The genus Diplurodes Warren (Geometridae, Ennominae) of Sumatra

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eq8k Tiuns, lepid S,oc. Jmpan 57 (2) 9:2-104, March 2006
The genus Diplurodes Warren (Geometri dEnaneo,minae) of Sumatra
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2-27-29Shindori-nisNhiii,gata,950-2036 Japan

Abstract Fiftcen species of the genus Diplurode asre rccorded fi'o mSumatra, Thc followin. asix
new species ure described :D. seinoi, D. h4fidisaccutu sD,. cristatus, D. indentodes ,D. sommereri,
and D. diehli .D. submontana Holloway and D, indentata Warrcn are newly recorded,
Key words Diplttrodes B,oarmiini, Geometrida en,ew species, new record. Surnatra.
The genus Dipturodes Warren was strictly define dby Holloway ( 1994) by close comparison
with allie dgenera in the eleyenth part of "`The moths of Borneo" series, and ten species
were recorded from Borneo, with descriptio nofs four new species. Seven of them were
also recorded from Sumatra ,partl ybased on my material. Two species were lef tunde-
scribed because they were represented by single males having very similar genitall ato
those of the congeners. One of them (Hollow a1y9,94, fig .561) fbund also from Sumatra
wM here be described as a new species, but another species (Hellow a1y99,4, fig .562) still
remains unidentified.
For detaile ddefinitio nof Dipluivdes ,see the paper by Holloway (1994 )and my previous
paper on the PhilippineDiptutvd (eSsat o1,999).
In this paper, fifte esnpecies of Diplurodes will be recorded from Sumatra ,includin gsix
new species and two species newly added to Sumatran fauna.
Detailed notes on the localit ioefs collecting sites in Sumatra were given by Diehl (1982,
1 997), Kobes (1985 ,1992) and Schintlmeist (er 1994).
The fo11owin gacronyms are used to indicat tehe locatio nof the spe ¢ imens. BMNH: The
Natura lHistor yMuseum, London, UK. MS: Manfred Sommerer collection, Munich,
Gei'rnany.NIAES: Natural ResourcesInventoryCenter,NationalInstitutefor Agro-

Environmental Sciences,Tsukuba,Japan. NSMT: National ScienceMuseum, Tokyo.

ZFMK: Zoologische sForschungsinsti tunudt Museum Alexander Koenig, Bonn, Germany.
ZSM: ZoologischeStaatssammulungM,unich, Germany.

Unless stated otherwise, all the spccirnens includin tghe type material recorded in this paper
will be deposite din NIAES.
Dipturodes inundutaProut(Figs1-2)

DiplurodesinunckitaProut,1929:74.

This and the next three new species (sein obiC,fidisaccut ucrsis,tateis) share the following
characteristics: both wings greyish ,medial area of fbrewing and basal half of hindwing
paler than the rest; strong pair sof coremata between abdominal segments 4 and 5, and 7
and 8 in male. This species can easily be distinguishe dfrom the others as fbllows.
Forewing roundish apically; undersides of both wings predominantl yblack on dista hlalf,
the rest paler with a slight ting eof pink. Male genital i(aFig 3.7) :Holloway (1994 f,ig .555,
Berneo),Sato(1999f,ig,45,Mindanao Is.).Female genitali(aFig5.2):Sato(1999f,ig.63,

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Diplt rtodes of Sumatra 93
Mindanao Is.).

Materia lexamined. 1 \, Berastagi ;3 9, Karo Highland 900 m; 2 8' 1 Y Dolok Merangir
,
170 m; 3 82\, Pematang Sianta r3;9 839 4, Holzweg II 1,050 m; 1 8 1 9, [Ibl e1,800 m;
3X2 ?,Prapat;19,Gunung Malayu; 18,BukitSubang; 1r3 ?,Lampung.

Geographica lrange. Peninsula rMalaysia ,Borneo, Sumatra ,Java ,Philippine s(Luzon,
Negros,Mindanao,Palawan).

Diplurodes seinei sp. nov. (Fig 3s-5)
Length of forewjn 1g3-17 mm, wingspan 24-29 mm. Most similar to inundata. ForexN'ing.
More elongate, les sroundish apically; darker between antemedial and postmedjaHines.
Hindwing. Darker basad of postmedia lline .Underside .Dista lblack part pale rand re-
stricted to dista dof postmedia lline m;edial lin emore distinct,
Male genital i(aFig 3.8). Somewhat similar to those of kerangati s(Fig 4.6) in having a
slender saccular proces sclose to ventral margin of valva, but ditferen tfrom them in the fo]-
lowing characteristics. Valva ampler; saccular proces sslenderer, tapering apicad, while in
kerangati supcurved at the apex. [Ibngue-lj kperoces sfrom tegumen more developed, in
form Medial broader.
of semicircle. portion of gnathos
Female genital i(aFig 5.3) ,Sterigma wider tban ]ength ,tapered anteriorly, heavily sc]ero-
tized lateral lyB.ursa copulatrix weakly sclerotized near sterigma, with a small bowl-like
signum posteriorl y.Somewhat similar to those of inundata, but in int{nda tsaterigma much
longer b,ursa copulatrix sclerotized in U-shape posteriorl ys,ignum larger.
Helotype. gZ, N. Sumatra ,Berastagi 1,500 m, 26-28. vii. 1985 (R .Sato) , Paratypes.
Berastag i1,500 m, 1 ?, 26-28, vii, 1985 (R. Sato) ,Mts Dair i1,500 m, 1 g, 27. iv .1986 (E.
W. Dieht),Mt SibayakII,1r,4. 1988(S.& A, Saito)S,amosir,Tbbasee,2 8,20. ix.
v.
1991(Graul& SchintlmeisterS)a,mosir,1g, 24.ix.1991,R Siantar1,8, 20,ii,1991,

Tble,1r,22. 1990(E.W, Diehl),W. Sumatra,Mt Talamau,Westseite1,2km S.[falu1,
v,
r, 11. ix ,1991 (Grau l& Schintlmeiste rB)u,kit Subang, 58, ]9. x. 1981 (A,
Schintlmeiste r)`t.Air Putih Quell SeUd 97030 E/4e09 N 1,650 m", 2g, 27. i. 1989,
"Gunung Leuser, Bepangi 2,200 m, 3e55'26"N197elO'53"E", 1 \, 27. ii .1997 (PIOs s&l
Tarmann), '`Dairi Berge 1,800 m, 30 km E Sidikalang", 1g' 29, 8-9. i , 1981 (M,
Sommerer), `CHolzweg Il 1,050 m, 28 km SW Sianta 9r8059 E12046 N", 2 8 1 9, 1. vi, 1986
(E ,W, Diehl), "Kerinci, Kayu Aro 1,800 m, nOrdl. Sungaipenuh" ,3 e 1 ?, 20-23. ii .1976
(M. Sommerer), "Ostl, Tbbasee Tele ca. 1,600 m", 1 \, 6. ix .1975 (E .W. Diehl) ,"Holzweg
IH ],150 14km NE Prapat98058E12"46N",]?,10-21. 1983,18,21-26. 19g3,
m, xi. x.
24, 13-21. iii .1984, 1 X, 24. iii .1986 (E .W. Diehl) ,MS, "Dairi-Berge, 30 km NO v.
Sidikalan g1,500 m", 3 g" 1 ?, 5, v, 1972 (Roesl &e rKtippper) ,ZFMK.
Geographica lrange. Sumatra.
Etymology. This species is dedicate dto my frien dMr Akio Seino ,a specialist of the
Psychidae ,who was my companion during the fiel dwork (1985 i)n Sumatra.
Diplurodes bijidisaccu tsup.s nov, (Fig s6-8)
Length of forewing i3-17 mm, wingspan 25-28 mm. Simila rto seinoi, but distinguished
from it by paler wings, especially distad of postmedia llines .Medial whitish area between
antemedial and postmedia llines slightly wider on forewing. Also similar to D. sugiltata
Prout, Abdominal coremata develope das well as in the othcr congeners, while in sugittata
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Figs1-36. Diplurodes spp. 1-2. D. inuttdat aProut. o T. 3 - 5 . D . he in o i s p, noN. 3-4. dA,holot>pe.
5. ?.paratyp e6.-8, D.bij7disacculu ssp. nov , 6 - 7 . S ,h ole type . 8 ?,paratype. 9-1O .D.
tnangulata Hollowa.v ,e' 1 1-]3. D. cristatus s p. no v . 11-12 ,X',holot yp13e. .\.paratype.
14-15, D. semicircularis Holloway, X' ,16-18 . D , i nd en t od es s p. no v . 16-17. Z holotype.
18 .I, paratype . 19-20. D. indenrata Warren , i ' ' 2. 1- 2 2 . D . p e t fu , s(Meyrick) S,, 23-25.
D. sommereri sp. nov. 23-24. g, holoty p e . 2 5 . f, pa r at yp e 26-28 .D. kerangatrs
Holioway. ;' .29-30, D, submontana HoLlo w a y . S . 3 1 - 34 . D . d i e h i isp. nov. 31-32 .,r,",
holotyp e3.3. I. paratype ,34. ,L paratypc.35-36. D. decttrsari a(Walker g),.
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vestigial to absent (Hollowa y1,994: 262), D. sttgillala vv'as describe dby Prout (1932 1:00)
from Borneo. To confirm the identificat iofo nthis new species, I examined the male holo-
type of sugiltala (Borne Soa,bah, Mt Kinabalu ,Lumu Lumu, 5,500 ft )and it sgenital islaide
(Geometri gdeaneitali aslide No. 13171) at BMNH.
Malc genital i(aFi g3.9) ,Simila rto those of sugiUata Prout ,but uncus much longer ,a pair
o'f sinuous, interwine sdaccular processes much shorter, bifurcat aepically,
Female genitali a(Fi g5,4) ,Sterigma subelliptical; bursa copulatrix with belt-shap esdclero-
tization near posterio ernd and a small bowl-like signum at lef tside posteriorly,
Holotype. r, N. Sumatra ,Holzweg II 1,050 m, 30. v. 1990 (E .W, Diehl). Paratypcs. N.
Sumatra ,Berastag i2, 8 1 - ' 21. viii. 1979, Mts Dairi 1,600 m, 1 ?, 28, viii, 1979 (Dieh &l
,
Schintlmeiste Hro)l,zweg II 1,050 m,1 \,30-31 .vii. 1985 (R .Sato), 14, 12, v. 1986, 1 8'
1 4, 12. vi. 1986, 1 .7･, 6. ix .1986, 1 g･ ,28. x. 1986, 1 g,2, ix ,1989, 1 4,28. x. 1989, 1 g,
23.i,1990, 1g,21,iii.1990,14,23.iii.I990,1!,14.ii.1991,151,20. 1991,19,2.
v.
ix .1991, Sitahoan ,181 Or,26. vi. 1990, Prapat,19, 18, vii, 1983 (E .W. Diehl) ,Prapat,
Holzweg IV 1,050 Se,7, 1994(H,Inoue),P,Siantar1,g',20.ii.1991,[Ible,
m,1 vi, ncar
Tobasee,1cl",3.vi, 1990(E.W. Diehl),W. Sumatra,BukitSubang,18 (RS-2743)1iPT

19-20, x. 1981 (A ,Schjntlmeiste r`C)Holz.weg III 1,150 m 14 km NE Prapat 98058 Ef2"46
N", 1 8, 10-25. vi, l985, "Tbba-See Ostufe rParapat ca, 1,100 m", 1 [1', 6. vi, 1975, "Bei
Bragtag i1,200 m'1 1 g, 26. ii .1985 (E .W. Diehl), "Brastagi 1,200-1,400 m", 2 \, 28-29.
xi. 1975 (Bende &r Diehl), "Holzweg II 1,050 m 28 km SW Siantar 98e59 E12046 N", 1 9 ,
9. xi. 1970, 1?, 27. ii ,1992, 1\,13. ix. 1995, "Holzweg III 1,150 m 14 km NE Prapat
98058 E12C46 N", 1 di 1a ?, 24, ii i1,984 (E ,W. Diehl) ,"Kerinci, Kayu Aro 1,800 m, n6rdl.
Sungaipenuh" ,1 4, 20-23. ii .1976 (M, Sommerer), `tAir Sirah vic. Padang 1,200 m", 1 ?,
2. 1984(Widagdo)M,S.
vi.
Geographical range. Sumatra,
Etymology ,The specific name, b(fidisaccul ruesfe,rs to the apically bifurca tseacculus in
the male genitalia,
Remarks. Holloway (1994 :262) include dSumatra in the geographica rlange of sLtgiilata
by examining my genital isalide (RS-274 3of) the male specimen taken at Bukit Subang.
However, this specimen should have been identifie dwith this new species. TherefOre
Sumatra should be delete dfrom the range of sugillata.
Diplurodes eristaturs sp. nov. (Figs11-13)
Length of forewin g13-14 mm, wingspan 24-27 mm. Also similar to seinoi. Smaller in
generat .Both wings paler with les sdefine dmarkings; les scontrast between dark and pale
areas; dark area more unifOrmly grey ;basa lgrey shading extending to medial lin eon
forewing,
Male genital i(aFi g4.2). C]car]y djstinguishe dby a large protuberanc weith several short
crested processes at mid-valva. [[bngue-lik eproces sfrom tegumen inverted U-shaped ;me-
dial spinous part of gnatho slonger than width, tapered apicad; sacculus short with a digjtate
process,
Female genital i(aFig 5.5), Sterigma narrow; bursa copulatrix slender, cylind[ical, weakly
sclerotizcd and ribbed posterior lwyit,h a small signum at lef tside posteriorly.
Holotype.ge,SumatraUtara,Sitahoan1,200 4-5. 1994 (H.Inoue).Paratypes.
m, vi.
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Figs37-45 Malegemtahaot Diplurodesspp 37 D inundataProut RS-2706 38 D sein(n sp
nov RS 4059 39 D bifithsac(u lsup hnov RS 6805 40 D semiLircular!sHolloway
RS 2697 41 D trtang"lataHolloway RS 1844 42 D (ristatussp noN RS 604g 43
D tndentodes sp nov RS 4057 44 D tndentata Warren RS 60S9 45 D petray
(MeynLk )RS 6060
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Figs 46 -5 1 MalLgLnitdliaofDipturodesspp 46 D kerangattsHolloway RS 1846 47 D sttb
inontanaHo]loway RS 2712 48 D sornmereits p no N RS 2745 49 D diehh sp noi
RS 2717 50 l) decuisana(Walker )RS-2741 5 1 D sinecoremata Holloway RS 2694
Samosir ,1 8, 20 ix 1991 (E W Diehl), W Sumatia, Bukit Subang, Bergurwald 1,200 m,
22 km E Padang, 1 8, 19 x 1983 (Schintlme iRsoteselre, r& Widagdo) `Kermci, Kayu
Aro 1,800 m, nordl Sungaipenuh" ,4 r 1 ?, 20-23 ii 1976 (M Sommerei) `eHo17weg III
1 150 m 14 km NE Prapat 98058 E12C46 N", 1 8, 21-31 vii 1984, "Sipirok III 1,300 m, 10
km NE Sipirok"19, 29 1995(EW Diehl),MS S Sumatra Berastagi1,?,4
i vi
1972, "Dam-Berge", 1 g, 5 v l972 (Roesl &e rKupper) ,ZFMK
GeographicalrangeSumatra

Etymology This bpecdic ndmc, cnstattts, retcrs to the crest-1ike protuberanc oen the valva
in the male genitaha
DrplurodegtriangulataHolloway(Fig9s-1O)

Dipiurodes tnangttlata Hulloway 1994 264
Ths and the next three gpecies (semtaizul ainrdeintbat,a, indentodes) are similar in uni
fbrmly brown wingg, and dre characterized by the presenc eof a pair of coremata between
abdommal segments 3 and 4, in addition to the two parr s(4- 5and 7-8) in male
This species is ditstinguis hferdom the others by the strong black medial band on the hind-
wing and the maggive coremata between abdominal segments 3 dnd 4 m the male Male
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genitali a(Fi g4.1): Holloway (1994 f,ig .564, Borneo) . Female genitali a(Fig ,56):
Holloway(1994f,ig,566,Borneo).

Materia lexamined. 1 g' ,KaTo Highland 900 m; 1 oi, SRII, 11 km off Sindar Raya 400 m; 2
8' 8?, Holzweg II 1,050 m; 4"29, Prapat ;1 e' ,Sitahoa n1,200 m; 1 r (RS-1835),
Gunung Malayu.

Geographica lrange. Borneo, Peninsular Malaysia, Sumatra,
Remarks, Holloway (1994 i)nclude dSumatra in the geographica rlange of this species
based (RS-1835, Gunung Malayu).
on my genitali aslide male,
Dipturodes semicircularis Holloway (Fig 1s4-15)
Diplurodes semicircutaris Holloway, [994: 264.
Very similar to triangulata .Coremata between abdominal segments 3 and 4 much smaller
or vestigial in male; media] black ljne les sdefine don hindwing ;dista dlark and basal pale
regions more clearly diyide dby distin cptestmedial line son underside, Male genital i(aFig,
40)iHolloway(1994f,ig.563,Borneo),Sate(1999f,ig.52,Palawan Is,).Female genitalia

(Fig5.7):Sato(1999f,ig.66,Paiawan Is.).

Materia lexamined. 1 S, Berastag i1; 9, Karo Highland 900 m; 3 r, Dolok Merangir 170
m;1 9, Ketambe;1 g, SRII, 11 ktn eff Sindar Raya 400 m;9 g9 9, Holzweg II 1,050 m;2
2 ?,Prapat;1g',Gunung Ma]ayu.
or
Geographica lrange. Peninsular Ma]aysia, Borneo, Sumatra ,Philippin e(sPalawa [nI,lawi
[lbwi).
DiplurodesindentataWarren (Figs19-20)

DiplurodesindentatWaarren,1897:93.

This species was describe don the basi sof a single malc from Penungah in N, E, Borneo.
Holloway (1994 )recorded three specimens (se uxnknown) from Borneo with redescription
of the male, but did not mention the female, The female has not yet been described,
Male genital i(aFi g4,4). Most similar to those of petras, but tongue-like process from
tegumen shorter, medial spinous part of gnatho sbroade rdistall y.Also illustra tbeyd
Holloway(1994f,ig.560,Borneo),

Material exainined, N. Sumatra ,Holzweg Il 1,050 m, 18 km to Prapat, 1 8i ,4, xi. 1989 (E,
W. Diehl). "N. Sumatra ,N,W, v, Medan, Stabat" l, 8, 30. iv ,1972 (E ,W. Diehl), ZFMK.
Geographica rlange. Borneo, Sumatra (new record),
Remarks. Holloway (1994 r)eported this species to be rare in Bomeo. Also in Sumatra it
seems to be rare, because only two male specimens were recorded above from Sumatra,
Diplurodesin`lentodes (Fig1s6-18)
sp. noy.
Length of forewing 13-15mm, wingspan 21-25mm. Simila rto indentata .Medial part on
fbrewing and basal part on hindwing paler than the rest, while in i.ndentat aalmost unifOrm-
ly brown on both wings.
Male genital i(aFig 4,3). Similar to those of indentat aand petras, Tbngue-lik eprocess
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Figs S2-64 Female genital oifaDrpimodey spp 52 D !nund at a P r o u t RS 6 0 4 6 5 3 D yetnoi
sp noy RS 4060 54 D btfidtsacculu snsoNp RS 2721 55 D (msta tM s s p nov RS
6049 56 D triangutataHollo"ay RS 2930 57 D s em tc i tc tt t ar is H o l lo w a v J RS 2746
5S D vzdentodeg sp nov RS 6064 59 D petras (M e y r i L kR)S 6057 6 0 D kerangattg
Holloway RS 2744 61 D yubmontana Hollowdy RS 1353 62 1)soinmelert sp noN
RS 27tS 63 D diehtisp nov RS 6034 64 D decu rs a r i a ( W a l ke r ) RS S307
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from tegumen much shorter; medial spinous part of gnathos much wider, les stapered apic-
ad; valva costa more strongly sclerotized; saccular process asymmetrical, lef tone longer
than the right.
Female genitali a(Fi g5.8) ,Distinct from those ofpetvas, Sterigma longer than wide, al-
most parall eslided, with ridge-like protuberanc elsateral lbyu;rs acopulatrix weakly sclero-
tized posterier lwiyth small signum near II posterio ernd.
Holotype .g, N, Sumatra ,Mt Sibayak ,4. v. 1988 (S .&A, Saito )P.aratypes ,Sumatra .W,
Sumatra ,Bukit Subang, 1 \, 19. x, 1981 (A. Schintlmeister )S,, Sumatra ,Lampung, 1 g' 1
4,x. 1995 (nati vcoellector). P, Malaysia, Genting Highlands ,2\, 6-8, iv .1986 (K.
Yazaki),Came-ron Highlands, 1if1'･tt,19-20.iv.1990 (N.Bito).

Geographical PeninsulaMralaysia,Sumatra.
range.
Etymology ,This species is similar to D, indentat Waarren in colour and maculation of both
wmgs.
Diplurodespetras(Meyric(kF)ig2s1-22)

Ecn'(rpi psetra s"Ceyrick ,1897, 7}'ans .ent. Soc ,Lond, 1897 (1) 7:5.
Diplurodes petras: Holloway, 1994: 263.
Similar to indentata and indentodes, but distinguish efdrom them by much paler brown
wings and the absence of coremata between segments 3 and 4, Male genital i(aFi g4.5):
Holloway(1994f,ig,559,holotypeB,orneo).

Female genital i(aFig 5.9). Il]ustrat ehdere fbr the firs ttime, Sterigma short, almost paral-
le lsided; bursa copulatrix sclerotized and ribbed, restricted near the posterior end.
Material examined, 1 4, Karo Highland 900 m; 1 8, Dolok Merangir; 38 o7' 6 ?, Holzweg
II 1,050 m; l 8, Ketambe; 1 r, Prapat; 5 r 1 9, R Sjantar ;1 S, Aek [[laru Iml 150 m; 2 8,
Paklca t1 ;8' ,SRIL 1 1 km off Sindar Raya 400 m; 2 ?, Barus, Sibolga ,Mt Pinapan,
Geographial Borneo,Sumatra.
¢ range,
Remarks. This species was treated as ajunior synonym of D. dect{rsar i(aWalke r1,863) by
Parsons et al, (1999 :234), but had already been recognized as a distinc tspecies by
Ho]loway (1994),

DipturodeskerzingatiHoslloway (Fig2s6-28)

DiplurodeskefungatisHo]loway,1994:262.

In this and the next two species (submontan sao,mmereri), the fbrewing is reddish grey and
the hindwing whitish especially in basal half ,corernata develope dbetween segments 4 and
5, and7 and 8, but lacking between 3 and 4, In some specimens (Fi g2,8) ,outer edge of
postmedia lline banded with black on both wings, Male genital i(aFig .46): Holleway
(1994fi,g.565,Borneo),

Female genital i(aFig 6.0). Illustrat ehdere for the firs ttime. Sterigma tapered pesteriorly,
latcr arlnargins heavily sclerotized; posterio rhalt o'f bursa copulatrix cylindrical with scle-
rotization at right near the end, the anterior half globular with a 1arg ebowl-like signum at
Ieft.Matcrial
examined. 2g, Dolok Merangir 180 m; 35 84\, Holzweg II 1.050 m; 2g,
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TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan
Dipt"rodesofSumatr a 101
Prapat ,Holzweg IV 1,050 m; 7 81 4, Prapat 1,150 m; 2 r1 9, Gunung Malayu 80 m; 1
8,Aek [farumII180 2 S,BukitSubang.
m;
Geographical range. Peninsular Malaysia, Borneo, Sumatra,
Diplurodes Holloway(Fig2s9-30)
submontana
Dipturodes suhmontana Holloway, 1994: 262.
Simila rto kerangati s.Forewing Ies ssuffused with grey; line smore defined .Hindwing
with more sinuous postmedia lline ,Male genitali a(Fig ,47): Helloway (199 4f,ig .558,
Borneo).
Female genitali a(Fi g6,1) ,Illustra theedre for the firs ttime, Clearl ydiffere nftrom those of
kerangatis ,Sterigma large rwith triangular depressio nanteriorly; bursa copulatrix globular
as a whole, sclerotized and ribbed near the posterio ernd, with a larg enarrow signum medi-
ally.
Materia lexamined. 26 S 15 9, Holzweg II 1,050 m; 1 8' ,Prapat ,Holzweg IV 1,050 m; 3
r 13 9, Prapat 1,150 m; 1 9, P, Sianta r2; 8, Sitahoan 2; g, Bukit Subang,
Geographical Borneo,Sumatra(new
range. record).
Dipluroctes sommereri sp. nov. (Fig s23-25)
Length of forewin g13-16 mm, wingspan 21-26 mm. Simila rto subnzontana, Forewing.
Postmedial line outcurved between veins CuA, and CuA,, while in submontana deeply in-
curved from M, to inner margin, Hindwing, Postmedial lin emore weakly incurve dthan in
submontana, especially between CuA, and CuA2. Underside. Distal black area wider than
insubmontana,
Male genitali a(Fi g4.8) .Distinguish efdrom those of the other congeners by strongly con-
vex ventral margin of valva and tightl ycoiled saccular process .Also illustra tbeyd
Holloway(1994f,ig.561).

Fema!e genital i(aFi g6.2) .Sterigma shorter than width, parall eslided; bursa copulatrix
pyrifOrm, sclerotized and ribbed near the posteri' eonrd, with a small signum.
Holotype. r, N. Sumatra ,Holzweg II 1,()5 0m, 2, ix ,1986 (E .W, Diehl) . Paratypes.
Holzweg II 1,050 m, 1 r, 30-31, vii, 1985 (R ,Sato )1, 8, !8, vi. 1986, 1 4, 21. ii .1987,
Sitahoa n1,200 m, 2\, 26, iv ,1990 (E ,W. Diehl) ,1 if'19 ,4-5. vi. 1994 (H. Inoue), R
Sianta r2, 82?, 20. ii .1991 (E W. Diehl), Prapat, 1 if 3' 9, 15. v-3. vi. 1983, 1 9, 16. viii.
1983, Gunung Malayu 80 m, 1 g 1 ?, 4-5. v. 1983 (E .W. Diehl), W. Sumatra, Bukit
Subang, 2 g, 19-20. x. 1981 (A .Schintlmeiste rS). ,Sumatra ,Brastagi 1,200 m, 1 r, ]7, ii,
1985 (E .W. Diehl) ,"Holzweg II I1,150 m 14 km NE Prapat 98058 Ef2046 N", 1 9, 2-10,
xii, 1983, 1 r, 10-21 .xi. 1983, 1 8A ,25-31. i .1984, 1 9,11-30, v, 1984, 1 9,10-25. xi
1984,18, 1-16. iv .1985,1?,5. vi. 2000 (E ,W, Diehl) ,MS, "Holzweg III T,iga-Dolok",
1 9, 2. vi. 1972 (Roesl &e rKUpper), ZFMK.
Geographical Borneo,Sumatra.
range.
Etymology. This new species is dedicated to Mr Manfred Sommerer, who is an excellent
lepidopter iisn tGermany, and kindl ysupplied me with a large number of specimens used in
this study.
Remarks. This species was recorded as Diptt{rode ssp. from Borneo and Sumatra on geni-
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