The types of Recent and certain fossil opisthobranch molluscs in the Muséum national d'Histoire naturelle, Paris

48 Pages·1998·17.8 MB·English

by  Héros, Virginie

The types of Recent and certain fossil
opisthobranch molluscs in the Muséum
national d'Histoire naturelle, Paris
Ángel VALDÉS & Virginie HÉROS
Laboratoire de Biologie des Invertébrés Marins et Malacologie,
Muséum national d'Histoire naturelle,
55 rue de Button, F-75231 Paris cedex 05 (France)
[email protected]
[email protected]
Valdés A. & Héros V. 1998. — The types of Recent and certain fossil opisthobranch mol
luscs in the Muséum national d'Histoire naturelle, Paris. Zoosystema20 (4): 695-742.
ABSTRACT
Three hundred and fifty seven lots of Recent and certain fossil opisthobran
ch mollusc type-specimens deposited in the Laboratoire des Invertébrés
Marins et Malacogie of the Muséum national d'Histoire naturelle (MNHN)
are catalogued by original binomen and arranged alphabetically within fami
lies. Most of the fossil type specimens are housed in the Laboratoire de
KEYWORDS
type specimens, Paléontologie, and therefore are not included in this catalogue. The essential
opisthobranchs, bibliographical, geographical and taxonomic information is provided for each
Mollusca,
MNHN, Paris. taxon.
RÉSUMÉ
Les types actuels et de quelques fossiles de mollusques opisthobranches du Muséum
national d'Histoire naturelle, Paris. Trois cent cinquante-sept types d'opistho-
branches actuels et quelques fossiles déposés au Laboratoire de Biologie des
Invertébrés Marins et Malacologie du Muséum national d'Histoire naturelle
(MNHN) ont été identifiés et listés par ordre alphabétique d'espèces à l'inté
rieur de chaque famille. La plupart des types fossiles se trouvent au
MOTS CLES
types, Laboratoire de Paléontologie et ne sont pas mentionnés dans cette liste. Les
opisthobrancnes, principales références bibliographiques, géographiques et taxonomiques
Mollusca,
accompagnent chaque potte-nom.
MNHN, Paris.
1998 -20 (4) 695
Valdés Á. & Héros V.
INTRODUCTION whose types ate now in MNHN, and new océa
nographie expeditions [Cap Horn, Travailleur et
In recent years, opisthobranch systematists have Talisman, Diguet expedition to Baja California,
become increasingly interested in examining type French Antarctic expeditions, Gravier Expedition
material as part of their research. However, the to Djibouti) were an additional source of type
scarcity of updated museum opisthobranch type specimens, mostly described by A. Locard,
catalogues make this task arduous and time P. Fischer, J. B. M. Vayssière and J. Risbec.
consuming. Most museum opisthobranch type During the First and Second World Wars, unlike
collections consist of historically important speci several natural histoty museums in Europe, the
mens for which the identification of type speci MNHN collections remained untouched. In
mens, and the determination of their status, is these and in the following years, many type spe
usually difficult to resolve with certainty. Often, cimens were added as a result of the work of
old type specimens remain unrecognized in the A. Pruvot-Fol who described material collected
general collections, which may house types of abroad and from the French coasts. However the
species not expected to be present at that types of many species that she described could
museum. never be found, possibly because the specimens
The objective of this paper is to document the were discarded after dissection; the extant mate
present collection and scope of the Recent and rial is mostly that of formal expeditions ot
certain fossil opisthobranch name-bearing type museum holdings that were entrusted to her for
specimens in the Museum national d'Histoire description. Material from the Calypso expedi
naturelle of Paris (MNHN). This will facilitate tions was also described by Ev. Marcus. In the late
future systematic research based on this material. 1950s, the Fischer family shared the property of
Journal de Conchyliologie. E. Fischer-Piette, then
professor and director of the Laboratoire de
A BRIEF HISTORY OF THE COLLECTIONS Malacologie, donated the type collection of the
The Museum national d'Histoire naturelle in Journal, containing a number of opisthobranch
Paris (MNHN) was founded in 1793, and short rypes. At that time, the Muséum wet collections
ly thereafter, the first opisthobranch type speci were stored in high glass containers, at the
mens were deposited in its collections. The ear- Galerie de Zoologie, the main exhibit building.
liest major collections extant are those of the When this gallery was closed to the public in
French expeditions to exotic countries, carried 1965 the collections remained there, and many
out during the early nineteenth century specimens went dry, or were otherwise damaged.
{['Astrolabe, 1' Uranie, Savigny expedition to A new age in the MNHN opisthobranch collec
Egypt, d'Orbigny expedition to South America, tion, characterized by the internationalization of
Peron expeditions to the Indo-Pacific), described the scope of the type collection, started with the
by the French classic authors: G. Cuvier, arrival of P. Bouchet on the staff in 1975. He not
J. C. Quoy & J. P. Gaimard, J. V. Audouin, A. de only added his own types, but also was instru
Ferussac, P. S. Rang, A. d'Orbigny, L. F. A. mental in encouraging authors from other coun
Souleyet. tries to deposit their type specimens in MNHN.
In the second half of the nineteenth century, and Many significant types of European and African
the beginning of the twentieth century, the species were contributed through the Spanish
French carried out a significant collecting effort school (J. A. Ortea, J. C. García-Gómez, and
in their colonies in the Caribbean, Africa, Eastern others), and Indo-Pacific material collected by
Asia and South-West Pacific. This material, French scientists were described by W. B.
added to MNHN collections, was also mainly Rudman (some of them still on extended loan).
described by French authors (G. Deshayes, In 1985, the MNHN scientific collections were
L. Morlet, A. T. de Rochebrune, P. Fischer). At moved to a new, large underground storage buil
about the same time, L. Morlet and H. Crosse ding, and the type specimens of Recent and cer
described several opisthobranch fossil species tain fossil molluscs were separated and placed in
696 ZOOSYSTEM A • 1998 - 20(4)
Opisthobranch mollusc type specimens in MNHN
a repository in the Laboratoire de Biologie des species-group names, they are listed separately,
Invertébrés Marins et Malacologie. However, the unless if they are identically spelled. The essential
opisthobranch collection still remained in need taxonomic, geographical and bibliographical
of curation for several years. The older type spe information is provided for each species-group
cimens were stored together with other material name. These data include the name of the species
in antiquated fashion, and no longer suitable cited exactly as published in the original descrip
containers, and it was difficult to find specific tion, the author, date and bibliographic reference,
lots. In fact, there are some examples in the recent the type locality as originally printed and its
literature of type specimens considered lost modern geographical equivalent, the category
(Wàgele 1985, 1990) that have recently appea and number of specimens, state of conservation,
red. For this reason, the first author of this paper accessory material (e.g. photographs, microscopy
was kindly invited in 1995 to work at MNHN as slides, spawn or food of the specimens, paratypes
an assistant curator, to reorganize the opistho and paralectotypes), and the name of the collec
branch collection, and separate the type speci tor when it is known. The old geographic names,
mens. This task, carried out in collaboration with no longer used, are printed between quotation
the second author, was concluded during another marks. The names of large geographic areas,
short-term stay in 1997, with the final separation countries or archipelagos are translated into
and inventory of all type specimens. During this English, but other names are printed in their
work, we have also found several types of species local spelling. The state of conservation includes
described by authors whose type material was not two main items: "specimen", which refers to spe
expected to be in MNHN. Some of them are cimens conserved in alcohol complete of soft
replacement names, names introduced under the parts, and "shell", when only a shell remain in the
Article 70c (ICZN 1985), or specimens appa dry collection. When the type specimen held is
rently borrowed by A. Vayssière from other insti also known to be originally or subsequently figu
tutions, never returned and later deposited at red it is indicated. All shells (with the exception
MNHN. of four nominal species catalogued while the
paper was already in press) are illustrated, even if
they have been previously figured. Figured speci
TYPE CATALOGUE mens are individually identified in the collection.
The catalogue of Recent and certain fossil types Short remarks with the currently accepted taxo
of opisthobranch molluscs in MNHN covers nomic status of the species, lectotype designa
three hundred and fifty-three lots of name-bea tions, status as type species of genus-group
ring type specimens. However, it does not inclu names, or any other useful information are given.
de most of the fossil species in MNHN which are No attempt has been made to list the synonyms
stored in the Laboratoire de Paleontologie (e.g. of each species, but opinions on the taxonomic
the Cossmann collection). status of several names have been included (only
This catalogue is restricted to name-bearing when they are based in the examination of the
types: holotypes, lectotypes, neotypes and syn- type material or are derived from an exhaustive
types. Other type lots (paratypes and paralecto- revision). All names have been grouped in fami
types) are in (or will be transferred to) the gene lies on the basis of available literature. When it
ral collection, and are only included in the cata was not possible we have been obliged to adopt
logue as accessory material of the name-bearing our own view, which obviously requires confir
types. All type specimens are listed by species- mation by further studies. This is not a critical
group names and arranged alphabetically within review of the material, but merely is an attempt
families. To facilite the search, the complete list of to make its usage easier.
names, arranged alphabetically, indicating the The families are enumerated in the order of the
family, higher category taxa and page number classification followed by Rudman & Willan
where each name has been placed is included. (1998), with nomenclatural adjustments
When the same lot is the type of two different (Bouchet & Rocroi, pers. comm.).
ZOOSYSTEMA • 1998 -20(4) 697
Valdes A. & Heros V.
CEPHALASPIDEA Family RINGICUI.IDAE Philippi, 1853
Family ACTEONIDAE d'Orbigny, 1842 admirabilis. Ringicula admirabilis Morlet, 1883:
203, 204, pi. 9, fig. 3. Type locality: Mediter­
azoricus. Actaeon azoricus Locard, 1897: 85-87,
ranean Sea. SYNTYPE: 1 shell, leg. Morlet
pi. 3, figs 8-11. Type locality: Talisman (1883)
(Fig. IE). Remarks: This is the type species of
Expedition, stn 126, 38°37'N - 28°21'W, bet­
the genus Plicatra Nordsieck, 1972, by original
ween Pico and S. Jorge, Azores, 1258 m.
designation. According to Ciccone & Savona
HOLOTYPE (by monotypy, figured by
(1982), this species is a junior synonym of
Locard, 1897, pi. 3, figs 8-11): shell (Fig. 1A).
Ringicula conformis Monterosato, 1877.
Remarks: This is the type species of the genus
bourgeoisi. Ringicula bourgeoisi Morlet, 1878b:
Inopinodon Bouchet, 1975, by original desi­
261, 262, pi. 8, fig. 5. Type locality: Pontlevoy,
gnation.
Paulmy, Ferrieres FAnjon, Manthelan (France,
maltzani. Actaeon maltzani Dautzenberg, 1910:
middle Miocene), Mandillot (France, early
10, pi. 1, figs 1, 2. Type locality: Baie de
Miocene). SYNTYPES: 3 shells, leg. Crosse
Rufisque and Goree, Senegal. SYNTYPE
(Fig. IF).
(figured by Dautzenberg, 1910: pi. 1, figs 1,
bourguignati. Ringicula bourguignati de
2): 1 shell (Fig. IB). Remarks: Dautzenberg
Rochebrune, 1883: 178. Type locality:
(1910) placed Tornatella senegalensis Petit de la
Casamance, Senegal. SYNTYPES: 2 shells
Saussaye, 1851 in the genus Acteon de
(Fig. 1G).
Montfort, 1810 (cited as Actaeon) and therefo­
caledonica. Ringicula caledonica Morlet, 1880:
re Actaeon senegalensis Maltzan, 1885 becomes a
154, 155, pi. 5, fig. 1. Type locality: "Plage de
junior secondary homonym of A. senegalensis
la Baie Pouen" (= Baie de Pritzbuer), New
(Petit de la Saussaye, 1851). Therefore,
Caledonia. SYNTYPES: 14 shells, leg. Morlet
Dautzenberg (1910) introduced the replace­
(Fig. 1H).
ment name Actaeon maltzani for it. This new
crossei. Ringicula crossei Morlet, 1878b: 273,
species was based on Maltzan's material and
274, pi. 7, fig. 11. Type locality: various locali­
newly collected specimens studied by Dautzen­
ties in central and western Europe (middle and
berg. All of them are synrypes of this species.
upper Miocene). SYNTYPE: 1 shell, leg.
morelletorum. Actaeon morelletorum Gougerot et
Crosse (Fig. II).
Braillon, 1968: 200, pi. 1, fig. 3. Type locality:
gaudryana. Ringicula gaudryana Morler, 1878b:
Barisseuse, France (middle Eocene). HOLO­
283, 284, pi. 7, fig. 12. Type locality: various
TYPE (by original designation, figured by
localities in central and western Europe (mid­
Gougerot & Braillon pi. 1, fig. 3).
dle Miocene to Pliocene). SYNTYPE: 1 shell,
vagabunda. Tornatella vagabunda Mabille, 1885:
leg. Crosse (Fig. 1J).
208. Type locality: Magallanes, Chile. SYN­
mariei. Ringicula mariei Morlet, 1880: 152, 153.
TYPE: 1 shell (Fig. 1C).
Type locality: "Nossibe" (= Nosy Be), Mada­
gascar. SYNTYPES: 9 shells, leg. de Folin
Family APLUSTRIDAE Gray, 1847
(Fig. 2A).
[= HYDATINIDAE Pilsbry, 1895]
minutula. Ringicula minutula Locard, 1897: 89,
eximia. Bulla eximia Deshayes, 1863: 55, 56, 90, pi. 14, figs 7-9. Type locality: Talisman
pi. 7, figs 23, 24. Type locality: La Reunion (1883) Expedition, stn 84, 22°54'N -
(Indian Ocean). SYNTYPES: 3 shells (Fig. ID). 17°26'W, off Sahara, 860 m. SYNTYPE:
guamensis. Bullaea guamensis Quoy et Gaimard, 1 shell (Fig. 2B).
1824: 423-425, pi. 66, figs 10-12. Type locali­ noumeensis. Ringicula noumeensis Morlet, 1880:
ty: Agana, Guam (Pacific Ocean). SYNTYPE: 155, 156, pi. 5, fig. 3. Type locality: Noumea,
1 specimen, leg. Quoy and Gaimard. New Caledonia. SYNTYPES: 18 shells, leg.
Remarks: Pruvot-Fol (1934b) placed this spe­ Morlet (Fig. 2C).
cies in the genus Micromelo Pilsbry, 1894. oehlertiae. Ringicula oehlertiae Morlet, 1880:
698 ZOOSYSTEMA • 1998 -20(4)
Opisthobranch mollusc type specimens in MNHN
FIG. 1. — A, holotype of Actaeon azoricus, 11.3 mm; B, syntype of Actaeon maltzani, 3.1 mm; C, syntype of Tornatella vagabunda,
8.2 mm; D, syntype of Bulla eximia, 10.2 mm; E, syntype of Ringicula admirabais, 3.6 mm; F, syntype of Ringicula bourgeoisi, 3.9 mm;
G, syntype of Ringicula bourguignati, 4.0 mm; H, syntype of Ringicula caledonica, 1.9 mm; I, syntype of Ringicula crossei, 3.7 mm;
J, syntype of Ringicula gaudryana, 7.2 mm.
ZOOSYSTEM A • 1998 - 20(4) 699
Valdes Ä. & Heros V.
156, 157, pi. 5, fig. 4. Type locality: Seas of savignyi. Ringicula savignyi Morlet, 1878a: 117,
China and Japan. SYNTYPES: 3 shells, leg. 118, pi. 5, fig. 1. Type locality: Suez, Egypt
Morlet and Saint-John (Fig. 2D). (Red Sea). SYNTYPES (one figured by
passiert. Ringicula passiert Morlet, 1880: 157, Bouchet & Danrigal 1982, fig. 59): 10 shells,
158, pi. 5, fig. 5. Type locality: Cap Breton leg. Savigny (Fig. 3C).
Canyon, Bay of Biscay SYNTYPE: 1 shell, leg. schlumbergeri. Ringicula schlumbergeri Morlet,
de Folin (Fig. 2E). 1883: 204-206, pi. 9, fig. 4. Type locality:
paulucciae. Ringicula paulucciae Morlet, 1878b: Mediterranean Sea. SYNTYPES: 3 shells, leg.
266, 267, pi. 6, fig. 6, pi. 8, fig. 9. Type locali­ Morlet (Fig. 3D). Remarks: According to
ty: Saucats and Dax, France (early Miocene). Ciccone & Savona (1982), this is a junior
SYNTYPE: 1 shell, leg. Crosse (Fig. 2F). synonym of Ringicula conformis Monterosato,
pirulina. Ringiculapirulina Locard, 1897: 87, 88, 1877.
pi. 14, figs 1-6. Type locality: Talisman (1883) senegalensis. Ringicula senegalensis Morlet, 1883:
Expedition, stn 33, 32°31'N - 09°48'W, off 202, 203, pi. 9, fig. 2. Type locality: Senegal.
Atlantic coast of Morocco, 1350 m. SYN­ SYNTYPES: 2 shells, leg. de Folin (Fig. 3E).
TYPES: 2 shells (Fig. 2G). Remarks: terquemi. Ringicula terquemi Morlet, 1880: 159,
According to Bouchet (1975b), this is a junior 160, pi. 5, fig. 7. Type locality: "Baie de
synonym of Ringicula nitida Verrill, 1874. Smyrne" (= Bay of Izmir), Turkey. SYN­
ponteleviensis. Ringicula ponteleviensis Morlet, TYPES: 5 shells, leg. Terquem (Fig. 3F).
1878b: 274, 275, pi. 8, fig. 8. Type locality: Remarks: According to Ciccone & Savona
various localities in France (middle Miocene). (1982), this is a junior synonym of Ringicula
SYNTYPE: 1 shell, leg. Crosse (Fig. 2H). auriculata (Menard de la Groye, 1811).
pulchella. Ringicula pulchella Morlet [ex Jeffreys toumoueri. Ringicula tournoueri Morlet, 1878b:
MS], 1880: 158, pi. 5, fig. 6. Type locality: 287, 288, pi. 6, fig. 10. Type locality: various
Atlantic European. SYNTYPES: 4 shells localities in central and western Europe
(Fig. 21). Remarks: This is the type species of (Neogene). SYNTYPE: 1 shell, leg. Crosse
the subgenus Ringactaeon Notdsieck (1972) by (Fig. 3G).
original designation.
quadriplicata. Ringicula quadriplicata Morlet,
Family CYLICHNIDAE H. etA. Adams, 1854
1878b: 286, 287, pi. 7, fig. 1. Type locality:
[= TORNATINIDAE P. Fischer, 1883]
several localities in Europe (middle and uppet
Miocene, and lower Pliocene). SYNTYPE: abyssicola. Mamillocylichna abyssicola Bouchet,
1 shell, leg. Crosse (Fig. 3A). 1975b: 349, 350, fig. 14. Type locality:
rosildae. Ringicula rosildae Rosso et Saubade, Biagores Expedition, stn 251,
1985: 305, 306, figs 8-11. Type locality: 47°38'N - 08°56'W, Banc de la Chapelle,
14°36'30"N - 17°15'30"W, off Senegal. 3360-3600 m. HOLOTYPE (by original desi­
HOLOTYPE: said to be deposited in MNHN gnation): shell (Fig. 4F).
in the original publication, but never presented crossei. Cylichna crossei Bucquoy, Dautzenberg et
to the museum. Dollfus, 1886: 526, 527, pi. 64, figs 9-11.
roussellae. Ringicula roussellae Rosso et Saubade, Type locality: not specified; cited from Canet,
1985: 303, 304, fig. 2. Type locality: France, Viareggio, Italy and Alger. SYNTYPES
14°22'40"N - 17°09'W, off Senegal. HOLO­ (One figured by Bucquoy, Dautzenberg &
TYPE: said to be deposited in MNHN in the Dollfus 1886: 526: pi. 64, fig. 9): 2 shells from
original publication, but never presented to the Alger.
museum. imperceptus. Meloscaphander imperceptus
salleana. Ringicula salleana Morlet, 1880: 153, Bouchet, 1975b: 341-343, figs 9, 10, pi. 3,
154. Type locality: Cap Breton Canyon, Bay of figs G-H. Type locality: Talisman (1883)
Biscay. SYNTYPE (probably): 1 shell, leg. de Expedition, off Mauritania. HOLOTYPE (by
Folin (Fig. 3B). original designation, figured by Bouchet,
700 ZOOSYSTEMA • 1998 -20(4)
Opisthobranch mollusc type specimens in MNHN
FIG. 2. — A, syntype of Ringicula mariei, 1.7 mm; B, syntype of Ringicula minutula, 2.9 mm; C, syntype of Ringicula noumeensis,
3.7 mm; D, syntype of Ringicula oehlertiae, 3.9 mm; E, syntype of Ringicula passieri, 4.2 mm; F, syntype of Ringicula paulucciae,
3.4 mm; G, syntype of Ringicula pirulina, 6.8 mm; H, syntype of Ringicula ponteleviensis, 3.2 mm; I, syntype of Ringicula pulchella,
2.7 mm.
ZOOSYSTEMA • 1998 -20(4) 701
Valdes A. & Heros V.
1975b, pi. 3, fig. H): shell (Fig. 4G) + 1 para- Travailleur (1880) Expedition, drag. 2,
43°46'N - 01°55'W, Cantabrian Sea, North of
insperata. Bulla insperata P. Fischer in Locard Spain, 1019 m. SYNTYPE: 1 shell (Fig. 5F).
1897: 54, 55, pi. 1, figs 19-22. Type locality: scaphandroides. Roxania scaphandroides Staadt
Talisman (1883) Expedition, stn 76, 25°01'N - in Cossmann & Pissarro 1913, pi. 55,
16°55'W, off Sahara, 2638 m. SYNTYPES: fig. 242-12. Type locality: Chenay, Ftance (late
2 specimens (1 of them dissected). Remarks: Paleocene). HOLOTYPE (by monotypy, figu­
Accotding to Bouchet (1975b), this is a junior red by Cossmann & Pissarro 1913, pi. 55,
subjective synonym of Scaphander mundus fig. 242-12): shell (Fig. 5C).
Watson, 1886. villersii. Bulla villersii Audouin, 1826: 39. Type
lemchei. Cylichna lemchei Bouchet et Waren, locality: Egypt. SYNTYPES (one figured by
1979: 228, 229, figs 16, 18G, H, J-L. Type Savigny, 1817, pi. 5, fig. 4 and Bouchet &
locality: Norbi Expedition, stn CP02 Danrigal 1982, fig. 53): 2 shells (Fig. 5D).
(64°26'-64019'N, 0T°36'-0r44'E) Norwegian voluta. Bulla voluta Quoy et Gaimard, 1833:
Sea. HOLOTYPE (by original designation, 359, 360, pi. 26, figs 33-35. Type locality:
figured by Bouchet & Waren 1979, fig. 18J-L): Guam (Pacific Ocean). SYNTYPE: 1 shell, leg.
shell (Fig. 4H). Quoy and Gaimard (Fig. 5G). Remarks:
millepunctata. Bulla millepunctata Locard, Pruvot-Fol (1934b) placed this species in the
1897: 52-54, pi. 2, figs 3-6. Type locality: not genus Acteocina J. E. Gray, 1847.
specified; cited from Talisman (1883)
Expedition, several stations, off Sahara,
Family RETUSIDAE Thiele, 1925
Senegal and Azores. SYNTYPES: 1 specimen
and 2 shells (Fig. 41). Remarks: This is the canariensis. Cylichnina canariensis Nordsieck et
type species of the subgenus Bullocardia Garcia-Talavera, 1979: 178, pi. 44, fig. 46.
Nordsieck, 1972, by original designation. Type locality: Tenerife, Canary Islands.
According to Bouchet (1975b), this is a junior SYNTYPE: 1 shell (Fig. 5H).
subjective synonym of Scaphander nobilis caudidida. Cylichna candidula Locard, 1892: 28.
Verrill, 1884. Type locality: Atlantic, coast of Ftance. SYN­
mirabilis. Tornatina mirabilis Locard, 1897: 72, TYPE: 1 shell (Fig. 51).
73, pi. 2, figs 20-24. Type locality: Travailleur desgenettii. Bulla desgenettii Audouin, 1826: 39.
(1882) Expedition, stn 23, 38°21'N - Type locality: Egypt. SYNTYPES (one figured
09°27'W, off Portugal, 2000 m. SYNTYPE: by Savigny 1817, pi. 5, fig. 6 and Bouchet &
1 shell (Fig. 5A). Danrigal 1982, fig. 55): 5 shells (Fig. 5J).
morelletorum. Cylichna morelletorum Gougerot dilatata. Retusa dilatata Pallary, 1904: 215, 216,
etLe Renard, 1983: 82, 85, fig. 12. Type loca­ pi. 7, fig. 8. Type locality: Gulf of Gabes,
lity: Ferme de l'Orme, France (middle Tunisia. SYNTYPES: 11 shells (Fig. 5K).
Eocene). SYNTYPES: 24 shells. Remarks: Pallary (1904) selected a 6 mm long
olivaeformis. Tornatina olivaeformis Issel, 1869: shell as the "type" of this species. However, in
171, 172. Type locality: Gulf of Suez, Egypt MNHN collection thete are two vials with 4
(Red Sea). SYNTYPE (figured by Savigny and 7 shells respectively, all of them similar in
1817, pi. 6, fig. 25 and Bouchet & Danrigal length (about 6 mm), which therefore are
1982, fig. 56): 1 shell (Fig. 5E). considered syntypes.
petiti. Cylichnapetiti Dautzenberg, 1923: 70, 71, fourierii. Bulla fourierii Audouin, 1826: 39.
fig. text. Type locality: "Fenerive" (= Fenoarivo Type locality: Egypt. HOLOTYPE (by mono­
Atsinanana), Madagascar. HOLOTYPE (by typy, figured by Savigny, 1817, pi. 5, fig. 5 and
monotypy, figured by Dautzenberg, 1923, fig. Bouchet & Danrigal 1982, fig. 54): shell
text): shell (Fig. 5B). (Fig.5L).
pusillina. Tornatina pusillina Locatd, 1897: girardi. Bulla girardi Audouin, 1826: 39. Type
75-77, pi. 2, figs 29, 30. Type locality: locality: Egypt. HOLOTYPE (by monotypy,
702 ZOOSYSTEMA • 1998 - 20 (4)
Opisrhobranch mollusc type specimens in MNHN
FIG. 3. — A, syntype of Ringicula quadriplicata, 8.9 mm; B, probable syntype of Ringicula salleana, 5.0 mm; C, syntype of Ringicula
savignyi, 2.8 mm; D, syntype of Ringicula schlumbergeri, 3.1 mm; E, syntype of Ringicula senegalensis, 2.3 mm; F, syntype of
Ringicula terquemi, 1.8 mm; G, syntype of Ringicula tournoueri, 4.3 mm; H, syntype of Bulla australis de Ferussac, 41.6 mm;
I, syntype of Bulla australis Quoy et Gaimard, 49.4 mm.
ZOOSYSTEMA • 1998 -20(4) 703
Valdés Á. & Héros V.
FIG. 4. — A, holotype of Bulla mongli, 1.4 mm; В, syntype of Bulla orbignyana, 16.1 mm; C, syntype of Bulla ovoidea, 13.6 mm;
D, syntype of Haminea perrieri, 16.4 mm; E, holotype of Cylichnium waldae, 28.1 mm; F, holotype of Mamillocylichna abyssicola,
2.8 mm; G, holotype of Meloscaphander imperceptus, 19.3 mm; H, holotype of Cylichna lemchei, 7.1 mm; I, syntype of Bulla mille-
punctata, 41.0 mm.
704 ZOOSYSTEM A • 1998 - 20 (4)