The Group of Trichotichnus leptopus (Coleoptera, Carabidae) of Japan

65 Pages·1997·63.4 MB·English

by  Morita, S

Elyt・,a,Tolkyo,25(2):521-585,November15,1997
TheGroupof zchotzc usfeptopus
(Coleoptera,Carabidae)ofJapan
SeijiMORITA
Motoazabul-3-28-405,Minato-ku,Tokyo,106Japan
Abstract HarpalinecarabidbeetlesbelongingtothegroupofTric11otichnus lep-
topus(BATES)areenumeratedfromJapan.Tli'1chotlc1/nus leptopus isredescribedon the
basisofthetypeseriesandnewlyobtainedspecimens.Twentyspeciesarenew toscience
anddescribedunderthenamesof「1/1saA,oae,「afp加Is,「abet,「a,7mge,-,「1,nω-al,
r,,, e1ioae, r osa・af yesﾍ,i・0 lats,,e1lsfs, f,,・,i(cid:0)fa, r/,,・,asa,l,a, sﾍ,motel
.
a , si/vesf,・Is, r /1f1-amsh,, r g,,aclffs, r oilｽ、al,al /1fgo川s, na,・M ｽ 1.1,a, aspe,,and r
/fat,a ｽalt,al
Introduction
Themembersofthegenus Tricholichnusarewidelydistributed throughout the
worldwiththeexceptionofAustralia,andareclassified into threesubgenera, Tri-
chotichnus,Be11ogenusandHarpa1oxenus. Thisgenushasbeenrepresentedbymore
thanonehundredspecies.AnoutlineofthegenuswasdelineatedbyNooNANin1985.
IntheJapaneseIslands,thisgenusisoneofthemostimportantharpalinegenera,
sinceit includesthespecieswith reducedhindwings,whicharemuch localizedin
mountainousareas.Besides,theyareverycloselysimilar tooneanothernotonly in
theirexternalmorphologybutalsoingenitalicfeatures,makingitquitedifficulttode-
termine them.
Needlesstosay,thefirstspeciesofthisgroupknowntoscienceisTrichotichnus
1eptopus(BATES),originallydescribedin1883 fromNikko,Central Japan. Itwasre-
describedbyHABU(l961,p.150)inhisrevisionalstudyofthegenusTrichotichnus,
but his careful account wasbasedon elevenspecimens from five localities, oneof
whichwassent toLondonandwascomparedbyBRITToNwith thetypeofT leptopus.
Unfortunately, therewasagrave fault intheprocedure for identificationofHABUs'
specimen,sincethedirectcomparativestudywasbasedonlyontheir external mor-
phology.ThesameviewwasstillheldbyHABUin1973,whenamonographofharpa-
1inecarabidswaspublished.A量or thispioneerwork,thesameauthoraddedonenew
species,Tkasaharai fromCentralJapan.
Morethantenyearsago,KAsAHARAredescribedTyukihiko1andT leptopus,and
theircollectingdatawereshown.At thattime, Ihadanimpressionthatthelattercon-
522 SeijiMORITA
sistedofat least four differentspeciesbasedon theextremely variablecharactersin
theirbodysizeandtheshapeofaedeagus.Inthepast fewyears,KAsAHARAdescribed
severalnewspecies fromCentralJapan.Recently,KAsAHARAand ITOdescribedtwo
morenewspecies fromShikoku.Sinceall ofthemareboth taxonomicallyandgeo-
graphically remote fromT leptopus, thesespeciesarevail eventhough their own
view of T leptopus was basedonHABU's inadequate treatment. Last year,several
speciesmainlyrelatedtoTpacifcatoriusHABUweredescribedbyN.ITO.Inmyopin-
ion,itisprematuretostudythegroupofTleptopusfromWestJapanwithoutacritical
studyofall theknownmembersofthisgroup fromCentralandNorthJapan.Afterall,
taxonomyof theJapanesemembersofT leptopusisnotwellworkedout, thougha
totaloftwelvespecieshavebeendescribedsinceHABU'smonograph.
Toundertake、thistask, Ihavetogivea full redescriptionofT leptopusandto
bringitstruesystematicpositiontolight.Then,twentynewspeciesandnineteenhith-
ertoknownspecieswillbereported.
Materials
Thisstudy is basedonexaminationofapproximately 800specimens. Most of
themwerecollectedby friendsofmineandbymyself. Thetypespecimens ofTli-l-
chotlchnus lept〔)pus borrowed from the Natural History Museum, London.
were
HABU'sspecimenswerestudiedin theNational Instituteof Agro-environmenta1Sci-
ences,Tsukuba.Forcomparison,specimensfromKoreaandPrimorskijTerritorywere
offeredfromDr.IMURAandMr.AKITA,respectively.
Method
SincethemembersofthegroupofT leptopusaremoderatelylarge,itisrelatively
easy toremove thegenitaliaofa thoroughly relaxedspecimen.Whenseparating the
parameres fromtheaedeagus,acarefuloperationisrequirecしsincetheproximalpart
ofaedeagus istransparentand fragile. Inorder tostudystructureof thecopulatory
piece,thedorsalmembraneouspartshouldbecutopenfromtheproximalparttothe
apicalorificewithasharppinandtheinnersacistakenout.
Onlywhenthecopulatorypieceisremoved fromtheinnersacandmountedona
slidewitha fewdropsoflacticacid, itsdetailedstructurecanbeobserved. Aftera
carefulexaminationanddrawing,thegenitaliaarepreservedinindividualvialsin70%
alcohol.
Abbreviations
Inthedescription, thefollowingabbreviationsareused:L-body length, meas-
ured fromapicalmarginofclypeustoapicesofelytra;HW-greatestwidthofhead;
PW-greatest widthofpronotum; PL-lengthofpronotum,measuredalongtheme-
GroupofTti'ic1/otic11nuslptoptls 523
dianline;PA-widthofpronota1apex;PB-widthofpronota1base;EB-widthofely-
tra1base,measuredbetweenlateralendsofbasalborder;EW-greatestwidthofelytra;
EL-greatest length of elytra; WL-greatest length of hind wing; TL- length of
metatarsus; TV-lengthofclaw segmentofmetatarsus; TI-lengthofsegment I of
metatarsus;FL-lengthofmetafemur;MTL-1engthofmetatrochanter;M-arithmetic
mean; NHM-Natura1History Museum, London;NSMT-Nationa1ScienceMuseum
(Nat. Hist), Tokyo; NAS- National Institute of Agro-environmenta1 Sciences,
Tsukuba.
Acknowledgments
I wish toexpressmydeepgratitudetoDr.Shun-Ichi UENo oftheNationalSci-
enceMuseum(Nat.Hist),Tokyo, forcritically readingthemanuscriptofthispaper.
ThanksarealsoduetoDr. NigelSTORKandDr.StuartHINEof theNaturalHistory
Museum,London, forloanoftypematerialundertheircare,andtoDr.TakeshiMATsu-
MURAandDr.Koji YAsUDAoftheNational InstituteofAgro-environmenta1Sciences,
Tsukuba, forgivingmetheopportunityofreexaminingthematerialstudiedbyHABU.
Mydeepindebtednessisalsoduetothefollowingcolleaguesand friends,whose
kind aid and support enabled me to complete this study: Dr. Yuki IMURA, Dr.
KazuyoshiKURosA,Dr.Yoshiro KuRosA,Dr.KimioMAsuMoTo,Dr.GeraldR.Noo-
NAN, Dr. Katsuyuki TERADA, Dr. Shin-ichi YosHIMATsU, the lateMr. Atsuo IzUMl,
Messrs. Azuma ABE, Hideto ARAMAKI, Hirofumi FUJIMoTo, Takashi FUNAKosHl,
Michiaki HAsEGAwA, Hirofumi HAYAKAwA, Katsuo HIRAl, YukihikoHIRANo,Hanmei
HIRAsAwA,FuminoriHIRoKAwA,KatsuhikoHoRI,NoboruITO,Yoshiyuki ITO,Sumac
KAsAHARA, ToshioKlsHIMoTo, TakuyaKURITA,KazushigeKUsANo,Koichi MATsUl,
Yutaka MIYAzAwA, Keiichi MATSUMOTO, Mitsuyasu N1sHIDA, Masaaki NlsHIKAwA,
Hideo OHKAwA, Isao OHTsUKA, Toshihiro OzAKI, MasahiroSAIT0,Shohei SHIM1zU,
ToruSUDA,YuJiTOMISHIMA,KojiTOYODA,TeruhisaUEN0,SatoshiYAMAUcH1,Masa-
takaYosHIDA,andNaohiroYosH1zAwA.
Iamalsoindebtedtomywife,Eiko, forheraidinsearchingforcarabidbeetlesin
thefield.
ClassificationoftheGroupofT lept.opus
Bodysize. L: 8-14mm.Thebodysizeshowsratheranarrowrangewithinasin-
glepopulation.
Co1ou1-. Colour black to blackish brown; elytra with iridescent lustre; ap-
pendagesblackishbrowntoreddishbrown.
Colorationonthedorsalsideandlegsfurnishesanimportantcluefor identifica-
tion.
Head. Headmoderately convex, withshallow frontal furrows; lateral grooves
deep andusually straight; eyes rather flat tomoderately convex; microsculpturenot
524 SeijiMORITA
sharplyimpressedthoughconsistingofisodiametricorwidemeshes,partiallyobliter-
ated;apexoflabrumandshapeofmentumtoothvariablewithinasinglepopulation.
It isverydifficulttodetectslightdifferencesofheadwithout topotypicalspeci-
mens.
Pronotum. Pronotumtransverse; apical marginemarginateor almoststraight;
apicalanglesalittleproducedand roundedat thetips;sidesarcuate,convergent to-
wardsbase,andthenweaklysinuatebeforehindangles,whicharedentate,sharpora
littleproducedlaterally;basal foveaeshallowtodeep,andwithcoarsepunctures;me-
dianlineclearlyimpressed;basealmoststraight,orslightlyemarginateatmedianpart
andslightlyobliquejustinsidehindangles,andborderedthroughout;surfacedensely
andcoarselypunctate,thoughminutelysoonthediscandalmostobsoletealongapical
marginandmedianline;microsculpturecomposedofwideortransversemeshes.
Thepronotum ismutuallysimilar tospeciesbelongingtothesamecomplexso
thatslightdifferencescannotbeexpressedbythestandardratiosofparts.
Hind、、,ings. Thestateofreductionofhindwings isoneofthemost important
charactersforseparatingcomplexes.SofarasIamaware,itisalmostconstantwithin
aspecies.
Elytra. Elytraoval, ovate,elongatedoval,orelongatedovate,moderatelycon-
vex,andwidestataboutmiddleoralittlebehindmiddle;sidesgentlyornarrowlyar-
cuate,andslightlysinuatebeforeapices; intervalsslightlyconvexto flat;striaeim-
punctate;basalborderweaklycurvedoralmoststraight;interval3withadorsalpore,
adjoiningstria2;marginalseriesvariableinthenumberofpores.
Theshapeofelytragivesanimportantclueforseparatingcomplexes.
Malegenitalo・fgan. Aedeaguselongate;basalpartlargeandtransparent; right
wallhigherthantheleft;apicalmarginofmembraneousdorsalpartsimplytowidely
roundeli, or slightly emarginate at apex in dorsal view; apical lobemoderately to
strongly produce andalmoststraightor inclinedto theright indorsal view; apex
thickorborderedabove,sometimesinterruptedatmiddleindorsalview;viewedven-
trally,apexalmostflat,thick,orborderedatthesidesandnotchedatmiddle.
Malegenitaliaserveasoneof themostimportantcharacteristics for identifica-
tion.However,dorsalmarginoftheaedeagusisusuallyslightlyreducedindriedspeci-
mens(cf.Figs 6a,12c).
Structu1-eofaedeagus. Innersaccoveredwithscalesandarmedwithacopula-
torypiece,andsometimeswithamatofspinulesnear thebasal partofcopulatory
piece;copulatorypiecevariableinsizeandshapewithinasinglepopulation,andhol-
lowandopenat thebase. (Thiswasobservedunderamagnificationofatleast100di-
ameters)
Ifthecopulatorypieceislargeinsize,it furnishesanimportantcluefor identifi-
cation.
Unfortunately,recentauthorsincludingmyselfhavenotbeenabletoshow relia-
bility of both external and genitalic characters for separatingspecies. However, six
GroupofT''lcltotic11nusleptopus 525
groupsbasedon theshapeofbody, apexofaedeagusandstateof reduction ofhind
wingsarerecognizableasfollows: -
1. Thelept〔)puscomplex:Bodyrelativelysmall(L:7.7-10.8mm);elytraovaland
ovate; WL/EL0.20-0.27;apical lobeofaedeagusalmoststraight orslightly curved
ventradinlateralview;apexofaedeagusthintothick,notborderedinventralview.
11. The hirasawazcomplex: Body medium to large (L:10.2-12.5mm); elytra
elongatedovate;WL/EL0.24-0.28;aedeaga1apical lobeslightlycurvedventradinlat-
eralview;apexratherthin,notborderedinventralview.
111. The kasahara1complex: Body large (L: 11.0-14.5mm); elytraelongated
ovate;WL/EL0.20-0.25;aedeaga1apical lobestronglyproducedandreflexe
or ro-
bustandthickinlateralview;apexthickandnotorslightlyborderedinventralview.
IV Thetranqui11uscomplex:Bodysmall(L:8.4-9.5mm);elytraovate; WL/EL
0.30-0.37;aedeaga1apical lobeslightly reflexedin lateralview; apexthinandnotbor-
dered in ventral view.
V The daibosatsunls complex: Body medium-sized (L: 9.5-11.0mm); elytra
elongatedoval;WL/EL0.60-0.73;aedeaga1apical lobeelongateandreflexedorpro-
ducedinlateralview;apexnotborderedinventralview.
VI. Thepaaficatoriuscomplex:Bodymediumtolarge(L:9.5-l2.5mm);elytra
elongatedoval;WL/EL0.50-0.68;aedeaga1apical loberatherelongateandproduced
inlateralview;apexthick,borderedatthesides,andnotchedatmiddle,eachsideof
thenotchdentate,sometimesobtuseinventralorapicalview.
Complexofnrichotichnusleptopus
nrichotichnus(Trichotichmls)imafukuiHABU
[Japanesename:Imafuku-tsuya-gomokumushi]
(Figs.1-2)
Trlichotichnus(Trichotichnus) lmafukuiHABU,1961,Bull natn. Inst agric.Sci.,Tokyo,(C),(13): l34,
151,169,figs7,38,39;typelocality:Mt.Nishikoma-ga-take;1973,Carab.HarpaliniinFaunaJapo-
nica,Tokyo,285,figs515,521,530,535,540,pi t9-4.
Trichotichnusimafukui:KAsAHARA,l985,Coleopt.JapanCol.,Osaka,2:147.
Diagnosis. Bodysmallandrobust;sidesofpronotum lesssinuatebeforehind
angles;elytraoval; legsbrowntoreddishbrown;aedeagustubular;ventralsurfaceof
apical lobealmostflat.
Description. L: 8.22-8.74mm. Colour black; ventral side blackish brown to
black;legsbrowntoreddishbrown.
Head weakly convex; frontal furrows very shallow and not reaching lateral
grooves;eyesrather flat;mentumtoothroundedat thetip;apexoflabrumemarginate
at themedianpart,or almoststraight; dorsal surfacefinely or microscopically and
sparselypunctate;PW/HW138intheholotype,1.35in1d,136,1.40in2 ; rela-
tive lengths of antennal segments as follows: - I:II:III:IV :V:VI:XI≒l:0.53:
526 SeijiMORITA
,
b
a c
Fig. 1. Tt・ichotichnus(Trichotichnus)spp. - a,T.(T)imafukuiHABUfromMt.Kisokoma-ga-take;b,
T.(T)hisakoaeMORITA,sp nov., fromMt.Senmai-dake;c,T.(T)alpinusMORITA,sp nov.,fromMt.
Sugoroku-dake.
0.89:0.89:0.90:0.91:1.09.
Pronotumtransverseandflat;PW/PL138intheholotype,1.38in1d,138,1.45
in 2 ;PW/PA t43intheholotype,1.38in1d,136,1.39in2 ;PW/PB122 in
theholotype,1.20in1,3,1.23,1.24in2 ;PA/PB0.86intheholotype,0.87in1?,
0.88, 0.91 in 2 ; sides weakly arcuate, convergent towardsbase, and then very
weaklysinuatejustbeforehindangles,whicharedentate;marginalgutterclosetoside
marginfromapicalangletothewidestpart,andthenbecomingnarrowlydistantposte-
riadandobliteratedatabout1/4 frombase;anteriormarginalsetasituatedatthewidest
part;basal foveaemoderatelydeep,butrathershallowin ;anteriortransverseimpres-
sionshallowbutobsoleteat thesides;basal transverseimpressionobsolete;baseal-
moststraight,orslightly emarginateatmedianpartandslightlyobliquejust inside
hindangles,andborderedthroughout;microsculpturecomposedofwidemesheson
theapicalpart,andoffinetransverseonesonthediscandthebasalpart.
Elytraoval,moderatelyconvex,widestataboutmiddle;EW/PW1.25inthehole-
type, 125 in1(S,128, 1.33 in2 ; EL/EW 153 in theholotype,1.46 in1(3,1.41,
1.48 in2 ;shouldersrounde EB/EW0.70intheholotype,0.68in1d,066,0.70in
2 ;sidesratherstronglyarcuate,slightlysinuatebeforeapices;scutellarstrictoshort,
impunctateand freeattheposterioron andwithbasalpore;intervalsrather flatand
almostsmooth;apicesseparatelyandsimply rounded;basalborderweakly curvedor
GroupofT1i・ic11otichmlslep1opus 527
b
Fig. 2. Malegenital organof Tli・ichotlchnus(Tli・ichotichmts) imafuku1HABU from Mt. Kisokoma-ga-
take. - a,Aedeagus, left lateral view;b,apicalpartofaedeagus,dorsal view;c,apicalpartof
aedeagus,ventralview;d,rightparamere,leftlateralview;e, leftparamere,Ie量lateral view.Scale:
1mm
almoststraight,andjoiningstria1,orclosetoscutellum;marginalseriescomposedof
19-23pores.WL/EL0.24.
Prosternum ciliate; apical part of mesepisterna, metepistema and sides of
metasternumsparselypunctate;basaltwosternitesrugose;medianpartofsternites2
and3,andapicalpartofmetasternumciliate.
Legsmoderatelylong;TL/HW1.03in1d,092,0.93in2 ;protibiaesulcateon
theexternalface;MTL/FL0.44in1d,043,0.45in2
.
Aedeagus tubular and elongate, almost straight in dorsal view; ventral margin
slightlyarcuateinlateralview;apicallobesimplyroundedindorsalview;ventralsur-
faceofapical lobealmost flat (notconcave);copulatorypiecesmallercone-shaped
(cf.HABU,1961,fig39).
Specimense)camined. 1 (holotype), “VII i8, 1958, Mt. Nishikomaga-take,
NaganoP.S.Imafuku”/“20pores”/“Holotype71-1chotlchnuslmafukuiHabu”(NAS);
1d,samedataas for theholotype(NAS);1d,2 Mt.Kisokoma-ga-take,Nagano
,
Pref.,26-VII-l976,S.MORITAleg.
Range. ChubuDistrict(NaganoProf.),CentralJapan.
Notes. Thespecimenscollectedbymyselfwereobtained from thesubalpine
zoneofMt.Kisokoma-ga-take
nrichotichnus(nrichotichnus)hisakoaeMORITA,sp
nov.
[Japanesename:Ashiguro-tsuya-gomokumushi]
(Figs.1,3)
Tlifchotichnusimafukui:MORITA,1986,Shizuoka-no-Kochu,Shizuoka,4:1,3.
Diagnosis. Small species with narrow elytra; body black; femora blackish
brown; tarsi andtibiaemoreor less lighter than femora;basal foveaeofpronotum
small andlinearatthebottom.
528 SeijiMORITA
Description. L: 7.71-8.86mm.Smallspecieswithnarrowelytra;body black;
palpi,antennae, labrumandmandiblesbrown; femorablackishbrown;tarsi and tibiae
moreorlesslighterthanfemora.
Headas inT imafukui,but thegenaearelessconvex;PW/HW135-1.40 (M
1.37)in to d,1.33in19;eyesratherflat;antennaerelativelythick;relativelengthsof
antennal segments as follows: - I:II:III:IV:V:VI:XI≒1:0.50:0.94:0.87:0.93:
087: 1.08.
.
Pronotumconvex;PW/Pi t35-1.41(MI40)in toed,1.36in19;sidesweakly
arcuate in front,andusuallyweaklysinuatealittlebeforehindangles, or rarely con-
vergenttowardshindangles;basal foveaesmallandlinearatthebottom,alittlediverg-
inganteriorly;PW/PA t36-1.44(MI42) in toed,1.39in19;PW/PB122-1.34(M
1.26)info,3(3,1.23in1?;PA/PB0.86-0.96(M0.89)info 0.88 in1
, .
Elytraoval; sidesgently arcuate, veryshallowly sinuatebeforeapices; apices
weaklyproduced;EW/PW123-1.30(MI28)intoed,1.26in1e;EL/EW145-1.54
(MI51) info3d,1.49in19;EB/EW0.66-0.71(M0.69) intoed,0.70in19;mi-
crosculpturecomposedoffinetransversemeshesorpartiallyobliterated.WL/EL0.20.
Protibiaeusuallysulcateon theexternal face, rarely vestigial;TI/TV 1.10-125
(M1.17) in 5ed, 1.20 in19; TL/HW 0.95-1.05 (M 1.00) in5ed, 0.85 in 19;
MTL/FL0.42-0.44(M0.43)in5ed,0.48in l 9.
Aedeagus small and robust, high at about middle in lateral view; apical lobe
slightly inclinedtotherightandmoderatelyroundedatthetip;apexslightlyconcave
inventralview;copulatorypieceverysmall.
Typeseries. Holotype: (S,allotype: ?,Mt.Senmai-dake,22~23-VII-1981,S.
MORITA leg. (NSMT). Paratypes: 7ed, Mt. Chausu-dake, 21~23-VII-1978, S.
MORITA leg; 71(S(S, Mt.Senmai-dake,22~23-VII-1981,S.MORITA leg;1?, 2
,
Mt.Nitta-dake,22-VII-l981,S.MORITAleg.
Localities. Mt. Senmai-dake (type locality), Mt. Chausu-dake and Mt. Nitta-
dake,Shizuoka-shi,ShizuokaPrefecture,CentralJapan.
Furtherspecimensexamined. 1 3 Mt.Kita-dake,alt 2,300m,Ashiyasu-
, ,
mura, Yamanashi Pref., 31-VII-1982, S. MORITA leg; 1,3, Mt.Toyoguchi-yama,
Ooshika-mura,NaganoProf.,25-VII-1993,N.YosHlzAwAleg.
Range. ChubuDistrict (ShizuokaPref.,Nagano Pref., and Yamanashi Pref),
CentralJapan.
Notes. Of the 72 fromthetypelocality, twospecimenshaveanadditional
lateralsetaononesideofpronotum.Fivespecimenshavenodorsalporeontheelytra.
Elevenspecimenshaveanordinaryporeononesideoftheelytra. 0nlyaspecimenhas
anordinaryporeoninterval IIIoftheleftelytron.Asfor thechaetotaxyoftheanal
stemite,onlythreespecimensbearanadditionalsetaononeside.
ThespecimensfromMt.Kita-dakeareslightlydifferent fromthetypeseries:the
elytraarewider,EL/EW 142-1.48(M I45) in3 ,and the legsare lighter. 0ther
standardratiosoftheirbodypartsareincludedwithintherangesofthoseinthetype
series. 0nly a single specimen from Mt. Toyoguchi-yama is relatively large (L:
GroupofTli'ic/1otichnllsleptoptls 529
I - - 」
B
t J
Fig.3. MalegenitalorganofTrichotlchnus(T・litchotlchnus)spp. - a-(i,g,T.(T)hisakoaeMORITA,sp.
nov;e-f,h,T.(T)alpinusMORITA,sp nov. - a,c,e,Aedeagus,leftlateralview;b,(i,f,apicalpart
ofaedeagus,dorsalview;g-h,copulatorypiece. - a-b,g,SpecimenfromMt.Senmai-dake;c-
specimen fromMt.Chausu-dake;e, f,specimenfromMt. Sugoroku-dake; h,specimen fromMt.
Nishihodaka-dake.Scale:A-1mmfora-f;B-02mmforg-h.
8.86mm)andthebaseofpronotumhasdenserandcoarserpunctures.
Theample typematerialwas found fromunderstones lyingat thesidesofa
mountainpath,togetherwithArmatolelridesasymmetricusTANAKA.
Afterseventeenyearsoffightingagainstoralcancer,mymother,Hisako,passed
awayonDecember8,1996,attheageof69.Thisremarkablenewspeciesisnamedto
hermemory,asshehadwatchedmystudyofcarabidbeetlesforalongtimewithboth
affectionandencouragement.
530 SeijiMORITA
nrichotichnus(nrichotichnus)alpinusMORITA,sp nov
[Japanesename:Takane-tsuya-gomokumushi]
(Figs.1,3)
Diagr1osis. Similar toThtsakoaeincoloration,especiallyoflegs,butcloser to
T im(;1fukuiintheshapeofprothorax.Fromtheformer,itisdistinguishedbythefol-
lowingpoints: thebodyisrobust, thesidesofpronotumaresuddenlysinuatebefore
hindangles,whicharedentate, themedianlineofpronotumisdeeper, theelytraare
wider,andtheaedeagusisslenderer.
Des〔lript1on. L: 8.71-9.00mm. Colour as in T hisakoae; eyes rather flat;
PW/HW139,1.45in2(5d,1.43in19;pronotummoderatelyconvex;sidessimilar to
thoseinT imafuku1;basal foveaelargeandshallow;apexwidelyemarginate;PW/PL
1.40,1.47in2ed,1.49in l9;PW/PA t40,1.44in2ed,1.43in19;PW/PB 122,
1.26in2ed,1.25inl9;PA/PB0.87,0.87in2ed,0.88in1!.
Elytrabroad;WL/EL0.25;EW/PW130,1.32in23d,1.36in19;EL/EW144,
1.47in2ed,1.36in19;EB/EW0.68,0.70in2(5d,0.67in19;protibiaesulcateon
external face; TI/TV1.04-1.07in2 and19;TL/HW0.89-1.03 in2 and1 !;
MTL/FL 0.41-0.46in2 and1
.
Aedeagusshortbutslender;viewedlaterally,ventralmarginofaedeagusalmost
straight frombasal t/3tonearapex;apicalpartslightlyinclinedtotherightoralmost
straight; apexslightlybordered indorsalview; ventralsideofapical lobenarrowly
concave;copulatorypieceverysmall.
Typeset・ies. Holotype: ,3,Mt.Sugoroku-dake, 13-VIII-1981,S.MORITA leg.
(NSMT).Allotype:9,Mt.Yumiori-dake,13-VIII-1981,S.MORITAleg.Paratypes:1d,
Mt.Yumiori-dake,13-VIII-1981,S.MORITAleg;19,Mt.Nishihodaka-dake,Azumi-
mura, Nagano Pref., 24-VIII-1985, S. MORITA leg; 2 d, Mt. Nishihodaka-dake,
Kamitakara-mura,GifuPref.,22-VI-l988,Y.KuRosAleg.
Locauties. Mt.Sugoroku-dake,2,660malt., 0omachi-shi,NaganoPref; Mt.
Yumiori-dake, 2,300m alt., Kamitakara-mura, Gifu Prof.; Mt. Nishihodaka-dake,
Kamitakara-mura,GifuPref.,andAzumi-mura,NaganoPref.,CentralJapan.
Furtherspecimenexamined. 1?,Kamikochi,NaganoPref., 9~10-VII-1983,
S.MORITAleg.
Range. ChubuDistrict(NaganoPref.,GifuProf.),CentralJapan.
Notes. Tostudy copulatory piece,only asinglemale fromMt.Nishihodaka-
dakewasdissected.Anothermalehasateeth-patchinapical1/5ofaedeagus.Unfortu-
nately,theaedeaguswasnotcutopenandnotdirectlyobserved.
Thisnew specieswasobtained fromthealpinezoneof theNorthernJapanese
Alps.