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THE VASCULAR FLORA OF MADISON COUNTY, TEXAS PDF

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THE VASCULAR FLORA OF MADISON COUNTY, TEXAS Amanda Hugh and D.Wilson 1 Neil! K. = USA. 'Station.^?7843-3258, College ABSTRACT Cyperaceae spp). ( )' spp.). Tin ]\ u> arc ..m eel list that includes i i n name, collection citations, anc Twlicihei thai specie i W duded,n,he h S e P ; tottrrru rspedetlnd n en ra re a d S ( RI'SrMFN !m on [inn <o. noii ii 1 i \ i. i u 1 ' i i I l i i i i ii i m up l,o, [>,.„, u-< SA p n„ u linuompil kirn n un. h-t u -nieiit ulaqiu Inv. 1 i I i i , m nil. v< ml in u nl d. la ih ioik in' u ( i« I i ( i \ ii t ii i 1 INTRODUCTION many Madison Count) like oihei o until ol ha n ed an in-depth flo- 1< i 1 I lom>k Texasbota- evaluation \u>ni| u im ha n u,oalof ristic U id< it - ,t r f 1 t M) was intended irn-volume but nists.Lundell's Flora \n-a h> to be a series, oj /, Ma the project was abandoned in 1955. Correll and Johnston's nualofthe Vascu- and remains only Plants Texas (1970) incorporated Lundell's work, the lar of reference with dichotomous keys encompassing the entire state. Updated check- Gould Hatch of the Texas flora have been compiled by (1975a), et al. (1990), lists and Jones Although these reference species by region or vegeta- et al. (1997). due no county-by-county species information available to the tional area, is Manual Reed compiled the the Dicot dearth of local floristic studies. (1997) of Madison Flora Brazos and Surrounding Counties, technically including of County. However, than 200 collections from this county were available for less A&M (FTC reference from Texas University's two herbaria at the time Her- Bnn barium ipeeimen inten reference). M ei Almost a century ago, Texas botanist William Bray stated that "before the kxas loia suffers further radical changes, the schools of the ought ol state to I coopciaic ni securing a complete and authentic species represented by oi lisi carefully collected and well-preserved specimens" While t,Brav E)0b). the col- haw and leges universities of Texas a great n adit ion ecological and system- oi ,u ic siudies a century has brought us but little closer to a modern state-wide loia. radical changes which Brav spoke continue unabated and lie lorist ic o\ i I i often unobserved, and the labor intensive loras of Texas counties are woe- still I lully tew. The were and objectives of this count)' flora identify preserve to collect, ) I specimens ol all vast u la plants occurring in Madison ,ount\ lexa m r ( ' g< ' i ate a species lor he count v and delineate pal terns plant communities by list oi I and habitat preference; 3) analyze the flora for disjuncts, notable range exten- endemics, and sum- sions, recent invaders, rare taxa. This article provided as a is mary of the first authors M.S. thesis research (.Neill 2000)* information also is available on the Madison County Flora Website <http:// at B10COURSE.B10.TAMU.EDU/graduatc-students/neilla/madhome.htmxThis site includes annotated species searchable in several formats, as well as lists, mans and links images. to Description Site Madison County comprises 302,451 acres (122/398 ha) in east-central Texas W between 30'49' and "06' N and 9537' and 96 (Ramos latitude 3' longitude "1 3 1 The 1997). county bordered on the west by the [\a\asota River and Brazos is County, on the east by the Trinity River and louston County, on the north by I County Leon and on the south by Bedias Creek and Ci nnesand Walker coun- The county and ties (Fig. seat largest city Madisonville, situated about 90 1). is NNF km) mi (142 Houston. of The land surlaci flat to gent rolling unl genetal .lopes to the south- l\ l\ i east, with elevations ranging from 420 ft (128 m) above sea level south of Normangee to 140 (A in above sea lead along the lower Trinity River and 3 i t » Bedias Creek (a tributary of the Trinity) CU.S.G.S. I9b2-I989). The eastern two- thirds of the county are drained by these watercourses, which converge the at While southeastern tamed corner. the Trinity hydrologically with a river a is now historical flood plain 10 miles wade and largely under cultivation, Bedias Creek most deep for length a small, creek with extensive bottomlands is of its oxbow ol lakes, swamps, and frequently flooded ridge and -swale topography. u The Navasota drams River tributary ol the Brazos River the western third of J Madison Count}', where meanders through wide and wiyxled loodplam a it ol I sloughs, natural and lakes, terraces, ravines. d The geology of Fast Texas the result alluvium washed southward by is m and rivers deposited layers upon he advance and retreat of coastal waters in t the Gulf of Mexico. This deposition occurred in the region during the Eocene Madison County, Texas and and map bottomland and vegetation zones. Adapted from Gould et (1960) Neitsch Location of soils al. (1 Fig. 1 . Epoch (58 to 37 mya) of the Tertiary Period C pearinj L991) The Eocene layers members Group and com- deposited Madison County are of the Claiborne are in and (Neitsch Neitsch posed sandstones, lignites 1994). of clays, shales, siltstones, and with upland three (1994) 12 associations in the county, six soils lists soil loamy sands and Three bottomland soils of alluvial clay prairie soils, all clays. modern un young cover flood pi tncl terra; i : The Madison County growing season averages 250 days, with a frost-free maximum 96T mean minimum mean temperature and Janu- July of (35°C) a and and ary temperature of ~]tVF TC). Precipitation peak:, in the spring fall av- { cm) (Ramos annually erages 41.6 in (106 1997). Cultural History Madison Although no major archaeological sites have been found in County, some Indian camp and kitchen-middens have been found that date from sites Caddo 3000 (Madison County Historical Commission 1984). Tribes of the B.C. Confederacy formed permanent settlements in the eastern part of the county, where the people ultivated corn and squash lorn n ibes that occupied sdic c I who Creek and confluence the county include the Bedias, frequented Bedias its with the Trinity River, and apparently had a village in this peninsular area Almanac (Neitsch 1994; Texas 1951). French and Spanish explorers, missionaries, and soldiers began leading The Spanish expeditions through the area in the mid-sixteenth century. first Bucareli-where hghway settlement, present-day Texas 21 crosses the Trinity I River-was established in 1774 as an outpost against French invasion. Bucareli was abandoned in 1779 due 10 Idocxlint; epidemics, and raids by the nomadic Comanches (Madison County Commission listoneal 1984). 1 Madison County most is historically significant for containing the inter- two section of earl}' roads. Originally Indian these roads were used by trails, missionaries, explorers, and eventual!)' The Old San Antonio Road settlers. known (OSR), also as Hi Cainino Real, or the Kings Highway, connected settle- ments in the hast with those in San Antonio and Mexico Today, this modern highway forms most northern Madison town o\ the bordei oi ounty. At the of I Midway, another road known La Balna from OSR and as Trail split led south- west to the mission at Goliad (U.S. Dept. o{ the Interior 1998). Madison County was organized and in 1853, settlement increased rapidly to a population of 3000 by 1800 (Texas Almanac 1951). By 1900, the population had grown and to 10,432, 71,000 acres were being farmed (.Richardson 1940). Over 20,000 acres of this was cotton. Three railroads were through in built the and county, the population peaked in 1930 at 12,227 inhabitants (Ramos 1997; Richardson and 1940). Pasture haylancl for beef cat prod action have replaced le t most cultivated crops today. Seventy percent of the county used as rangeland, is and hayland, pasture (Neitsch 1994). Fishing and hunting are important land many in ihccounty with in propertic leased easonal duck and deer hunting. to i Botanical History map Every vegetation available represents Madison an '.omit v as ecotone. Past i and ecologists botanists in Texas recognized that he area around the 96 th me- I which ridian, divides the county in half, a zone where western xerophytic is grasslands mingle with mesophytic eastern forests (Bra)' 1906; Blair 1950; Tharp Cronquist 1982; 1920). Allied and Mitchell (1955), Correll and Johnston Cory and Parks and Gould upon (1970), (1937), et (1960) generally agree the al. boundaries of the three major vegetation zones converging Madison County; in commonly these vegetation zones are called the Pineywoods, Blackland Prai- and Oak Savannah Post ries, (Fig. 1). m Correll anil Johnston 1970) described the vcgctational area-, largely ac- ( cordance with The Pineywoods past descriptions. region of Hast Texas repre- sents the most mesophytic area of Texas and characterized by extensive pine is and pine-hardwood swamps Crests wit intermittent Madison ountvsoast li ( M ern edge one the most westward-extending, tongues is ol this vast vegeta- nonal Blackland \<i\c lie Prairies region lies in hive diagonal bands trending "I t NE-SW Of wo across east-central Texas. the southern bands Blackland of Prai- t San Antonio the Prairie occupies the border northwestern Madison rie, of County, and the Payette Prairie is sometimes shown to extend into the south- central portion the county. The Post Oak Savannah region mterdigitates with ol" the Blackland Prairies. A belt of Post Oak Savannah occupies most central of Madison County. Although no \hn iu tnuonal ndu U mi onducted Madi m \egi in lia\ l * County, past lescan h h included Holistic ispeel found within the eount\ Launchbaugh Both 952) and High tower 987) changes J invc ligated in veg- (J I etation structure n ;pon etodisturbaua Anion Diamond in in th< u 10 Prairie. ., remnant communities which (1980) described plant of the Fayette Prairie, is tht south* rn disjunct of the Black land proper. ing the northern limits Pi pla< Grimes and Walker McCaleb of the Fayette Prairie in counties. (1954) analyzed m m m the vegetation of t!u outhernP tOal ih hid( d Madison County \ i I within his Oak-Hickory Forest Region. Allen (F)7! vcgetational study of the s ) lower Navasota described plant ommunitie dong he watershed Grimes in i . i County and pointssouth. Apparent v none the above investigators conducted o\' 1 research within Madison County. There no voucher specimens are resultant for (TAMU) the county in the Tracy (TAES) or Biology Department herbaria Texas at A&M (FTC Herbarium Specimen University Browser, internet reference). Mi-inons Vascular plants were collected frequent and regular intervals in Madison at May County, from 1996 to April 2000. Public lands (roadsides, railroad rights- unm of-way, city parks ami Miiemru n io ml habitats identified as \\ i > unusual were With news- revisited over several seasons. the assistance of a local paper article (Levey 1998), private landowners were contacted for permission to collect on their prop* tn s >thei pi ivate land wen hosen with the aid of ( i ; < photographs and and aerial .,vm lh gin in a .in bartered plain topographic ill soils maps, in order to u.r\ bot h gi ultural h\< toed n id-use areas and to y i I unusual undisturbed Urban investigate possible or habitats. areas (Madisonville, Normangee, North Zulch. and Midway) provided disturbed upland sup- sites weedy porting a unique, flora Riparian ind lacustrim terns were accessed \ bridges, on private property, and by canoe. at At the time of plant collection, data were recorded on the specific habitat and abundance relative of the species in the area. Publications describing similar research were reviewed for exposure to floristic protocol. References include Allen McCaleb Reed Starbuck and Wilson UQ84). (1974), (1954), (1997), (1972). o R< it r< rn - us< d for pla t mil iti n u orr< 11 hn to 1°, to ii * t I « < | i i [ et al. (1999), and Reed (1997). Grasses were identified with The Grasses of Texas (Gould 1975b), and identifications were verified by Stephan Hatch and Dale Kruse at the S.M. Tracy Herbarium in College Station, Texas (TAES). Sedges (Carex sp.) were identified with an unpublished key to the Carex of Texas provided by Stanley Jones, curator the Botanical Research Center allege Station, Texas oi ( (BRCH), and identifications were verified by him. Other experts consulted were Momque Reed (TAMU). Theodore Barkley the Botanical Research Institute at Texas (BRIT), and Richard Rabelerai (he University Michigan Herbarium of o! (MICH).Taxonomtc onquistian system classifit ation nti follow* d ilu \ i ; ;i 1 1 1 ( i U 'ronquisi 1088). wirh the exception ol familial treatment ol the Tupelo family Chiekenspike family (Sphenocleaceae and Dodder family iNyssaceae), ), (Cuscutaceae). Voucher specimens collected throughout the project were de- A&M posited in the herbaria at Texas University in College Station, Texas (TAMU, TAES), and are available for examination. A search the Flora of Texas Consortium Browser C.OOOa.) revealed some ol m species that the authors did not find thccounty,and these are included in the TAMU TAPS Only those specimens held and were seen by the au- checklist. at Numerous specimens had been landed "Madison County" but older ac- thors. were northwest Normangee Icon County. The location tually collected of in SMU and ASTC, TEX, SHST, and identification of species collections held at LL, have not been verified. AND DISCUSSION RF.SULTS CUT Madison County vascular plant collections resulted in a total ol speci- mens. The flora comprised of 1041 taxa with 985 species in 488 genera and is m The 136 27 Poaceat families. fiv't largt pi ntlainili< Vstci ic (1 species), t i Fabaceae spp Cyperaceae and (123 spp.), Uscn.su stricto) (70 (63 spp.), ). 44% Fuphorbiaceae (32 spp.). These live families con tain approximately of the species found in Madison County. Further numerical distributions of the flora are found in Table 1. Major Vegetational Habitats Thirteen major habitat types were classifiable in Madison County. These habitats are essentially those described, lor Robertson Count)', Texas, by Starbuck (1984). Riparian habitats and lowlands include habitats in water, near water, and low ARM), and areas within floodplains. Aquatic emergent aquatic floating (AFF), ( submerged (ASUJ and aquatic are habitat categories obligate aquatic species lor wen dm those plants that found in landing water io loot ling Habitats along the banks and frequently (.two or more times a war) flooded areas of riverine and lacustrine bodies oi water were divided into major riparian (RUM) and (RUM) minor riparian habitats. Major riparian habitat that of the Navasota is and Trinity whereas minor riparian habitats are those along creeks and rivers, ponds. Rowland habitats flooded once year or [uenily were divided by less a re. i woody lowland woods (TIW). lowland open cover into terrestrial terrestrial (TFO),and lowland mixed (TFM). Uner upland areas were similarly terrestrial divided upland woods upland open (TUO), into terrestrial (.TUVV). terresii ia.1 and terrestrial upland mixed (II VI) [wool uirbucl habitat groups, parasite UTI (PAR) and epiphyte better describe plant habits, but were maintained for ), BOO wen consistency in this te> [wool tarbucl habitat H used. Hi bog (, Sphagnum habitat applies to the acidic bogs found the deep sands of in Robertson County (Bryant 1977; Starbuck 1984). This habitat not present in is Madison Count) was St rbucl rn trial prairie(TPR) habitat used in his text i specimens by Hightower remnant solely for collected (1987) at a prairie locality. mm m Although certain pi ni p kIm m uuld In linn i_i d \ i > ! i i < > t \ i no undisturbed county prairie species, prairie habitats are intact in the today. oxbow Aquatic habitats such as lakes, stock ponds, reservoirs, drainage Uncom ditches, and shallow creek and river margins supported a unique flora. - mon mCid herbs found ju.ur lumbolutea,Thaha tin h h bitai h * />. , > dealbata, and Utricularia radiata. Lowland and riparian habitats contain the most diverse flora in Madison County. The extensive wooded loodplams of the f Navasota River and Bedias Creek exemplify typical bottomland hardwood veg- common etation. Trees most in low, moist areas of the county include Carya aquatica,Fraxinuspcnn yiviimca Nyssa sylvatua Plan ra aquatica, Querent lyrata,Q nigra, phellos, Tilia americana, and Ulmuscrassifolia. Disturbances Q. and Due to this area include logging, flooding, cattle grazing. to agricultural use and the construction of levees and dams, the Trinity River floodplain for- om umnd when ests are less extensiu vu >Mtion ol sjk n w comparing ( < i collections from the two major riparian habitats (RLM). Species found only Amtoma within the Navasota River floodplain include Aristolochia illuttris, tomentosa,Baptisiii alba Brunniehia ovata Carex cnmpl<iihita erut-eorvi. ( Mikaniascandens.lHaticniaquatica^PrunusearoliiuumKTiUandsiarecurvata, mnmm mdma onb Urticachamacdiytudi lounb aiinn <uu\ u << >\> • > the Trinity River floodplain include Amorphafruticosa, Ampelopsis cordata, Carex am,h retroflexaXlcmah nln uluui qui iuu< hu viyai imi 'l'i c uto I noodmnhmm Panicum virgatum,. .m nhitn .ttnl low land open habitats (l r< I | (TLO) were typical urnd id Forested areas or river flood plains, low pas- I md tures, moist roadsid Inch oth ropenarea near bodies of water. These mowing and and habitats are often disturbed by grazing us support variety th a mm pm pn wlli weedy invade mini ml of n i i | upland Madison County prime and Level sites in are locations for pastures m haymeadows,asthey were mostly cleared the century cropland. Most last for wooded with what would con upland always found areas be idered nearly in in along creeks, and so \vou Id be included in die terrestrial lowland classifications ITUM) or this stud True wooded upland 'W'iand mixed upland habitats siios^.'1'l v. I may do occur, and these be carcgoi ized into h ve general association types: t Oak - Winged Elm - Yaupon Post Association 1. This association encompasses nearly upland wooded sites in the county, and all huypical [ho 1'ostOak Savannah vegetation type he association found in ol is I areas with sandy loam topsoil and, gra\ clay subsoils anil characterized by is Quercus Ulmusalata/and vomitoria. Ilex stellata, Oak - Oak - Winged Elm Blackjack Post Association 2. Qucnusmanlandu. Characterized by the presence o^ sites exhibiting this as- u, Oak - sociation arc generally more poorly drained than [hose with the Post Winged Elm Yaupon The - association. aspect of these sites also that of the is Post Oak Savannah vegetation pe but was ivquent ly iound with tall grasses in 1 such Andropogonternarius, seen in the Blackland Prairie vegetation type, as Soiyhastrum and Schizachyrium scopanum. nutans, Gum Southern Red Oak - Sweet - Post Oak Association 3. This association occurs the eastern third the county and character- is in ol is by dominance Quctx and u/mT.mdv/ These ized a oi us jalcata styraciflua. / sites have acidic sandy soils Nvith \iood drainage. The best examples of this as- unwu Cobb sociation arc found the uplandsalong Youngs reekc and Creek in (. The Canyon. These localities are discussed in the Unique labitats section. I Padina series of deep sands frequently iound these sites. Sassafras dl bid urn is at and Ptnuslacda are also found this association. in upland open habitat dFl'CYis ubiquitous and easi accessed in Madi- lei'ivsii iul In son ounty. ike open lowland sues, these generally roadside and pasture habi- ( I mm num ma are characterized heib and hough heic that tats In hi is it many showy may Open of the and popular wild! lowers Texas also be seen. ol summer when upland habitats are first to suffer in drought conditions, even mown narrow county roadsides arc lor hay in [Ins county difficult to find is It examples of this habitat type that have not experienced disturbance. Pasture are frequently invaded by Prosopisglandulosa and Acacia Jarncsiana. sites Epiphytes found county (EP1) in the are restricted to Pleopletis and (PAR) found polypodioidcs, Tillandsta rccurvata, usneotdes. Parasites in /'. the county include Cuscuta cuspidatd, C.glomcrata. C. oblusifolia, Orobanchc and Phoradcndron tomentosum. multijlora, Unique Habitats m Several localities Madison County primarily on private lands, exhibit un- usual characteristics that deservespccial mention. Isolated populat lonsol I'mih tacda are found at several places in the countv. but the only area that deseiwes woods lound designation as the Hast k as Piney vegetation typ< n out h i eastern Madison Count), on the peninsula formed by he eon luenceol the Trin- I i River and Bedias Creek and extending up Bedias Creek for several miles, ity hardwood lowland Species although area o contain ;onn forest col- this il Caicx na lected only thi habitat include \nsioloihiu ei pentai ui. hvali in. Dioscorea ixgodhun japonicum ninth, pracco\ xillasa in pi ' 1 Nointact,undistui'jfflpn(. oft an \ntonio Blackland Prairie are pre- ha ; OSR served in Madison County. The narrow band oi prairie that once followed way 45 has been from the Navasota River to a few miles oi Intel tat< tgh t i ' 1 1 woody Although and encroachment the road- lost to grazing, crops, of species. mown, some road contain pock- sides are occasional portion oi tin .rretehol \ I support herbs along the fencelmes lh sites spe- ets of prairie rid reel in e - Eriogonum including Astragalus Buglossoides arvensis, cies distortus, m cum Orbexilum Pen- Eryngt m, latum, simplex, longijolium, u vucetjolt u ivi h a maxima tcmou lan'ijhna R and Silphium radula. in <Wis Uiiiliu t I > OSR county wildflower viewing Roadsides along are the best places in the for and summer. in the spring ome Hit Ka\ ota IV will continue to host the most unique habitat -r if i in Madison Count}' he wild stretch of south o\ ake milestone is allowed if it t I I remain unimpouncled. The upper terraces along the consist deep to rivet oi spn sands where a high water table produces my, a nil nearl v bog-like seeps. Here, two springs supported a number of species not found elsewhere in the county: Galium obtusum. tlvdnxoiylc ranunculoidesjuncus Baccharis halimijolia, Saccharum Ranunculus baldwinii, debilis,Polypogon mouspclicnsis. scclcral us. Sagittariaplatyphxlla. and laxndium distich lows oi these springs have urn. lie I I been altered by landowners to provide water lor livestock, but the habitat is m m p.m rounded by Myrica and rich unique nu h o h id a spi nig still i i , , and Rubus argutus and harbored a few species typically associated with cerifera such hea Rhexia mat tana and b'uirena squarrosa. true bogs, as Plan foetida, and Unfortunately, h)°° an v oat* d to u an small tank, the in tin a \< i i < may unusual hydrology not continue support these plants. altered to Numerous and oxbows found along Navasota River natural lakes are the m Oxbows low woods near the river host several aquatic and terrestrial species m Some rarely or never seen elsewhere the count}'. of liese are Didiplisdiandra, t and pcnxOldcnlandia Sauru ruscernuus, uncus Eragrostishypnoidcs. re host ii, j OSR Utricularia radiata. Just south of on the Navasota River lies a string of natural lakes in the open loodplam. These lakes have a unique flora differing f from that of the oxbow lakes, Species found in and around these lakes include Nelumbo americana, Bacopa Eleocharis palustrisjusticia lutea, rotundifolia, Nuphar Sphenoclea zeylanica, and Zizianiopsis miliacea. Most of these lutea, water species were found nowhere else in the county. Plants in these bodies of somewhat are protected Irom curious hotaiusts by snapping and turtles nest- ing alligators. The Navasota River has created ancient high winch appear river terraces, as a series of hills that are easily seen from OSR. These have bald tops with hi lis summit a white sand ringed by scrubby Oak woods, and rocky with Post sides These rich prairie-like pockets. unusual lulls host species with distinctly xenc, pe< ie: n .trictcd to e\n\ in< In.i \rg< inane alhiflora, ' li l bra, Croptilon ngulijaliuin, and Mirabihs Rivina linearis. mmit one of of these Species hills. only seen on the rocky hillsides include Loejlmgia and Opuntia saiairrosd The wa humifusa. lath ivcred with tin cale insect U loinopi .occoidea.e) r ra:(. ( from which pigment The the red cochineal isextracted. grassy areas of the hill- side were the only Dalca eampaila conn sites lor in lie v t t In the northern portion o( the county are two creeks that drain into the Cobb Youngs and down Trinity River, ireek Creek. These creeks have cut into ( the ancient terrace sand, and mudstonc deep ol clay, to iorrn canyons, creating a landscape unlike thai in any other part of Madison County. lowever, the two \ creeks have very different histories, and this is reflected in their floras. Youngs Canyon Creek formed within 70 the past years as the result an excavation by o'i the owners an attempt and in ro divert the creek reduce looding of the sur- I rounding cropland (Leon Wakefield, com The pers. L997). easily eroded in. soil was washed down quickly to the ironstone bedrock, creating sculpted walls more than seven meters high, bizarre monolithic pinnacles of sandstone, and The uncommon waterfalls. surrounding land exhibits the Southern Red Oak - Gum Oak Sweet - Post association, but the canyon itself has not yet developed a unique Only twospeeies, llydnnvt urnhelhaa and flora. Najasguadalupensis, vie bund were solely Youngs Creek Canyon. Future monitoring at of species re- cruitment canyon at this easily accessible could provide an interesting study ol colonization in a situation of unusual disturbance. The Cobb uplands around Creek are similar to those around Youngs Creek, maze but the of spring-led gullies and canyons wees created without human interference. This locality supports a flora that is allied with that of northeast- ern Texas and has produced cveral peck range extensions. Species found only along springs and low banksol CohhCreekme Apiasamentana,Araliaspinosa, Carex emoryi, Onoilea Quotas shumardir num and sensihilis, Tr Utile walteri, WaodwardiaarealaiaOne very unusual thiscanvon wasa witha habitat at site hanging garden on the canyon wal was continued some on that in aspects the I The woods cliff above. on the were mostly composed Quercus cliff of marilandica. Species found nowhere in the county other than this include site AreolariagrandiflorajLchiniiccaangustijoliajMLtucLi hirsuta,Liatriselegans, Osmunda cinnamomea, and Spiranthes tuherosa. Other species unique to

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