\\inl~r:!:OlIO THE JAGUARS OF ALTAR Q, COpAN, HONDURAS: FAUNAL ANALYSIS, ARCHAEOLOGY, AND ECOLOGY DIANE1\. BALUNGER Ofllll/~ \/,4" Nor/II 7i.'.\n:' /-Im/lfl Cn)"c51/.~klll Dq!IJrllllcJlI nfP1Iy:>icnl Mt'dicilll'rllld 1~1'I/Illlilill/ti(l1l LIT501l1/1'1'1'~/1"'" Ml'diml emll'l" Dallll:', TX JI::f-:FREY STOMl'ER ColIl'S" (~rLllkl'COIIIII.ll Gmy.~ll/kc, II \lJ~TR,\CI.-.\n l'\(",1\,II;ll(l ,11 COp,1I1, H"ndm'b, ,I I,lk Cl.h~ie \1.1~.1 ~itl" 1\'1'''II,'d Ill<' rjlll,11 (",ldll' "I b'111l·... III ,II 11.',1,.1 (,IUrlv"n bi;.: (",11,. ,1",,,"cj,lh'd with \Iloll" Q <;\'I\'r,11 uf IhL' l.lf).~,. r,ll:> "erl' idel1liik,d ,1'" j,I~U.tr. ('Jlllthem 1!ll"1. I'n'limlll;lry,In<1l~,.i:- "hUII"'d th,ll !h~,111illl,ll,. \\'~r,· 111 ~{'od 11<',11111 ,Il thl'Iim..·o( lill'ir d,·,nh,.. All b,1! "Ill' \\"'r,' ,ldult". 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L,'", \·I·nt,lil,. d,' 1,1 qu~'u,' d~' I.·... oi,.,·<1U" .1 ,1("1I11p,I).';11L; I,·... ,11,·, "'~t1wnt" dl"~ ch,lt"~r"nd ",....(I(IL;... I,'nypl d.e,.I"".1I1iIl1l'.1l1'" il \ 0111 d'·I1.\ ,,"'i,'I1.... I.,.,. I'd;I,·... "llt"I"l",'I1l,'nbtil' I" dt'll" 111"1/ ,.1'. <lllll'lJr...1lllleonrlll" '1"t'Ct· Il'd,'"!1-',,,,,I",,.,.ibk·1'1)111'I,· l.l;.:;lIolr,.fin'"btilll'rd.llhk \",1II.·l'ekC,'1',111 ~,.He~·'Il"" ,I,·...(I'lItf.lint,·...d'!.'n\"in'11111111'Il!. 224 BALLINGERand STOMPER Vol. 20,No.2 INTRODUCTION Thediscoveryofanoffertorycacheoffelinebonesyieldedanexcitingglimpse ofancient Maya ritual behavior toarchaeologists. Felinebones fmed themasonry crypt to the east ofAltar Q at Copan, Honduras, an altar whose sides bear high reliefportraits ofthe sixtccn ruters ofCopan'sClassic Period dynasty. The cache at the foot ofAllarQ contained the remains ofat least fourteen large felines, two macaws,and thetail fansofsevenotherbirds.Thisisa reportofthediscoveryand preliminary analysis of the offertory complex associated with AltarQ and Struc turelQL16in theAcropolisareaoftheMainGroup.JeffreyStamper,thenagraduate student under the supervision of Dr. William Fash, the Director of the Copan AcropolisArchaeological Projectinstituted undertheauspicesofthegovernment ofHonduras, excavated theoffertorycomplex in1988.Theofferingconsisted ofa scaled cryptcontaining the remainsofat least14large felines, including Pallthera onca, and macaws (Amsp.)associated witha stonealtaron a low, round platform that was placed on the central axis of the western side of Structure 10L16in the Main Group atCopan. Until now, the evidence that large felines were involved in Maya ritual has consisted ofskeletonsofsingleanimalsand portionsofskeletonsofcatsburied in ritual contexts (pohI1983, 1990). Landa (Tozzer1941)discussed animal sacrifices statingthatanimalswereused in ritualsandweresacrificed.They werepresented to the gods either alive or not, sometimes dismembered but also whole (Tozzer 1941).ThecacheatCopanisthelargestcacheoffelid,\('bonesdiscoveredbyMaya archaeologists todate. JAGUARSYMBOLISM Maya art and iconography are rich with depictions of jaguars. The same is trueofother Mesoamerican civilizations. Copan, Palenque, Uxmal, and Ttkal are among the lowland Maya sites where jaguarsare featured prominently (Morley, Brainerd,andSharer1983;Spinden1975;Tonerand Glover1910).Spinden (1975) considered them second only to the snake in symbolic importance to the Maya. Cae(1972) pointed out the linksbetween rulers and jaguarsand theseparationof king and commoner in native Mesoamerican religions. He further remarked on the antiquity of the jaguar as il religious icon and its ties to Mesoamerican reli gionsand the ruling lineage (Cae1972). Images of jaguars frequently decorate ceramics recovered from Maya sites, appearing on a variety of vessels. On these, men often wear clothes with jaguar markings or jaguar pelts (Spinden 1975: 149). Many times jaguar symbolism is partofa ritualorreligiouscontextonvessels,suchasonefrom AltardeSacrificios. On thisvessel,therulerofYaxchilan isdressed injaguarskin trousers,miltsmade of jaguar paws complete with claws, and a jaguar headdress. Asecond figure in jaguar regalia is near that has the arms, hands, tail and feet of the jaguar on a humanbody (Saunders 1989: 146). Thejaguarepitomized twodifferent kindsofstrengthtotheMaya.Thejaguar motif was associated with the underworld and its supernatural power and also with physical strength. The association with physical strength derived from the Winter2000 JOURNALOF ETHNOBIOLOGY 225 fact thatjaguarsarcpowerful,nocturnal hunters. Ideasofthesupernaturalpower ofthejaguararosefrom theearly totemic traditionofMesoamericaand thejaguar nagllal's (spiritual co~essence)relationship to the shaman's power. Possession of supernatural powerandphysical strength was important to Maya rulersbecause their political power rested on theirability toactas a priestly bridgebetween the ancestors, the underworld, and the livingworld (Scheleand Freidel1990). Maya artists juxtaposed rulers and warriors with jaguar symbolism in art. Strength and prowess ofcombat were required ofboth the ruler and the warrior but especially of the ruler. Afearsome beastof the tropical forest, thejaguar per sonifiedadualsymbolism:controlofthesupernatural,anecessarypowerfor Maya kings,and thephysical prowessneededby thesuccessful warrior(Saunders1989; Hassig1985). Mesoamerican cultures from Dlmec to Aztec revered the jaguar. The Dime<: associated jaguarswith shamanic powerand filled theirart with imagesofwere· jaguars.TheAztecs linked thejaguartowar,sacrifice,and royalty(Saunders1989: 150;Hassig1985).Furthermore, theyassociated jaguarswithjade, rain, and fertil ity (Sawlders 1989). At the time of the Spanish conquest, Aztec (Mexica) traders routinely transported peltsand liveanimals from outlying parts oftheempire to thecapital. Picturesofboth liveanimalsand peltsappearon tradeandtributelists of goods moving from Soconusco, the colonial province located on the coastal plain ofthestateofChiapas, Mexico, to theAztec Empire (Voorhies 1989). Relictsofthissymbolismare found today in remoteareasofCentral America, Mexico, and South America. Jaguar symbolism is most visible in masks and cer emonies performed in isolated villages. Wearing a jaguar mask transforms the wearerintoanewcreaturethatcombinesanimaLhuman,and supernaturalquali ties(Saunders1989).Thus, thejaguarisan ancientandpotentsymbol permeating nativecultures of Mexico,Central, and SouthAmerica. ARCHAEOLOGY Aspartofthefirst season'sworkoftheCopanAcropolisArchaeologicalProject, William Fash conducted preliminary investigations of Structure lOL-16 and the adjacentPlazaareasin1988(Fash1991;Agl.lrcia,StoneandStomper1989).During thespringofthatyear,excavationsatthesiteofthewesternbaseofStructurelOL· 16and AltarQ were inprogress, supervised byJefferyStomperand veteranlocal excavator, Ismael Gon:.~alcz.Stamper concentrated the initial excavations on the area in front ofand beneath AltarQ, anticipating finding the dedicatory offering or other remains ofassociated ritualsat that locus. Otherexamples ofdedicatory caches had been found either directly in flunt ofor underneath altars and stelae elsewhereatCopan (Stromsvik1941).Anarea wasmarked offdirectlywestofthe altarand excavated to a depth of120em. Fearing for the integrity ofthe trench if hedugbetween thestone supports that uphold AltarQ, Stomperelected instead to tunnel beneath the altar from the eastern sidewall of the same pH. This mini tunnel excavation beneath the altar produced no evidence of a cache, nor was there anyevidence that the area under the altarhad been disturbed. 226 BALLINGERand STOMPER Vol. 20, No.2 - - .. .- .~. ,0: ~ - ~~~"'11- •-• - .~ :"/ : f ACROPOLIS - • • FIGURE 1.-The EastCourt. MainGroup,Copan, Hondur,lS. Supplied by William and Barbara Fash. Winter2000 JOURNALOFETIINOl3l0LOGY 227 Stamper began another pit that mirrored the location of the previous one. Immediately,hefound twofeatures. Thefirst wasaplain, round altar,CPN 13470, with a diameter of 36 em. Altars such as these were used as stands for incense burners elsewhere in Copan during the Late Classic period (Fash 1983:464). The second feature waswhat appeared tobeseveral roughslabsofslone, Feature8,in a line below the level of the last plaza floor. Theexcavation wasexpanded to un cover theextent ofthis feature. The capstones were aligned in a north-south direction, sloping from cast to west, and covering the masonry crypt that contained the animal bones. Probing between thestones revealed a hollowarea apprOXimately1.3m deep and at least 1mwide. WhenStamperremoved thecapstones,hefound thatdebrisfrom places where the walls had caved in filled the crypt. The crypt, measuring 131 em long and 48em wide, had wallsofeightcoursesoffinely cutbuildingstone rising toa heightofl17cm. Within thecrypt, theexcavators found animal bones.The removalofthe first level of bones revealed an irregular intrusion of lime plaster in parts of the cist. Excavation of this layer revealed that more bones were embedded in the layerof plaster with still more bones below it. These layers were removed in the plaster matrix in large sections and kept separated. Stompercontinued excavations into lhe fill under thecist toa depth of3m under theplaza level. He recovered only 7 fragmentsofceramicsfrom underthecisloApproximately319em below theorigi nal cist floor, the excavators uncovered an earlierplaster floor ofthe West Court. Excavations were terminated at this point. Twosmallshaftsadjacenttothecistwerealsouncoveredjustoutsidethenorth east and southeast corners of the cist. The inside of the roughly square shafts measured 22 em on each side. At the bottom of one shaft were nine whole pris maticobsidianbladeletsand partsoftwoothers. Located25cmabovetheobsidian werethebonesofa medium-sized bird.Thesecondshaftalsocontained thebones ofa bird butnoobsidian. Nothingelse was found. Stamperexcavated the entire area between AltarQ and Structure LlO-16, re vealinga round platform ofsmall, faced stones twocourses high in some places. Unfortunately, thewhole platform wasnot preserved. Located in thecenterofthe platform was an oval stone with a smooth upper surface, repeatedly charred by fire. Resting on the top of the platform was a small, highly polished fragment of jade. The area around the platform yielded incensario fragments, a ceramic in censeburner. Inanadjacent area tothe westoftheplatformjustbelow the levelof the last plaza floor, a small cacheofobsidian lancetswas found. FAUNALANALYSIS During thesummerof1988,the faunal material was identified and recorded, and a preliminary report was filed at the Central Office of the Project in Copan (Ballinger 1988). The bones were counted, sorted into skeletal clements, and ex amined forpathologicalconditionsand anomalies.t Ballingerusedweight-bearing bonesoftheappendagestodeterminetheminimum numberofindividuals(MNI) becauseweight-bearingboneshavelargeareasofdense,compactbone,and,hence, tend to be better preserved (Brain 1981). Left and right elements were identified 228 BALLINGERandSTOMPER Vol. 20,No.2 and counted. Ballingermatched the bones bysizeor age to determine if theybe longed to a single individual (Chaplain 1971). Juveniles were identified by the lack of epiphyseal closure. The analyst ignored most of the fragments because time was limited. The largest MNI of the elements was then determined as the MNI for thespecies (Klein and Cruz-Uribe1984).Actualbonecountsweremade atCopan. Photographsofthebestpreservedcraniaweretakenforlateridentifica tion. Final identification tospecies level was made in the Zooarchaeology Lab at Indiana University, whereacomparativecollection ishoused. Analysis was done under field conditions without the use ofa comparative collectionor manualscommonly used by faunal analysts. Neithera comparative collectionnor library resourceswereavailable in1988at the Copanlaboratory to aid in identification. Ballinger took notes supplemented with photographs and sketches. The initial identification ofthebonesasfeline resulted from herexami nationofthe teeth, crania, scapulae, and femora while inHonduras.2 Condition ofthe Bones.-Burialin aclosedcryptresulted in good preservation.Al thoughthebonesweresubjected tonaturaldecay,theywereprotected fromsome ofthe taphonomic processes that radicallychange relationshipsbetweenskeletal elements(Lyman1982).Thus,whilemanyoftheboneswerenotarticulated, they wereclose tothepositioninwhichtheywereplaced inthecrypt.Thecryptwalls keptthemfrombeingdispersedafterburial.Theywerealsoprotectedfrom water andheat,thetwomostpowerfulagentsinbonedissolution (vonEndtandOrtner 1980). The bones were dry and chalky but retained their shapes, so skeletal ele ments were easily identified. Exfoliation was present on some of the bones but mostofthem werewell preserved. Anassessmentofpathological lesionswas madeonthebones thathad mini malflakingonthem.Thebonesofthesecondlevel,excavatedasaunitandcurated asaunit,allowed Ballingertodetermine thepositionofthejaguarsofthesecond level in thecrypt by the association ofskeletal elements. The heads ofsome had beenlaid overthe rear legs and feet ofothers. ResultsoftheFaunalAnalysis.-TheMNIofthefelineswasatleast14.Thisnumber is conservative because the analyst was unable to perform a complete examina tionofallthefaunal material. Ballingerfound thatmanyofthebonesfound inthe cryptwerePantheraonca. Differencesinjaguarsand puma lieincranial morphol ogy. According to Olsen (1968), jaguar crania have a sagittal concavity on the superior aspect of the cranium that rises to a pronounced lambdoidal crest that gives a slight s-shaped curve to the jaguar's skull. The posterior crest gives the jaguarcranium a squared offappearanceand the skull appears longerand more rectangular. The puma,however, hasan ovalskull lackingthe massive,posterior cresting.Craniaofjaguarsandpumaaresimilarin theiranterioraspectsandteeth butdiffer intheposterioraspect. Itis the posteriorfeatures on thecrania thedis criminate between the species. The more complete crania from the crypt have distinctivenuchal robusticityand amoreelongatedarchitectureofthejaguar.AI· though there is not enough cranial material remaining to account for 14jaguars, an estimated 6animals in the assemblage were Panthera onca. The others are fe linesbut remain tobe positively identified as jaguar. The remaining bone in the assemblageisbirdboneand a few intrusive rodentbones. Winter2000 JOURNALOFETHNOBIOLOGY 229 The bones of the felines are very similar in size, indicating that the animals weresimilar inageandsex. Theywerealsohealthy animals. Assessmentofindi cators of general health routinely surveyed by investigators: cortical thickness, osteoporosis,and thefrequency and severityofperiosteal reactions, lesionsofre activebonegrowthresultingfromlocalizedandsystemicinfections,demonstrated that the cats were free from these pathological conditions that commonly mark thebones and indicateadecline inhealthstatus. Large cats are prone to bone diseases in captivity. Osteoporosis, thinning of thebones, isaproblemfor largefelines inzoos.Itresultsfrom inactivity, toolittle protein, old age, and metabolic disorders (Fowler 1986). Caged felidae are also highly susceptible to metabolicbone disease that results in a rickets-likebowing ofthelongbones(Fowler1986).Thecorticesoftheboneswerethickandonlyone periostealreactionwasfound. Thesingleperiostealreactionwasonthehind foot of one animal and had fused two metatarsals. Appendicular bones were robust withprominentmusclemarkings, andhad nosign ofrickets~likebowing. No cut marks, dismembering marks, or skiIUling marks were found on the feline bones that were examined. The removal ofpelts results in acharacteristic patternofskinning marks. SkiIUlingfor pelts often makes cuts ringing the lower metapodial where the knifehascircled theankleor wrist toloosen theskin. Car pals,tarsals,and thebonesofthedigitsarethen removedwith thepeltand,thus, aremissingfrom theassemblage. Ballingerfound noevidencethat thepelts were removedafterdeath. Furtherwork,however,should includecarefulexamination ofthebones for skinningand otherbutchering marks. Inaddition, thevertebraemustbeexamined. Ballingerconcluded thatcrania mayhavebeendisposed ofseparately for someofthecats. Thepaucity ofcranial fragments compared totheamountofpost-cranial materialleadsBallingerto the conclusionthatcraniamayhavebeendisposedofseparately.Furtheranalysismay reveal cut marks on the first or second cervical vertebrae if the heads were re moved beforeburial. Theavianbones in the assemblagewerecollected from twoplaces, thesmall shaftsadjacenttothecryptandfromthecryptitself.Thebonesofanadultmacaw, Amsp., were found ineachshaft. Both of themacaw skeletons were missing the pygostyle, thebone to which the tail fan attaches. Two pygostyles of the correct sizewererecoveredfrom thecrypt,possiblytheonesmissingfrom thebirdsinthe shafts.Thepygostyleisafragilebone,however,and mayhavebeendestroyedby postmortemdiagenesis.Therearesevenotherpygostylesfrom otherunidentified birdsinthecrypt.Threeoftheseprobablyarethesamespecies.Theotherfourare two matching sets and may represent two more species. Lack of a comparative collectionprecludedthecompleteidentificationoftheavianbones.Inall,ninetail fanswereburiedwiththejaguars.Otherbirdbonesburied in thecryptincludeda caudalvertebra,afragmentofa tarsometarsus,afragmentofa proximalradius,a ribfragment, and twophalanges.Alloftheseremain unidentified.Therewereno signsofrodentgnawing orcutmarks on theavianbones. Preservation of the macaw bones differed for the two shafts. Burial 1 lacks cranial bones. Thehead may have been removed at the time ofdeathbut the re mainsweretoobrokentoproperlycheckforcutmarks.Theremainsofmore than 230 BALLINGERand STOMPEU Vol. 20,No.2 one bird may be present in thesecond shaft. Thebones removed from thesecond shaft have more fragments of long bone and cranium than Burial 2. Differences between thetwoshafts'contents,however,mayhavebeenaffected themicroenvi moments of each shaft. Ballinger identified the macaws by the presence of the beak in Burial2and acomparisonofthehvobirds. [twasdearthatpost-cranially they were the same (Hargrave 1970). However, aU of the avian bone should be reanalyzed with a comparativecollection at hand. In summary, a MNI of at least fourteen large felidae, some of them Pall/hem Ollea, were identified from thebones in the crypt. The animals were healthy with no indicationsoflongterm protein deficiencyor inactivity. Accompanying the fe lines were the tailfans of nine birds, including two macaws. No evidence of butcheringor mode ofdeath was found in thisanalysis. Afull study ofthe feline bones may reveal sex and age differences, less obvious evidence of disease, and the mode ofdeath of the animals. Similarly, comparison of the avian material to comparative skeletal collections may reveal the species of the unidentified mate rial. DISCUSSION Natuml Historyalld EcologyoftheJagllar.-Thejaguar(Pufffhem01lea), is the largest ofthreespeciesofspotted catnativetoCentral America (Burton1987).Classed as abigcat, thejaguarhasashoulderheightranging from 110-155cm (Burton1987). Thejaguarprefersa forest habitatbutcan live insavanna environments ifthere is enough brush cover. They also live in slightly arid areas. Good swimmers and climbers, jaguarsare primarily nocturnal feeders, sleeping during the middle of theday (Burton 1987). Few investigators havestudied thenatural historyofthejaguarinthewild so most information comes from zoo studies. Many of the details about breeding patterns come from studies of zoo animals. jaguarsare very easy to raise in cap tivity. They breed easily and can live on two or three pounds of meal a day, according to David Ruhter,J former Curatorof LargeAnimals, Houston Zoo (Per sonal communication via telephone,1994). Less is known about theirbehaviorin the wild. Large felidae are long-lived animals whose prey is normally abouthalf theirbodyweight(SunquistandSunquist1989;Gittleman1984;Packer1986).They usually need largeareas in which to forage and, except for lions, are solitary. Today, the jaguar hasadapted to morecrowded habitats in CentralAmerica. In 1983, Rabinowitz and Nottingham (1986) tracked the movements of nine jag liars in Belize.Theauthors found that the homerangesofthejaguarsoverlapped but that as long as prey was available, they remained on them, avoiding each other.Analysisofscatdetermined thatin Belizejaguarsfeed onseventeenspecies of prey (Rabinowitz and Nottingham 1986). This is consistent with reports from otherareasto thesouth that jaguarsfeed on diverse prey: capybara, fish, peccary, and alligator (Ewer 1973). Generally, jaguars feed nocturnally but the female in the Belize study changed her feeding pattern to the daytime in order to exploit cattleasprey.Thisstudysuggeststhatthejaguarisflexibleand able toadoptnew habits as needed tosurvive (Rabinowitzand Nottingham 1986). Winter2000 JOURNALOFETHNOBIOLOCY 231 Naturalists do not know much about the behavior of the puma in Central America. Felisconcoloroccupiesawider rangeofhabitatsthan thejaguar, ranging from mountains to jungles to deserts (Ewer 1973). The puma is slightly smaller thanthejaguar. Pumadietisomnivorousintropicalclimateswheretheyconsume several typesofrodents,fish, andothersmall game.The maincomponentoftheir diet is deer and domesticated farm animals, although they have demonstrated behavior flexibility in the selection ofprey species. Thejaguarand thepuma arein almost directcompetition in CentralAmerica becauseofthesimilarityin size,habits, and diet. Pumaarealsonocturnal feeders unless forced to feed at other times. They are known to live nearhome ranges of jaguars(Rabinowitz and Nottingham 1986) in Belize,but in the North American west, their ranges are largerand donotoverlap. Theecologyofthejaguarhas important implications for explanationsofhow theMayaacquired atleastfourteen bigcats forritual use. Rue(1987)statesthatby theLateClassic, thevalley washeavilydeforested resulting in a lossofhabitatfor tropicalforestanimals. If,however,enoughbrushyareasstill existed in thevalley, jaguarscould have survived there, given the presence of adequate prey. In gen* eraL species that live in savanna or parkland areas have larger populations than forest livinganimals(Berkoffetal.1984),tllllSdeforestation could lead toa slight increase in the population ofcats in the valley. Jaguars have demonstrated their abilitytosurvivecrowdedconditionsand theabilitytochangefeeding schedules. Itishighlyprobably thatthey survived in a mosaicenvironment ofparkland and brush. Ultimately,however,access topreygovernspopulationsize.Adeforested but stillbrushyareacould havesupported sufficientdeerandotherspeciesofpreyto, inturn,supportasmalljaguarpopulation.Similarly,a mosaicofbrushandcleared spaceswillsupporta white*taileddeer(OdocoilclIsvirginimll/s) populationwhereas closed canopy forest will not (Smith 1975). Deer have small home ranges, often living near humans and thriving. Historica.lly, deer have been an important pari ofMaya subsistence(Mandujanoand Rico·-Gray1991)and only recentlyhavede clined in importance as numbers decreased. Both brocket deer and white~tailed deer feed in cleared areas of new growth near fields because tender shoots are present there. There also may be fewer insects in young forest (Mandujano and Rico-Gray 1991). MandUjano and Rico-Gray (1991) remarked that the decline of Yucatan'sdeerpopulationwasdirectlyattri.butedtoover~huntingand lossofhabi tat, a relatively new situation. Population decline was exacerbated by the loss of native farming methods that provided browse each season after the fields were burned,cleared, and farmers severely trimmed brushygrowth in and around the milpas. This growth sprouted anew with the rains. Pohl (1994) proposed that Copanecos were very likely to have raised deer in and around their homes and fields in the valley. Other animals available as prey are Brocket deer (Mazama sp.) and peccary (TayasslI tayacu). Both would have doneWE,ll in thisopen,brushy habitat. Brocket deerbrowseonthesamekindsofplantsaswhite-taileddeer: tendertwigs,shoots, and leaves of a variety of herbaceous plants and fruits. The peccary is omnivo rous, living in a variety of tropical habitats, including forests and dry savannas 232 BALLINGERand STOMPER Vol. 20, No.2 (Lawlor1979). Smaller prey, such aspaca, agouti, and armadilloare native to the area and can live inbrushy environments. TheCopan Valleycould havesupported asmall population ofbigcatseven if it were partially deforested. Whether itcould have supported a population large enough toprovideatleast14adultbigcats from local sources isanotherquestion. TheCopan Valley iscomprised of26squaremilesofterritory.Ofthat, theheavily populated Copanpocketwould havebeen too urban-likeforjaguarsor pumas to live there. Game such as deer would have been drawn to the areas surrounding fields furtheroutratherthan thekitchen gardensclosertothecentral site. Bigcats would have lived farther outfrom the Main Croup atCopan. Normally big cats have large home ranges of 15 square miles. If we assume thatcrowdinghas limited thesizeofthehomerangeahometo5squaremiles, the Copan Valley would only provide ranges for five jaguars at a time. Avoidance behavior asdemonstrated in Belizewould not make it possible for moreanimals tosurviveunlesseven with preypopulationsofveryhighdensity.Thus, the large number of felines found in the crypt most likely resulted from a combination of trade, hunting, and hand raising. THEOFFERTORYCOMPLEX ANDTHE JAGUAR BURIAL Vax Pasah, the last ruler of Copan, had begun an aggressive new bujJding programbyraisingTemple11and makingsignificantadditionstothe WestCourt, whereAltarQ is located includingAltarQ (Fash1991).Thealtarisan illustration of the succession of the last ruling dynasty of the polity. Along the sides of the altarthe namessixteen rulersareinscribed beginning with VaxKu'kMo' (Blueor First Quetzel Macaw) and ending with Vax Pasah. Vax Pasah, by his choice of iconography for the West Court, indicated its relationship with the underworld, Xilbalba, and, thus, to hisancestors (Sehelcand FrcideI1990). Vax Pasah faced serious problemsduringhisreign, includingdeforestation, a population that wasshrinking, and a diminutionofhispower(Fash1994).Sehele and Freidel (1990) note that most of his monuments state the cosmic sanction of hisrule,and hypothesizethat hisreign wasmarked bycrises. Hisimpressivesac~ rifice and building program may have been an attempt to restore Copan to its previous place in the hierarchyofmajorsites, Structure lOL-16 was the final product of the remodeling in the West Court. Thejaguarburial occurred atthistimeasaceremonial cache for thededication of the last version ofStructurc L10-16. The Copenecos placed AltarQ at the base of this structure. Vax Pasah buried at least 14 big cats in the crypt associated with AltarQ.OncethepeopleofCopan placed thefelines inthecrypt, tail fans ofbirds were placed there, and the crypt was then covered with three capstones. Asmall, round, carved altar was placed next to it directly between the two shafts, Afinal plaza floor was then laid, covering the capstones. Jaguars and other spotted cats had an unknown but important ritual significance atCopan. Pohl (1994) remarks that while skeletons of spotted cats were often cached in ritual contexts by the Maya, Copan has more such caches than any othel' site. All of these have so far been found in elite contexts. Pohl (1994) considers them a measure of the high
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