The Trechine Beetles (Coleoptera, Trechinae) from Mt. Wawu Shan in Central Sichuan, Southwest China

2006·12.4 MB·

by  Ueno, S-I

Ell,t1・a、T◆ok、・(),34(1):l3-32、May20,2006
TheTrechineBeetles(Coleoptera,Trechinae) fromMt.WawuShanin
CentralSichuan,SouthwestChina
Shun-lobi UE_No
Departmento、fZoology,NationalScienceMuseum(Nat.Hist.)
3 23-1Hyakunin-cho,Shinjuku,Tokyo,169-0073Japan
Abstract Fournewspeciesof trechinc beetlesarc described from Mt. Wawu
ShanincentralSichuan,SouthwestChina.Twoo、fthembelongtothegcnusEpap/1lopsls;
onenamed E inuit・alisamemberofthesubgenusPse1lc/_paphitlsandhasnocloserela-
tivesinChina;theother namedE. (/ao,belongstothe,11bagroupofthesubgcnusEpa-
p/11a,na.0ncoftheremaining、t、,oisamemberofSill()tl・ec/11a,IlabutresemblesT1,・ec/11a111a
c・,asst/)osatthe rstglance.ItisnamedSI,1011・ec1/1a111(リ)al・、1,1sinviewofitssmallsizeand
thelatterspeciesismovedfi-omthegenusTli・ec1/iamatothegenusP・lof,・ec/11ama.Thelast
newspeciesisaneyelessendogcantrechinebelongingtoanewgenus.which resembles
Aeplb/e川, l、.,andDu、,a/1ob/elmlsinmanyrespects,butcanberecognizedonthespotbythe
presenceofaseti、lerousdorsalporeonthe5thelytra1striaandthepeculiarmodificationof
striation.ThenewnamegivenisLc1ob/e1mlscl、,pt1(,,fis.
WawuShanisafamoustablemountainlyingonthebordersofHong'yaXianand
YingjingXianin central Sichuan,SouthwestChina. Itstandson theDaxiangLing
Mountains, and isabout 40km distant to the west-northwest in a beeline from Mt.
cruelShan.Thetablemountainissurroundedbyverticalcliffsmorethan t,000mhigh
attheeasternandsouthernsidesandextremelysteepslopesattheothersides.Itstable
topis formedattheelevationof2,700to2,800m,andiscoveredwithAbles forest
withthickundergrowthsofarrow-bamboosandvariouskindsofRhododendrontrees.
Mt.WawuShanisprotectedasanationalnaturereserve,andisregardedbyTaoistsas
asacredplace,sinceLaozi(orLaotzu),thefounderofTaoism,issaidtohavespenta
secludedlifeonthetabletopof thismountainandgoneup to theheavenridinga
crane
It isnoteasytomakesatisfactoryco11ectingsonthetabletopofWawuShan,
sjncethefloorsoftheAblesforestsarethicklycoveredwithentangledrootsofa「「oW-
bamboosanddeadleaves.Itispossibletomakesiftingsinthebamboothickets,pa「tiC-
ularlyunderRhododend1-o1, trees,butbeetleslivingunderstonesonclayey9「ound,en-
dogeanformsjnparticular,cannotbeexpectedinsuchanenvironment.Ba「e9「oundS
arefoundonlyontrajlsandonthebanksofnarrowstreams,butallthet「ailsa「eti9htly
pavedwithslatesca 30x l00cm insize,whichcannotbereadily replacedto thei「
orlgjna1posjtjonwhenturnedover.BanksofnarrowstreamsareusuallyverySteepo「
vertjca1,andveryfrequentlyrocky,sothattheyarenotfavourablefOr1oOkin9fo「t「e-
14 Shun-lobiUI_No
chinebeetles.Undersuchcircumstances,wewerecompelledtoexaminethegrounds
beneaththepavement,andmovedmorethan200slates. Thiswasapainstakingand
time-consumingwayoftakingraretrechines,but fourspecimensofthreenewspecies
tobedescribedinthepresentpaperwerefoundoutbythismethod.
Ofthefournewspeciestobedealtwith,twobelongtotwodifferentsubgeneraof
thegenusEpaphiopsls.Theyareprimarilyhumico1ousinnature,butatenera1speci-
menofthelargerspeciesofthetwowasfoundinacellmadeofyellowclaybeneatha
slate.Aswillbenotedlater,thisspecies,tobenamedE.11mlla1,seemstobeausual
inhabitantoflower placesthanthetabletop. 0neof theremainingtwospecies isa
memberofthegenusSinot1-echlama;twospecimensofthisspecieswere found from
beneathslatesonthetrail,andtheotheronewastakenbysiftingdeadleavesaccumu-
latedatthesideofthetrail.Thelastspeciestobedescribedisananophthalmicendo-
geanspecies,alsofoundfrombeneathaslate.Wehavevisitedthetablemountainfour
times, intheearlysummerandtheautumn,andcarefullysearched foradditionalspeci-
mensofthenon-humico1ousspecies,aboveallmalesoftheendogeanone. 0urefforts
were,however,notrepaidwithanyofthethreespecies.Sincethemountainisnoteasy
ofaccess, Ihavedecidedtointroduceallthesenewspeciesintoscienceinthepresent
paper.Theabbreviationsemployedhereinarethesameasthoseexplainedinprevious
papersofmine.
Beforegoinginto further details, I wishtoexpressmyheartfelt thankstoDrs.
YukiIMuRA,YoshiakiNIsHIKAwAandMasatakaSAT0 for theirkindhelpinsearching
fortrechinebeetlesonMt.WawuShananditsvicinities.HeartythanksareduetoDr.
IgorA.BELOUSOv,Dr.Ilia1.KABAKandIng.ArturGITzF_Nforgivingmeopportunities
toexaminethetypesofChinesetrechinesforcomparison.Deepindebtednessshould
alsobeexpressedtoMr.FANTingforhiskindarrangementinpursuingourinvestiga_
tions.
Epaphiopsis(Pseudepaphius)imuraiS.UENo,sp
nov.
(Figs.l-3)
Len9th:3.50-3.90mm(fromapicalmarginofclypeustoapicesofelytra).
Relativelylargespeciesnotdirectlyrelatedtoanyofthehithertodescrjbedchj_
noseSpeciesofPse1,ldePaPhiils(UENo, l962,p 70),externally resembling cerlajn
Japanesemembersofthesubgenus(e.g.,E
morimoto1・s. UENo (1g84,p l46, figs
1-3))thoughgenitalicaIlydissimilartothem.
BodyShe「tandbroad, withsmallboa transverseprothoraxand ovateconvex
elyt「a;aPte「ouS;mCi rosculpturefine,sharplymi pressedandcomposedoftransverse
meshesandlinesonhead,mostlyclearandformedbytransverseljnesonpronotum,
andIa「9elyobIite「atedonelytrathoughvestigeoffinetransverseljnescanbeobserved
he「eandthe「e・COnCo1orousIydarkreddishbrown,shiny,feeblyiridescentonelytraIn
theSin9lefullymaturespecmi en(holotype),withpalepaIpjandclawsegmentsoffore
Ie9S,middleandhindlegssomewhatlighterthandorsum.
TrechincBootiesfromMt.WawuShaninSouth、vcstChina
Fjg 1 jpap1/1ops1s(psellde/,ap/11111.,)川l ll・,alS.U1lN0,sp nov., fromtheShuangdongXiattheeastern
,
footofMt.WawuShan.
Headsmall,transverse,obviouslywiderthanlong,widestatthemid-eyelevela
1jttlebehjndmjddle,withdeepentire frontal furrowshardlyangulateatmiddlebut
wjdelydivergentinfrontandbehind;fronsandsupraorbitalareasgentlyconvex,the
latterbearjngtwopairofsupraorbitalsetae,theanteriorpairofwhichisfoveolateat
theroots;eyessmallandflat;genaeconvexthoughshort,evidentlyshorterthaneyes
(three_ejghthstofour-seventhsaslongaseyes);neckverywide,withdistinctanteriO「
constriction; labrumtransverse,slightlyemarginateat theapex;mandiblesstout, in-
16 Shun-lobi UI_No
wardlyhookedat theapices; palpi shortandthin; antennae fairlyslender, reaching
basaltwo-seventhsofelytrain basal fourthofelytrain , antennomeres2-7sub-
,
equal in length tooneanother, eachabout three-seventhsas wideas long, anten-
nomeres8-10slightlydecreasinginlengthtowardsterminalantennomere,whichisthe
longestthoughevidentlynarrowerthanscape.
Pronotum transverse, much wider than head, widest at about two-thirds from
base, and more gradually narrowed towards base than towards apex; PW/HW
1.49-1.60(M I 54),PW/PL139-1.42(M I41),PW/PA t 45-1.48(M I46),PW/PB
1.25-1.30 (M I 27);sides rather narrowly bordered throughout, especiallynarrowly
near frontangles,moderatelyarcuateinfront,very feeblysoornearlystraightbehind
middle,andnotsinuatebeforebase.withtwopairofmarginalsetae,theposteriorone
ofwhichisinsertedalmostonhindangles;apexnarrowerthanbase,PB/PA t.14-1.16
(M I.15),eithernearlystraightorslightlyarcuate,with frontanglesroundedandeither
hardlyorveryslightly advanced;baseeitherstraight or invisibly arcuateatmiddle,
brieflyandveryslightlyobliqueoneachsidejustinsidehindangle,whichisobtuse
but veryslightlyproduced; dorsumconvex,median linedeeplyimpressed, not reach-
ingapexbutextendingalmosttobase;apical transverseimpressionshallow,slightly
wrinkle basal transverse impressionnarrow but continuous, lightly arcuateas a
whole,andlaterallyextendingintobasal foveae,whicharedeclivousandfairlylarge;
nopostangularcarinae;basalareasmall,narrowandsmooth.
Elytraovate,wider thanpronotum, widestatabout four-ninths frombases,and
moregraduallynarrowedtowardsbasesthantowardsapices,withnarrowapicalparts;
EW/PW128-1.36(M I32),EL/PL2.57-2.64(M2.60),EL/EW137-1.43(M I40);
shoulderssquarethoughrounded,withprehumera1bordersnearlystraightandonlya
littleoblique;sidesmoderatelyreflexedinfrontbutnarrowlysobehindmiddle,nearly
straightbehindshoulders, feeblyarcuatebehindmiddle,andthennarrowlyandcon-
jointlyroundedatapices,eachwithnoappreciablepreapica1emargination; dorsum
stronglyconvex, particularly inapicalhalves,steeplydeclivousat lateral andapical
parts,andlightlyandnarrowlydepressedon thedisc,striaeclearlyimpressedand
rathercoarselypunctateonthediscbutbecomingshallowerat thesideandpartially
obliterated,striae1-3nearlyentire,4-5superficialandobsoletenearbase,6_7 frag_
mentaryandevanescent,8impressedonlyneartheapicalsetofmarginalumbilicate
Pores;scutellarstrictodistinct,apicalstrictoalsodistinct,hardlyarcuatein front,and
directedtothesiteofstria5;intervalsfiatevennearsuture,apicalcarinaobtuse;stria3
withtwosetiferousdorsalporesaft/8-I/7and1/3-3/7frombase,respectively;preapi_
Calporeusuallyadjoiningstria2af t/4-3/10fromapex;stria5orinterval5wjthasjn_
9leSetiferousdorsalporeat3/7-5/9 frombase;marginalseriesofumbilicatepores
regular.
Ventralsurfaceglabrousandsmooth;anal setaeordinary.Legs fairlyslender
thoughshort;protibiaestraight,lightlydilatedtowardsapices; tarsishort,tarsomere1
about as long astarsomeres2and3 combinedinmesotarsus, longer than that jn
metatarsus;in ,protarsomeresland2widelydilatedandstoutlyproducedinwardsat
TrcchincBeetlesfromMt.WawuShaninSouth、vcstChina 17
3
、
/
/ /
///一一 /
_
0.3”;”l
Figs 2-3. MalegenitaliaofEpap1/1opsls(Psetlc/epap1/1us)limi・talS.UENo.sp nov.. fromtheShuang-
dongXiattheeasternfootofMt.WawuShan:leftlateralview(2).andapicalpartofacdeagusdorso-
apica1view(3).
apices.
Malegenital organsmall thoughheavilysclerotizedLdifferinginconfiguration
fromthoseofanyotherknownChinesespeciesofthegenus.Aedeagusonlyone-third
aslongaselytra,gentlyarcuateasawholethoughnearlystraightforashortdistance
behindmiddle,moderatelydepressed,andwidelymembraneousondorsum;basalpart
fairlynarrowinprofile,lightlycurvedventrad,andmoderatelyemarginateatthesides
oftheposteriorpartofbasalorifice;sagittalaileronlargebutnotsowide;viewedlat-
erally,medianlobegraduallynarrowedfrombehindmiddletowardstheanteriorendof
apjca1orjfjce,thenweaklycurvedventradandratherabruptlyacuminatetopointedex-
tremity;vieweddorsally,apicalpartbroadattheleve1oftheanteriorendofapicalo「i-
fjce,narrowedposteriorly,andproducedintoashortapicallobebluntatthetip;Vent「al
marginslightlyemarginateatmiddel inprofile.InnersacdevoidofdifferentiatedCopu-
latorypiece,whollycoveredwithminutescalesandteeth,whicha「emode「atelySCle-
rot1zedalongtwo1ongjtudjna1foldsofsacmembrane formin9 twoeton9ateScale-
patchesoneabovetheotheraltheleftside;anteriormarginofapicalo「ifCieed9edWith
heavilysc1ero1lzedteeth,whichdelmi itthePosteriorendofaSmalldo「Salteeth-Patch
lyinghorizontallystylesunequalinsizeandshape,leftstylemuchta「9e「andb「cade「
thantherightandwithawelldevelopedventralapophysis,eachStylebea「in9Ofu「
apica ;a:,
, (boeth,fessom. eHwOhlaottytepnee:ra1),,2l9-_VXI_-22000044,,YY..NMIIusRHAIKAlew9A.Ale9ll.depAolslOitteydpei:nth,ePCao「al--
type:1
lectionoftheDepartmentofZoology,NationalScienceMuseum(Nat・Hist・),Tokyo・
18 Shun-lobiUENo
Typelocality. Shuangdong Xi,1,320m inaltitude,at theeastern foot ofMt.
WawuShaninWawushanZhenofHong'yaXian,centralSichuan,SouthwestChina.
tutthorspecimense.xlam1,7ed. 1 (tenera1),Mt.WawuShan,Esideoftabletop,
2,760malt.,Wawushan Zhen, Hong'yaXian,23-VI-2004,S. UENo leg. (NSMT).
1 FuxingCun,1,620malt.,WuzhuangXiang,Hong'yaXian,24-VI-2004,S.UENo
,
leg; 1 (teneraI and somewhat damaged), same locality and date, M. SAT01eg.
(NSMT).
Notes. Thisisaproblematicalspecieswhosetrueaffinity isnotcertainat the
presentmoment. Infacies,itcloselyresemblescertainJapanesespeciesofPseudepa-
phlus,buttheresemblancemayhaveresultedfromconvergenceseeingthatthepresent
speciesdiffersfromJapaneseonesinthemodeofelytra1striationandconformationof
themalegenitaIia. 0ntheotherhand,therestillremainsapossibilitythatE. lmu1・alisa
true relativeofJapanesemembers; inthatcase, thedifferencesmayhavedeveloped
duringlongisolation.Furtherinvestigationsareneededforclarifyingitstruestatus.
Unfortunately,onlytheholotypeofthisnewspeciesis fullymature,theremain-
ingfivebeingmoreorlesstenera1.Thespecimensofthetypeserieswerecollectedina
deciduousbroadleaved forestat theleftsideof theShuangdongXiRiverbysifting
deadleavesaccumulatedonarathersteepslope.Thelocalityisabout1,440mlowerin
altitudethanthatattheeasternsideofthetabletop.ThematuremalewastakeninOc-
tober,whilethetwotenera1femalesweretakeninJune.Judging fromthe factthatall
thespecimenscollectedinJuneatthetypelocalityandtwootherlocalitiesaremoreor
lesstenera1,emergenceofthistrechineseemstotakeplaceearlyinthesummer,even
thoughitismainlydistributedinmiddlealtitude.Aswasnotedintheintroductjonof
thispaper,theonlyspecimentakenonthetabletop,3.60mminthelengthofbody,
Wasfoundinasmallclayceilbeneathaslateonthetrail.Itagreeswellwiththetype
seriesthoughtheelytraaresomewhatbroader.Thestandardraljosare:pw/Hw
148,
PW/PL 146,PW/PA t46,PW/PB128, PB/PA1.16, EW/PW
143, EL/PL 2.80,
EL/EW133.
ThetwoSpecimens fromFuxingtotheeastofMt.WawuShanweresjftedout
f「omSmallheapsofdeadbambooleavesaccumulatedattheedgeofabamboogrove
nea「theSOu「CeOfanarrowstream.Theyareslightlylargerinsizethanthetypeseries
(4・00-4・15mminthelengthofbody),andaredifferent fromthelatterjnthesmaller
eyes,lesst「anSVe「SeP「othOraxwithmoreregularlyarcuatesides,aljttlelongerelytra,
andSoon・TheStandardratiosintheperfectspecmi enareasfollows:pw1Hw
153
PW/PL1・34,PW/PAt・57,PW/PB130,PB/PA l20,EW/pw133,EL/PL263,
EL/EW1・47・AnewSubspeciescouldberecognizedfortheFuxjngpopulation,espe_
Cia11yonthebasisofthedifferentconfigurationoftheprothorax,butlprefertopost_
PonetheP「oPOSa1untilfullymaturemalesareavailableformystudy
TrechincBodiesfromMt.WawuShaninSouth、、'cstChina l9
pap加'opsis(9pap/ll'a''Ia) aoS.UEN0,sp nov.
(Figs4-6)
Length: 2.75-3.15mm (from apical margin of clypeus to apices of elytra).
Belongjng to the川agroupOf thesubgenus ap/?Ia'Ilaandclosest to . 川フfa S.
UENo(l998,p 267,figs.1-3)knownfromMt.NibaShanontheDaxiangLingMoun-
tains,butdistinguishedfromitbybroader facies,widerhead,lesscontractedpronotum
atbothapexandbasewiththesideslessstronglyarcuate,andusuallywiderelytraless
pojntedattheapicesandwiththesideslessregularlyarcuateinbasal third.Alsodif-
ferent fromE nlbaintheconfigurationofmalegenitalia.
colourmoreor lessdarkerthanin c nlbt1,sometimesinfuscatedinheadandely-
tra,thelatterofwhicharehardlyiridescent.Microsculptureasinc nlba.
Headtransverse,obviouslywiderthanlong,widestatthemid-eyelevelwhichlies
aljttlebehjndmjddle:dorsumdepressed,withdeepentirefrontal furrowswidelydi-
vergentposteriad:fronsandsupraorbitalareasgentlyconvex;eyessmallandfla、talit-
tle1ongerthangenaein ,usuallyaslongasoralittleshorter thangenaein ; 9enae
moreorlessconvex,usuallytumidin ;neckverywide,withtheanteriorconstriction
deeplymarkedatthesides;labrummoderatelyemarginateattheapex,mandiblesstout
thoughsharplyhookedattheapices;palpishortandstout;antennaeshortandStout,
varjablejnlength,sometimesevenattheoppositesides,usuallyreachingbasalsixthto
fjfthofelytrain basal eighthto two-eleventhsofelytrain , antennome「oS2-4
subequa1inlengthto, oneanother,eachveryslightlylongerthansucceedingones,each
issubova1and four-seventhsaswideas long, terminal antennomere the
of whjch
longest,slightlylongerbutnarrowerthanscape.
pronotumtransverse, wider thanhead,wider thanlonginasimilar proportion,
wjdestatalevelbetweenthree-fifthsandtwo-thirds frombase,andmoreg「adually
contracted towards basethan towards apex; PW/HW 135-1.43 (M 1.39), PW/PL
l32_l45(MI38),pW/PA136-1.46(MI43),PW/PB132-1.43(MI37);Sides
moderatelyreflexedthroughout,gentlyarcuatebeforemiddle,lesssoinf「ontandbe-
hjn and hardly sinuatebeforehindangles,whichareobtuseandsometimesVe「y
slightlyproducedlaterad,withtwopairofmarginalsetae,thePosteriorPairofWhich
areverysljghtlyremovedforwards;apexeitherstraightorslightlyemarginate,usually
somewhatnarrower thanbase,PB/PA1.00-1.l l (MI.05),withf「ontanglesna「「oWly
roundedandsljghtlyproducedfo1wards:baseeitherstraightorveryslightlya「Cuateat
middleandsljghtlyemarginateoneachsideinsidehindangle;dorsumConvex,Steeply
decljvousatanterjorlatera1parts,medianlinedeeplyimpressedonthediscandbnefly
widenedjnbasalarea;apjca1transverseimpressionmat-defined,longitudinallyW「in-
k1ed;basaltransverseimpressionalsomat-define withalongitudinal OfVeoleoneach
sideofmedjanljne,andlaterallyarcuateposteriad;basal foveaefai「lyla「9eanddeep;
postangularcarjnaedistinctthoughobtuse;basalareamoreorlessuneven・
Elytrasubovate,widerthanpronotum,Ion9erthanwideinaSimila「P「ope「tOin,
widestatabout four_njnthsfrombases,weaklynarrowedante「ladtOWa「dSShOulde「S,
20 Shun-lobiUENo
Fig. 4. ipap1/1oりfsis( ap/11a,Ila)(/aoS.UENo,sp nov., from Mt.WawuShan
,
andwidelyroundedatapices,whichformasmallre-entrantangleatsuture;EW/PW
1.31-1.42 (M I34),EL/PL2.40-2.65(M2.52),EL/EW l29-1.43(M I36);shoul-
dersdistinctthoughrounded,withprehumera1bordersonlyslightlyoblique;sidesnar-
rowlyborderedthroughout,either very feeblyarcuateornearlystraightbeforemiddle
andmoderatelyarcuateinapicalthird,eachwithaslightpreapica1emargination;dor-
sum convex,steeplydeclivousat lateral andapical parts; striaenearlyentirethough
shalloweratthesidethanonthedisc,vaguelycrenulate,1-3deeplyimpressed,4and5
shallower than innerones,6and7 fineandoften fragmentary,8distinctbehindmid-
dle; scutellarstrictoshort butdistinct; apical strioleclearly impressed,moderately
T「CChineBeetlesfromMt.WawuShaninSouthwestchjna
21
6
、 ノ
、ーノ
0.3mm
Fi9S・5-6. Male9enitaliaofEpap/1lopsls(Epap/11a1nc1)ciaoS.UENo,spnov., tron、MI.wawushan;tefl
lateralview(5),andapicalpartofaedcagus,dorso-apicalview(6)
Cu「Vcd,and freeat theanteriorendthoughdirectedtostria5; intervalsflatevennear
Sutu「e,apicalcarinaobtuse;stria3withtwosetiferousdorsalporesat2/11_2/9(usu_
allyabout1/5)and2/5-1/2frombase,respectively;preapica1porelocatedattheapjca1
anastomosisofstriae2and3oradfoiningstria2behindtheleve1oftheanteriorendof
apicalstricto,andalittlemoredistant fromapexthanfromsuture;stria5withasingle
setiferousdorsalporeatabout3/5frombase,theporebeinglocatedoftenatananasto-
mosisofstriae4and5;marginalumbilicateporesaggregatedandregular.
Ventralsurfacesmooth;analsetaeordinary.Legsshortandstout;protibiaemod-
eratelydilatedtowardsapicesandshallowlygroovedoneachexternal face;tarsishort,
tarsomere 1shorter than tarsomeres 2and3 combined in both meso- andmetatarsi;
protarsomeres1and2widelydilatedandstoutlyproducedinwardsatapicesin(S.
Malegenital organsimilar inbasic conformationto that ofE nlba S. UENo
(1998,p 267,・figs.1-3),but theaedeagusismuchlessarcuate,especiallyinapical
half,compressedinapicalpart,andwithnarrowlyproducedapical lobe.Aedeagus
abouttwo-fifthsaslongaselytra,sigmoidallycurvedindorsalview,andstronglyarcu-
atejnproxjma1half,withlargebasalpartstronglycurvedventrad;basalpart「ounded,
wjthsmall basal orificewhosesidesarehardlyemarginate;sagittal aileronabsent;
vieweddorsally,apicalpartcurvedtotheright,andthentotheleftandP「educedintoa
shortnarrowapjca11obebluntattheextremity;viewedlaterally,apicallobeShe「t,not
curvedventrad,andabruptlynarrowedtowardspointedextremity;inprofile,Vent「al
margjndeeplyemarginatebeforemiddlebutnearlystraightbehindthe「e. Inne「Sac
armedwjthaverylongslendercopulatorypiecenearlyaslongasaedeaguS,Withthe
22 Shun-IchiUI・,No
basalpartgraduallydilatedproxmi adandthenarrowhyalineapicalportionP「educed
totherjghl fromapicalorifice;sacmembranewhollycoveredwithminuteScalesand
teeth, formjngalongsheathofcopulatorypiece.Stylesshortandnarrow, leftStyle
muchlongerthantheright,arcuate,anddevoidofventralapophysis,eachboa「ingfOu「
apicalsetae.
Typeseries. Holotype: ,allotype:?,centreoftabletop,23-VI-2004,S.UENo
leg paratypes:6 ,4 (incl twobadlydamagedspecimens(1d,1!)collectedby
SAT0), 23_VI-2004, S.UENo & M. SAT01eg; ld, 5 , NW Sideof table top,
23_VI_2004, S. UENo & M. SAT0 leg; 19, Eside of table top, 2,760m alt.,
23_vI_2004, S. UENo leg; 2(S(S(tenera1), 1?, Esideof table top, 2,640malt.,
18_X-2004,Y.NlsHIKAwAleg. Alldepositedatpresent inthecollectionoftheDe-
partmentofZoology,NationalScienceMuseum(Nat.Hist),Tokyo.
Type locality. Mt. Wawu Shan, 2,640-2,780m in altitude on table top, in
WawushanZhenofHong'yaXian,centralSichuan,SouthwestChina.
Notes. Judgingfromconfigurationofthemalegenitalia,thisnewspeciesseems
closer toE nlba fromMt.NibaShanthantoE btldhalca(DEuvE)(1988,p 256,figs.
6, 16, 23; UENo,1998, p 284, figs 6-7) fromMt.EmeiShan. Asin the former
species,themedianlobeofmalegenitaliaissigmoidallycurvedindorsalviewthough
toalesserextent,andtheapical lobeispointedat theextremity inlateral view.An-
otherproofofthisaffinityisthepositionofthepreapica1poreontheelytra,whichis
locatedat theapical anastomosisof the2ndand3rdstriae. Itstypelocality isalso
nearertopographicallytothatofEnlbathantothatofE budhaica.
Epaphiopsisdaoisasubalpinespeciesratherwidelydistributedonthetabletop
ofMt.WawuShan.Itdwellsunderdeadleavesinthethicketsofarrow-bamboos,par-
ticularlyunder Rhododendrontrees,intheAbles forest,butdoesnotoccurbeneath
slatesontrails.Suchahabitatpreferenceiscommontoall themembersof thenlba
group.
ThenewspecificnameofthepresentspeciesisderivedfromDao(usuallyspelled
“Tao” inEn9lish)ofTaoism,whosefounder,Laozi,issaidtohavespenthislastyears
onthetabletopofMt.WawuShan.
SinotrechiamaparvusS.UENo,sp
nov.
(Figs7-9)
Len9th:5.15-5.50mm(fromapicalmarginofclypeustoapicesofelytra)
Smalto「thananyofthecongenershithertoknown,andcomparablejnthisrespect
andalsoingeneralappearance,toP,,of,e-〔,hiamac・,ass・,pes(s
UENo, 1gg7)1) It Is
hOWeVe「,evidentlyditto「entfromtheMeiguspeciesinthetypjca1lycordatepronotum
リPl'ofree/1i'a″Iae-ssfpes(S.UENo,1997),comb
nov
「'eC1/la7'1aC'aSSIPeSS・UEN0,1997,J.SPele01.Soc.Japan,22,p
38,6g.1;typelocality:Dafengd1n-0n
theDaliangShanMountainsinMciguxjan 一