceae Amianthium muscitoxicum (Walter) A. Gray 00087495X Melanthiaceae Amianthium muscitoxicum (Walter) A. Gray Sher-0760 Melanthiaceae Amianthium muscitoxicum (Walter) A. Gray H.S. 212L29 Melanthiaceae Amianthium muscitoxicum (Walter) A. Gray H.S. 212L63 = Melanthiaceae Melanthium hybridum Veratrum species sensu 0008745 IP 1 Weakley = Melanthiaceae Melanthium hybridum Veratrum species sensu 00087482T 1 Weakley Melanthiaceae Melanthium hybridum = Veratrum species sensu H.S. 212L36 1 Weakley = Melanthiaceae Melanthium hybridum Veratrum species sensu H.S. 232 127 1 f. Weakley = Melanthiaceae Melanthium virginicum Veratrum virginicum 00087502M Aiton (L.) = Melanthiaceae Melanthium virginicum Veratrum virginicum 000875 10L Aiton (L.) 00087494W Nartheciaceae Aletris aurea Walter Nartheciaceae Aletris aurea Walter H.S. 232 105 f. BSP Orchidaceae Calopogon iuberosus (L.) Sher-2022 BSP Orchidaceae Calopogon tuberosum (L.) Sher-2022-2 & Orchidaceae Cleistesiopsis divaricata (L.) Pansarin F. Barros 00087515Q A Orchidaceae Habenaria quinqueseta (Michx.) Eaton Sher-2009 Orchidaceae Habenaria repens Nutt. H.S. 212L90 Orchidaceae Habenaria repens Nutt. Sher-2008 Orchidaceae Habenaria repens Nutt, 00087503N Orchidaceae Hexalectris spicata (Walter) Barnhart Sher-2024 Orchidaceae Malaxis unifolia Michx. Sher-2016 212L56 Orchidaceae Platanthera ciliaris (L.) Lindl. H.S. Orchidaceae Platanthera ciliaris (L.) Lindl. 00087499. Orchidaceae Platanthera ciliaris (L.) Lindl. 00087507R Orchidaceae Platanthera integra (Nutt.) A. Gray ex Beck H.S. 212L55 A Orchidaceae Platanthera integra (Nuttall) Gray ex Beck 0008749 IT 00087487Y 11M Orchidaceae Pogonia ophioglossoides (L.) Ker-Gawler 000875 Orchidaceae Spiranthes odorata (Nutt.) Lindl. Sher-2013 McMillan and Blackwell: Catesby plants, SI oane and Oxford her 6 Poaceae Andropogon tenuispatheus (Nash) Nash 00087576X Poaceae Andropogon tenuispatheus (Nash) Nash H.S. 212f.87 Poaceae Arundinaria tecta (Walter) Muhlenberg 00087480R Poaceae Arundinaria tecta (Walter) Muhlenberg Sher-0184 Poaceae Arundinaria tecta (Walter) Muhlenberg Sher-0 184-2 Poaceae Cenchrus H.S. 212f.84 sp. Poaceae Cenchrus 00087496Y sp. A Poaceae Cenchrus incertus M. Curtis Sher-0 132 Poaceae Chasmanthium latifolium (Michx.) Yates H.S. 232 103 f. Poaceae Chasmanthium sessilifolium (Poiret) Yates Sher-0 177 = Poaceae Chloris elata Desv. Chloris polydactyla 00087545T Poaceae Chloris radiata (L.) Sw. 00087549X Poaceae Coelorachis rugosa (Nutt.) Nash H.S. 212f.85 00087584W Poaceae Coleataenia species 1 in prep, ssp, rigidula per Weakley Wood Poaceae Ctenium aromaticum (Walter) H.S. 212 44 f. Wood Poaceae Ctenium aromaticum (Walter) Sher-0 139 Wood Poaceae Ctenium aromaticum (Walter) 00087537U Wood Poaceae Ctenium aromaticum (Walter) 00087541P Poaceae Danthonia sericea Nuttall Sher-0 187 Poaceae Digitaria sanguinalis (L.) Scopoli Sher-0 125 Poaceae Echinochloa H.S. 212 44 sp. f. Poaceae Echinochloa crusgalli (L.) Palisot de Beauvois 00087523P Poaceae Eleusine indica (L.) Gaertn. H.S. 212f.85 Poaceae Eleusine indica (L.) Gaertner 00087562S Poaceae Elymus 00087521N virginicus L, Poaceae Eragrostts cilianensis (Allioni) Vignolo ex 00087570R Janchen Poaceae Eragroslis Watson 00087566W elliottii S. Poaceae Gymnopogon ambiguus (Michx.) BSP Sher-0 186 Poaceae Hackelochloa granulans Kuntze 00087527T (L.) Poaceae Indet. H.S. 232f.61 Poaceae Indet. H.S. 212f.45 Poaceae Leptochloa panicea (Retzius) Ohwi Sher-0 182-3 Poaceae Leptochloa panicea (Retzius) Ohwi ssp. brachiata 00087529V Snow (Steudel) N. Poaceae Leptochloa panicea (Retzius) Ohwi ssp. brachiata 00087533Q Snow (Steudel) N. Poaceae Leptochloa panicea (Retzius) Ohwi ssp. brachiata Sher-0 182-2 Snow (Steudel) N. Poaceae Leptochloa H.S. 232 103 sp. f. Poaceae Paspalum floridanum Michx. H.S. 232 117 f. Poaceae Paspalum floridanum Mchx. 000875310 Poaceae Paspalum floridanum Michx. 00087539W Poaceae Paspalum floridanum Mchx. H.S. 212f.83 Poaceae Phalaris caroliniana Walter H.S. 232f.61 Poaceae Phleum arenarium L. H.S. 232f.61 Poaceae Saccharum brevibarbe (Michx.) Persoon var. 00087547V R um contorf (Elliott) Webster Poaceae Saccharum brevibarbe (Michx.) Persoon var. Sher-0 183 contortum R. Webster (Elliott) McMillan and Blackwell: Catesby plants, SI oane and Oxford he Poaceae Saccharum giganteum (Walter) Pers. H.S. 212L86 Poaceae Saccharum giganteum (Walter) Pers. Sher-0108 Poaceae Saccharum giganteum (Waiter) Pers. 00087543R Poaceae Saccioiepis striata (L.) Nash 000875 19U Poaceae Sacciolepis striata (L.) Nash 00087580S Poaceae Setaria corrugata (Elliott) J.A. Schultes Sher-0118 magna 212L82 Poaceae Setaria Griseb. H.S. Poaceae Setaria parviflora (Poir.) Kerguelen Sher-0 116 Poaceae Setaria parviflora (Poir.) Kerguelen H.S. 212 44 f. Poaceae Setaria parviflora (Poir.) Kerguelen H.S. 212L83 232L30 Poaceae Setaria parviflora (Poir.) Kerguelen H.S. Poaceae Setaria parviflora (Poir.) Kerguelen Sher-0 116-2 Poaceae Setaria parviflora (Poir.) Kerguelen 00087564U Poaceae Setaria parviflora (Poir.) Kerguelen 00087568Y & Poaceae Setaria pumila (Poir.) Roemer Schultes Sher-0 116 212L86 Poaceae Sporobolus indicus (L.) R. Br. H.S. Poaceae Sporobolus indicus (L.) R. Br. H.S. 212L85 Poaceae Sporobolus indicus (L.) R. Br. 00087572T Poaceae Sporobolus indicus (L.) R. Br. Sher-0 145 00095710M Poaceae Tripsacum dactyloides (L.) L. Poaceae Uniola paniculata L. 00087525R Poaceae Uniola paniculata L. Sher-0 176 Poaceae Uniola paniculata L. Sher-0 176-2 232L56 Poaceae Uniola paniculata L. H.S. 212L88 Poaceae Zizania aquatica L. H.S. Poaceae Zizania aquatica L. var. aquatica Sher-0 103 232L67 Pontederiaceae Pontederia cordata L. var. cordata H.S. Pontederiaceae Pontederia cordata L. var, lancifolia (Muhlenberg H.S. 212 19 f. ex Torrey Elliott) Ruscaceae \1 uanthemum tcemosum (L.) Link H.S. 212L60 i Ruscaceae Ma mthemun em Link 00087486X )sun (L.) i i Ruscaceae Maianthernum racemosum Link 00087490S (L.) Ruscaceae Nolina georgiana Michx. Sher-0762 Ruscaceae Molina georgiana Michx. Sher-0762-2 212L32 Ruscaceae Nolina georgiana Michx. H.S. Ruscaceae Polygonatum biflorum (Walter) Elliott 000875 14P Ruscaceae Polygonatum biflorum (Walter) Elliott H.S. 212L60 232L31 Smilacaceae Smilax auriculata Walter H.S. Smilacaceae Smilax bona-nox L. 00095796- Smilacaceae Smilax bona-nox L. Sher-2217 Smilacaceae Smilax bona-nox L. Sher-2222 Smilacaceae Smilax bona-nox L. Sher-2227-2 Smilacaceae Smilax herbacea L. Sher-2219 00095792W Smilacaceae Smilax hispida Raf. Smilacaceae Smilax lauriflolia L. Sher-2229 Smilacaceae Smilax pumila Walter 00087200H Smilacaceae Smilax pumila Walter Sher-2223 Smilacaceae Smilax pumila Walter Sher-2223-2 Smilax pumila Walter H.S. 212L95 Morong 00095800M Smilacaceae Smilax, smallii Morong Smilax smallii Sher-2221 McMillan and Blackwell: Catesby plants, Sloane and Oxford 8 Smilacaceae Smilax 00095788 sp. Smilacaceae Smilax Sher-2225 sp. Tofieldiaceae Triantha racemosa (Walter) Small H.S. 232 117 f. Sher-sn-ATG Trilliaceae Trillium catesbaei Elliott 00087470Q Trill iaceae Trillium catesbaei Elliott 212T59 Trilliaceae Trillium catesbaei Elliott H.S. Trilliaceae Trillium cuneatum Raf. 00087474U Trilliaceae Trillium discolor Wray ex Hooker 00087478Y Trilliaceae Trillium discolor Wray ex Hooker Sher-sn-ATH Trilliaceae Trillium maculaturn Raf. H.S. 212T59 Xyridaceae Xyris amblgita Bey. ex Kunth H.S. 212 42 f. Xyridaceae Xyris ambigua Bey. ex Kurith 0008755 1Q Xyridaceae Xyris ambigua Bey. ex Kunth 00087563T Eudicots Acanthaceae Dyschoriste oblongifolia (Michx.) Kuntze H.S. 232 129 f. H Altingiaceae Liquidambar styraciflua L. S 212f 79 H amar 73? 34 Altingiaceae iqui styracifua S f 00087448V Altingiaceae Amaranthaceae 00087426R ' y I y. t ' ,| .), • | ,vy, ,| Ml v , Clemants Anacardiaceae Toxicodendi on pubescensP. Miller 00087442P Anacardiaceae Toxicodendron radicans (L.) Kuntze H Anacardiaceae toxicodendron vermx (L.) Kuntze S 212f 25 Anacardiaceae toxicodendron vermx Kuntze 00087212K (L.) Anacardiaceae oxicodendron -vermx (L.) Kuntze 000872160 t Anacardiaceae oxicoaenaron vermx Kuntze 0009571 IN I (L.) Ap aceae Angelica venenosa (Greenway) Fernald 0008733 5Q Ap Angelica venenosa (Greenway) Fernald 00087347T Ap aceae Angelica venenosa (Greenway) Fernald Sher-0622 Ap Angelica venenosa (Greenway) Fernald H.S. 212f.39 Ap Cicuta maculata L. 00087323N Ap Cicuta maculata L. 00087330L Ap Cicuta maculata L. 00087343P Ap aceae Cicuta maculata L. 000873510 Ap Cicuta maculata L. Sher-0626-5 Ap aceae Cicuta maculata L. H.S. 212f.27 Ap Eryngium integrifolium Walter 00087334P Ap aceae Eryngium integrifolium Walter H.S. 212f.41 Ap Eryngium integrifolium Walter Sher-0614 Ap Eryngium yuccifolium Michx. 0008733 8T Ap aceae Osmorhiza longistylis (Torr.) AP. de Candolle H.S. 212f.32 Ap Osmorhiza longistylis (Torr.) AP. de Candolle H.S. 212f.34 Ap aceae Osmorhiza longistylis (Torr.) AP. de Candolle 00087339U Ap Osmorhiza longistylis (Torr.) AP. de Candolle Sher-0635 1M Ap aceae Ptilimnium capillaceum (Michx.) Raf. 0008733 Ap aceae Ptilimnium capillaceum (Michx.) Raf. Sher-0621 Ap Ptilimnium capillaceum (Michx.) Raf. H.S. 232 116 f. Ap aceae Thaspium barbinode (Michx.) Nutt. H.S. 212f.58a Ap Thaspium trifoliatum (L.) A. Gray var. trifoliatum H.S. 212f.37 Apocynaceae msonia ta ernaemontana a ter Sher-0551 Amsoma ™ Apocynaceae tabernaemontana 00087427S a ter Apocynaceae / msonia ta ernaemontana ajter var. H.S. 212f.37 Apocynaceae Apocynum cannabinum L. Sher-UjjU-z Apocynaceae Apocynum cannabinum 0008743 IN L. Apocynaceae Apocynum cannabinum L. H.S. 212f.57 Apocynaceae Asclepias amplexicaulis Sm. Sher-0582 Apocynaceae Asclepias amplexicaulis Sm. H.S. 212 f.30 a Apocynaceae sclep as hum,strata a Sher-0586 ' ^ H Apocynaceae ^, sc epias umisra a a er S 232 f 86 J' Apocynaceae Asclepias numistrata waiter H.S. 212 f.30 Apocynaceae Asclepias michauxii Decaisne 00087214M H Apocynaceae SC ep,aS °~'ovata '° S 232 f 1 14 H Apocynaceae sc epias perenms a S 232 f 122 ei Apocynaceae Asclepias rubra L. Sher-0587-2 Apocynaceae Asclepias rubra L. H.S. 232 f.83 Apocynaceae Asclepias rubra Asclepias lanceolata Sher-0565 L. (left) W alter (right) Apocynaceae Asclepias tuberosa L. Sher-0574-2 H Apocynaceae sc epias tuerosa S 212f 31 . H Apocynaceae S 212f 30 Apocynaceae Asclepias variTata Sher-0567 I Apocynaceae sc epias vertici ata Sher-0566 . Apocynaceae Asclepias verticillata L. Sher-0546 H Apocynaceae Asclepias verticillataU S 212f 30 Apocynaceae sc epias viri ora Sher-0583 i] Apocynaceae Asclepias viridijlora Rat. H.S. 212 18 f. Apocynaceae Cynanchum taeve (Michx.) Persoon Sher-0548 Apocynaceae Matelea carolinensis (Jacq.) Woodson H.S. 212 17 f. H Aquifoliaceae C S 212 f 15 qu o laceae //ax; "mbiZa Sli-hx Torrey //ex fearf/e/ 0008725 5R j W W Ashe qu forlaceae 0008725 5R W W Ashe qu aceae //ex cosine L. Sher-0250 . ! qu aceae ex cassine 0008720 8P . . ! qu aceae excassme 000874371 . ! qu aceae a cassme 0008744 lO * „ . ! Aquilohaceae C SS'"e 00095687Z f T' 000957030 Aquifoliaceae jlex cassine Aquilohaceae Ilex cassine L. Sher-0256 Aquifoliaceae Ilex cassine L. H.S. 212 f.65 H 212f 636 S 65 H AXraceae AmpTlmtercar^^nianus (Walter) Ncsom S 212 f 83 Asteraceae Ampelaster carolinianus (Walter) Nesom 00087344Q Nesom Asteraceae Arnpelaster carolinianus (Walter) H.S. 232f.41 Asteraceae Arnoglossum atriplicifolium (L.) H. Rob. H.S. 212 f.6 8U Asteraceae Arnoglossum atriplicifolium (L.) H. Rob. 0008725 Asteraceae Berlandiera pumila (Michx) Nutt. 00087358V Asteraceae Berlandiera pumila (Michx) Nutt. H.S. 212f.34 McMillan and Blackwell: Catesby plants Sloane and Oxford her Asteraceae Bidens bipinnata L. 0008725 4Q Asteraceae Bidens bipinnata L. 00087274S Asteraceae Bidens frondosa L. H.S. 212 f.8 Asteraceae Bidens sp. H.S. 212 f.7 Asteraceae Bigelowia nudata (Michx.) DC. H.S. 212 74 f. Asteraceae Borrichiafrutescens (L.) A.P. de Candolle 000873 18R 00087263Q Asteraceae Brickellia eupatorioides (L.) Shinners Asteraceae Brickellia eupatorioides (L.) Shinners 0008732 1L Asteraceae Carphephorus carnosus (Small) C.W. James H.S, 232 £30 Asteraceae Carphephorus 00087370P sp. Asteraceae Chaptaiia tomentosa Vent. H.S. 212f.35 Asteraceae Chaptalia tomentosa Vent. Sher-1982 Asteraceae Chrysogonum virginianum L. H.S. 212 17 f. Asteraceae Chrysoma pauciflosculosa (Michx.) Greene Sher-1686 Asteraceae Chrysopsis gossypina (Michx.) Sher-1762 Elliott Asteraceae Chrysopsis gossypina (Michx.) Elliott H.S. 232 42 f. Asteraceae Chrysopsis gossypina (Mchx.) 00087290Q Elliott Asteraceae Chrysopsis mariana (L.) Elliott H.S. 212f.96 Asteraceae Chrysopsis mariana (L.) Elliott 00087269W Asteraceae Chrysopsis mariana (L.) Elliott Sher- 1887-2 Asteraceae Chrysopsis mariana (L.) Elliott Sher-1889 Asteraceae Chrysopsis mariana (L.) Elliott H.S. 232 42 f. Asteraceae Chrysopsis mariana (L Elliott H.S. 232f.64 ) Asteraceae Cirsium repandum Mchx. Sher-1638 Asteraceae Cirsium Sher-1639 sp. 00087322M Asteraceae Cirsium sp. Asteraceae Cirsium virginianum Michx. Sher-1637 (L.) Asteraceae Conyza canadensis (L.) Cronquist var. pusilla Sher-1745 (Nuttall) Cronquist Asteraceae Coreopsis delphinifolia Lam. Sher-1960 Asteraceae Coreopsis delphiniifolia Lam. H.S. 232f.29 00087278W Asteraceae Coreopsis lanceolata L. Asteraceae Coreopsis lanceolata L. 00087376V Asteraceae Coreopsis lanceolata L. Hort-048-056c Asteraceae Coreopsis lanceolata L. H.S. 232 123 f. Asteraceae Coreopsis lanceolata L. H.S. 212f.20 Asteraceae Coreopsis major Walter var. major H.S. 212f.33 Asteraceae Coreopsis major Walter var. ma/or H.S. 232f.48 Asteraceae Coreopsis major Walter var. rigida (Nuttall) 0008725 IN F.E.Boynton Asteraceae Coreopsis major Waiter var. rigida (Nuttall) 00095700L F.E.Boynton Asteraceae Coreopsis major Walter var. rigMa (Nuttall) Sher-1968 F.E.Boynton Asteraceae Coreopsis major Walter var. ng/'da (Nuttall) Sher-1969-2 F.E.Boynton Asteraceae Coreopsis pubescens Sher-1965 Elliott Asteraceae Elephantopus carolinianus Rausehel Sher-1632 00087371Q Asteraceae Elephantopus tomentosus L. Sher-1991 Asteraceae Erigeron quercifolius Lam. 00087374T

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