The buccinid gastropod Deussenia from Upper Cretaceous strata of California

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THE VELIGER
© CMS, Inc., 2000
The Veliger43(2):118-125 (April 3, 2000)
The Buccinid Gastropod Deussenia From Upper Cretaceous
Strata of California
RICHARD SQUIRES
L.
Department of Geological Sciences, California State University, Northridge 91330-8266, USA
AND
LOUELLA SAUL
R.
Invertebrate Paleontology Section, Natural History Museum of Los Angeles County, 900 Exposition Boulevard,
Los Angeles, California 90007, USA
Abstract. Rare specimens of three new species of the Late Cretaceous buccinid gastropod Deussenia Stephenson,
1941, are reported from California. Deussenia sierrana sp. nov. is from lower Campanian strata in the Chico Formation
in the Pentz area, Butte County, northern California. Deussenia californiana sp. nov. is from upper middle to lower
upper Campanian strata in the Tuna Canyon Formation in the Garapito Creek area, eastern Santa Monica Mounains,
Los Angeles County, southern California. Deussenia pacificana sp. nov. is from uppermost Maastrichtian or possibly
lowermost Paleocene strata in the Dip Creek area, northern San Luis Obispo County, central California. These three
new species are the only known occurrences of this genus from the Pacific coast ofNorth America. Deussenia has been
reported before only from upper Santonian to lower Campanian strata at the mouth of the Mzamba River (Pondoland,
Transkei) in South Africa and from Campanian to Maastrichtian strata in Texas and the GulfCoast ofthe United States.
INTRODUCTION discussion as to why he chose this family. Sohl (1964)
assigned Deussenia to family Melongenidae, and he also
Late Cretaceous buccinid gastropods are relatively un-
common on the Pacific coast of North America. Recent gave no discussion for the basis ofthis assignment. Mel-
ongenids have pyriform to fusiform shells, usually shoul-
inspection ofpreviously collected material resulted in the
dered whorls, a long anterior canal, and a smooth colu-
detection of three new species of genus Deussenia from mella (Wenz, 1943; Davies & Fames, 1971; Rosenberg,
widely separated locales in California (Figures 1, 2). It is
the purpose ofthis paper to describe and name these spe- 1992). These morphologic features are present on all spe-
cies of Deussenia Stephenson, 1941, including the new
coifesD.euTshseeyniasi,gnwifhiiccahntlwyasextpernedvitohueslbyiokgneoowgrnapohnilcyrfarnogme mspeelcoinegsedneisdcsritboedbeheare.suPbofnadmeilry&ofWaBrucecnin(i1d9a8e8.) Arkegearrsde&d
Upper Cretaceous rocks in South Africa and southeastern
Akers (1997) did likewise and, furthermore, placed genus
United States.
Abbreviations used are: CASG, CaliforniaAcademy of Deussenia in subfamily Melongeninae.
Sciences, Geology Section, San Francisco; CSUC, De-
partment ofGeology & Physical Science, CaliforniaState Genus Deussenia Stephenson, 1941
University, Chico; CSUN, Department ofGeological Sci- Type species: Deussenia cibolensis Stephenson, 1941, by
ences, California State University, Northridge; LACMIP,
original designation; UpperCretaceous (upperMaastrich-
Natural History Museum of Los Angeles County, Inver- tian) Kemp Formation ofthe Navarro Group, easternTex-
tebrate Paleontology Section.
as.
SYSTEMATIC PALEONTOLOGY
Discussion: Deussenia resembles the Upper Cretaceous
Superorder CAENOGASTROPODA Cox, 1959 bucciniform Aliofusus Stephenson, 1941, but Deussenia
NEOGASTROPODA differs from Aliofusus by having a stronger subsutural
Order Thieie, 1929 collar, less inclined growth lines on the collar, and
Family Buccinidae Rafinesque, 1815 straighter axial ribs. Aliofusus has axial ribs that are
curved and follow the outline of the outer lip.
Subfamily Melongeninae Gill, 1871 Deussenia superficially resembles the Upper Creta-
Discussion: Stephenson (1941) questionably assigned his ceous volutid genus Volutomorpha Gabb, 1877, but
genus Deussenia to family Buccinidae, but he gave no Deussenia lacks volutid characteristics in that it has no
&
R. L. Squires L. R. Saul, 2000 Page 119
T3
Age ° Sub- Deussenia
(Ma) (5! Stage stage n. spp. Ranges
DANIAN
65
D. paciflcana
Upper
MAASTRICHTIAN
70-
Lower
Upper
75-
CAMPANIAN "I D. californiana
Middle
80
LowerlD. sierrana
I
85 SANTONIAN Upper
Figure 2
Age andstratigraphic positionsofthe three new speciesofDeus-
senia. Geochronologic time scale from Gradstein et al. (1995).
100 250 500 miles
Min
i I
Pondoland, Transkei, South Africa (Sohl, 1964). These
latter two species were questionably assigned by Sohl
Figure 1
(1964) to genus Deussenia.
Index map to type locality areas of the three new species of
Deussenia. Deussenia sierrana Squires & Saul, sp. nov.
(Figures 3-5)
fold(s) on the columella, no posterior notch, and no sig- Diagnosis: A small-shelled species of Deussenia with a
moidal deflection of growth hues near the suture. Fur-
prominent sutural cord, subtabulate body whorl shoulder,
thermore, Deussenia differs from Volutomorpha by hav-
and nearly obsolete body whorl sculpture.
ing ahigher spire, a shorterbody whorl, atwistedanterior
mm
end, and more dense ornamentation. Description: Shell small, 25 high (incomplete); fu-
Prior to this present study, only 10 other species have siform; spire of medium height, about two-fifths of total
been placed in genus Deussenia, and Sohl (1964:200) height of shell; spiral angle about 45°. Protoconch not
listed them. All are of Late Cretaceous age. Four of the preserved. Whorls 6 (estimated). Upper spire whorls low-
species are from upper Maastrichtian strata in eastern ly convex, smooth. Ramp concave, constricted, and well
Texas, but Sohl (1964) beheved that some ofthese names differentiated starting on more mature half of ante-pen-
might be synonyms because they are (1) distinguished on ultimate whorl and continuing onto penultimate and body
minor differences in shape and ornament, (2) all are from whorls; ramp widest (about 0.75 mm) on body whorl.
the same stratigraphic horizon in a limited geographic Ramp always borderedposteriorly by aprominent sutural
area, and (3) the number of available specimens of these cord and always bordered anteriorly by a subtabulate
four species are so few that it is not possible to determine shoulder Prominent sutural cord somewhat undulatory on
whether or not the minor differences are significant. body whorl. On penultimate whorl, subtabulate shoulder
Four of the other known species of Deussenia are with low axial ribs. Body whorl sculpture mostly obso-
found in upper Campanian to Maastrichtian or upper lete, with faint spiral ribbing near neck area on ventral
Maastrichtian strata in Texas, Mississippi, and Tennessee, face. Growth lines prosocline on ramp; sharply flexed
and are found, to a lesser degree, in similar age strata in (sinused) and opisthocine on subtabulate shoulder; broad-
Alabama and Georgia (Stephenson, 1941; Sohl, 1964; ly prosocline on most of body whorl; nearly straight on
Akers & Akers, 1997). The other two known species of whorl base; and strongest near outer lip where growth
Deussenia are from upper Santonian to lower Campanian rugae develop. Aperture elongate, anterior end (incom-
strata at the mouth of the Mzamba (= Umzamba) River, plete) slightly twisted to left. Columella smooth, with a
Page 120 The Veliger, Vol. 43, No. 2
w
12
&
R. L. Squires L. R. Saul, 2000 Page 121
light callus. Outer lip sinuous, notched slightly opposite Woods, 1906:321-322, pi. 39, figs. 2a-c; pi. 40, fig. 1)
subtabulate shoulder. from the Upper Cretaceous Mzamba Formation in South
Dimensions of holotype: Height 25 mm (incomplete at Africa. The genus Cryptorhytis Meek, 1876, is not a buc-
both extremities), width 11.2 mm. cinid because it has one weak fold on the inner lip and a
weaker fold in the interior of the aperture, and Wenz
Holotype: LACMIP 12717. (1943) believed it to be a volute. Sohl (1964) was the
Type locality: CSUC loc. PN32, latitude 39°39'08"N, first to suggest that "C." pseudorigida might belong to
longitude 12I°35'50"W. Deussenia. We also consider Rennie's species to be a
Deussenia because it has a fusiform shape, along anterior
Distribution: Chico Formation, informal Pentz Road canal, a smooth columella, and a concave ramp with a
member of Russell et al. (1986), Pentz area, Butte Coun- cord bordering the suture; all ofwhich characterizeDeus-
ty, northern California. senia. The new species differs from D. pseudorigida by
Geologic age: Late Cretaceous (early Campanian). being narrowerjust anterior to the ramp, by having only
a hint of spiral sculpture rather than strong spiral sculp-
Discussion: Only three specimens were found. Two are ture, and by having no axial nodes on the shoulder ofthe
from CSUC Iocs. PN31 and PN32, and the third is from body whorl.
LACMIP loc. 10833. Except for the holotype, the speci- Rennie (1930) provided the name Cryptorliytis pseu-
mens are internal molds. It is likely that the holotype has dorigida for specimens misidentified by Woods (1906) as
been slightly worn or weathered, but the shell does not
the closely allied, so-called "'Cryptorhytis'' rigida (Baily,
appear to ever have been strongly sculptured.
1855:459, pi. 12, fig. 14; Rennie, 1930:225-227, pi. 27,
Specimens ofDuesseniasierrana are known only from
figs. 9-12; [non Woods, 1906:321, pi. 39, fig. 2a-c, pi.
tRhuessCehlilcoetFaolr.m(a1t9i8o6n)inintfheerriendfotrhmaatltPheentmzemRboeard mweamsbedre.- 4r0i,gifdiag.di1ff=ers"fCr"omps"eCud"orpisgeiuddaorRiegnindiae]b)y. h'a'Cvriynpgtoarfhiiygthise"r
posited under estuarine conditions and contains mixed
spire, a much more tabulate body whorl shoulder, and
shallow-marine and brackish-marine faunal assemblages.
Many faunal elements ofthe Pentz assemblages, however, narrower and more numerous axial ribs. Based on the
suggest normal-marine conditions (Squires & Saul, 1997; very close morphological similarities between these two
Haggart et al., 1997), and the localities yielding D. sier- species, we also consider "C." rigida to be a Deussenia.
rana were considered by Watkins & Gohre (unpublished Both species are based on specimens collected from the
MS) to represent a fully marine, shoreface deposit. type section of the Mzamba Formation at the mouth of
In the Pentz area, the Chico Formation probably does the Mzamba River. Rennie (1930) reported "C." pseu-
not exceed 150 m (450 ft.) in thickness (Russell et al., dorigida from Beds 3 and 16, and Greyling (1992) re-
1986), and based on the presence of the ammonites Bac- corded "C" rigida from Beds 9, 11, and 16. Klinger &
ulites chicoensis (Trask, 1856) and Submortoniceras chi- Kennedy (1980), on the basis of the ammonite fauna,
coense (Trask, 1856), the rocks are early Campanian in placed the Santonian-Campanian boundary at the base of
age (Matsumoto, 1960; Russell et al., 1986). The early Bed 8, but Greyling (1992) placed it at the top of Bed
Campanian magnetic anomaly 33R is included within the 12. In spite of this debate, both species are of late San-
ranges of both of these ammonites (Ward et al., 1983). tonian to early Campanian age and are about the same
The gastropod Anchura callosa Whiteaves, 1903, is also age as D. sierrana.
present in these rocks, and suggests an early Campanian
age (Elder & Saul, 1996). Etymology: The new species is named for the Sierra Ne-
Deussenia sierrana somewhat resembles the so-called vada range, which rises immediately east ofPentz, north-
Cryptorhytis pseudorigida Rennie (1930:227-228; em California.
<r-
Explanation of Figures 3 to 12
All specimens coated with ammonium chloride. Figure 7. Abapertural view. Figure 8. Apertural view with apor-
Figures 3-5. Deussenia sierrana Squires & Saul, sp. nov., ho- tion ofbody whorl removed to bettershow the columella. Figure
lotype LACMIP 12717, CSUC locality PN32, Pentz area, height 9. Paratype LACMIP 12719, apertural view, height 10.75 cm,
25 mm, X2.6. Figure 3. Apertural view. Figure 4. Abapertural X0.8.
view. Figure 5. Right-lateral view showing outline ofouter lip. Figures 10-12. Deusseniapacificana Squires & Saul, sp. nov.,
Figures 6-9. Deussenia californiana Squires & Saul, sp. nov., holotype CASG 61598.01. CASG loc. 61598, Dip Creek area,
CSUN loc. 153, Garapito Creek area. Figures 6-8. Holotype X1.9. Figure 10. Apertural view. Figure 11. Abapertural view.
LACMIP 12718, height 11 cm, XI. Figure 6. Apertural view. Figure 12. Right-lateral view.
Page 122 The Veliger, Vol. 43, No. 2
&
Deussenia californiana Squires Saul, sp. nov. Deussenia californiana is also similar to Deussenia
pseudorigida (Rennie, 1930:227-228; Woods, 1906:321-
(Figures 6-9)
322, pi. 39, figs. 2a-2c; pi. 40, fig. 1) from upper San-
Diagnosis: A large-shelled species of Deussenia with a tonian to lower Campanian rocks in the Mzamba For-
rounded body whorl shoulder bearing about 20 low and mation in South Africa (See "Discussion" underD. sier-
narrow axial ribs, and with numerous spiral ribs over the rana sp. nov.). The new species differs from D. pseu-
entire body whorl. dorigida by being narrower just anterior to the concave
ramp, by having narrower and more numerous narrower
Description: Shell large, up to 13 cm high (estimated);
axial ribs, andby having weaker spiral ribs onthe anterior
fusiform; spire ofmedium height, abouttwo-fifths oftotal
half of the body whorl.
height of shell; spiral angle about 50°. Protoconch not The type locality ofDeussenia californiana in the Gar-
preserved. Whorls 6 1/2 (estimated). Spire whorls tabu-
apito Creek area in the eastern Santa Monica Mountains,
late, steep-sided with axial ribs stronger than spiral ribs.
Los Angeles County, southern California, plots in carto-
Posterior part of body whorl constricted to a moderately
graphic unit "Kss" (unnamed Upper Cretaceous strata)
broad and lowly concave ramp, with spiral ribs weaker
ofDibblee's (1992) map, which is the most recently pub-
on ramp than on area ofgreatest inflation ofwhorl. Body lished geologic map of the region. Rocks belonging to
whorl elongate, with greatest inflation from rounded
unit "Kss" in the Temescal Canyon-Santa Ynez Canyon
shoulder to medial part of whorl. Body whorl sculptured
just southeast of Garapito Creek were assigned by Col-
by about 20, narrow and widely spaced axial ribs, becom- burn (1996) to the Upper Cretaceous Tuna Canyon For-
ing obsolete toward medial part of whorl, and numerous mation ofYerkes & Campbell (1979). This formationwas
and closely spaced spiral ribs over entire body whorl,
deposited by turbidity currents on submarine fans (Yerkes
persisting onto the ventral surface ofthe neck. Axial ribs & Campbell, 1979, 1980; Dibblee, 1992), and unit "Kss"
more prominent than spiral ribs on posterior halfofbody
represents a dominantly sandy facies. Unit "Kss" in the
whorl. Aperture elongate-lenticular, anterior end twisted region of Garapito Creek corresponds to "member D"
to left. Columella smooth. mentionedby Popenoe 1973) andtothe so-called "upper
(
Dimensions of holotype: Height 11 cm (tip of spire and Chico" Formation utilized by Carey & Colburn (1978).
extreme anterior end both missing); width 5.8 cm. Popenoe (1973:26-27) reported that "member D" rocks
Holotype: LACMIP 12718. were probably deposited as turbidities and that the mol-
lusk fossils are shallow-marine forms that might have
Type locality: CSUN loc. 153, latitude 34°07'N, longi- been transported from theirregularhabitat into somewhat
tude 118°34'W. deeper water. Carey & Colburn (1978) reported that these
Paratype: LACMIP 12719. same rocks represent middle-fan channelized turbidites
containing lenses ofconcentrated shallow-marine moUus-
Distribution: TunaCanyon Formation, south forkofGar- can shells that appear to have been transported.
apito Creek, eastern Santa Monica Mountains, Los An- The paleoenvironment of the Tuna Canyon Formation
geles County, southern California. closely resembles that of the Chatsworth Formation of
Colburn et al. (1981) in the Simi Hills just to the north
Geologic age: Late Cretaceous (late middle to early late
Campanian) = Metaplacenticeras pacificum ammonite of the Santa Monica Mountains. According to Dibblee
(1992), the Tuna Canyon Formation is probably equiva-
zone.
lent to the Chatsworth Formation in the Simi Hills.
Discussion: Only two specimens have been found, and Based on the presence of the ammonite Metaplacenti-
both are internal molds. The paratype is larger (estimated ceras pacificum sensu stricto (Smith, 1900), Popenoe
total height 13 cm, width 6.5 cm), but most of its spire (1973) assigned a late Campanian age to the rocks at
is missing. The paratype shows spiral ribbing on the neck CSUN loc. 153. Subsequent detailed collecting at this
of the body whorl (Figure 9), whereas on the holotype type locality of D. californiana yielded this ammonite,
the spiral ribbing in this area is not preserved. as well as the gastropods Anchura phaba Elder & Saul,
Deussenia californiana is similar to Deussenia cibolo- 1996, Volutodernia magna Packard, 1922, and Zinsitys
ensis Stephenson (1941:332-333, pi. 64, figs. 13, 14; Ak- kingi (Gabb, 1864); the bivalves Crassatella elongata
ers & Akers, 1997:figs. 183-184) from the Upper Cre- Anderson, 1958, Indogrammatodon sp., Cucullaea sp..
taceous Kemp Clay [also referred to as the Kemp For- Pinna sp., (closed valved), Pterotrigonia evansana
mation] in eastern Texas. Modem workers (e.g., Sohl, (Meek, 1858), Inoceramus sp., Clisocolus dubius (Gabb,
1964:fig. 12; Elder, 1996) correlated the Kemp Clay to 1864), an isognomid, and a venerid. The presence oiMe-
the upper Maastrichtian Stage. The new species differs taplacenticeraspacificum sensu stricto is very age diag-
from D. ciboloensis by having a much less tabulate body nostic because this species constitutes the Metaplacenti-
whorl shoulder, no tubercules on the body whorl shoulder, ceraspacificum ammonite zone (after Matsomoto, I960),
and narrower spiral ribs on the body whorl. which is of middle to early late Campanian age (Elder &
&
R. L. Squires L. R. Saul, 2000 Page 123
Saul, 1996). The geologic ranges of both Zinsitys kingii shore ofLake Nacimiento, San Luis Obispo County, cen-
and Anchura phaba are correlative to the M. pacificum tral California.
&
zone (Saul, 1988; Elder Saul, 1996).
At LACMIP loc. 27002. which is in the general area Geologic age: Late Cretaceous (latest Maastrichtian) or
ofCSUN loc. 153, the bivalves Glycymeris (Glycymerita) possibly earliest Paleocene.
veatchii (Gabb, 1864), Pterotrigonia evansana (Meek,
1858), Cytnbophora triangidata (Waring, 1917), Calva Discussion: Only a single specimen has been found. It is
small in size and could be ajuvenile form.
sp., and Yaadia sp. were also found. The exact location
of LACMIP loc. 27002 is not known, but is undoubtedly Taliaferro (1944) referred the Dip Creek strata to his
in the immediate area of CSUN loc. 153. "Dip Creek Formation," but Durham (1968) referred to
The mollusks at CSUN loc. 153 must have undergone them as unnamed Upper Cretaceous and lower Tertiary
post-mortem transport from shallow-water sites into rocks. At Dip Creek, the mollusks are shallow-marine
deep-water, submarine-fan paleoenvironments ofthe Tun- forms that have undergone post-mortem transport and are
ca Canyon Formation. Several of the species found at within deep-waterturbidites inbeds ofcoarse-grainedgrit
CSUN loc. 153 or in the immediate vicinity (e.g.,Anchu- or conglomerate (Grove, 1986). More detailed geologic
ra phaba, Volutoderma magna, Crassatella elongata. mapping is needed in the area before the Dip Creek sec-
Cymbophora triangidata, Glycymeris (Glycymerita) veat- tion can be assigned to a formation. The outcrops along
chii, Pterotrigonia evansana, and Pinna sp.) were nor- Dip Creek are usually covered by waters behind the Lake
mal-marine, shallow-depth dwellers (10to50m) thatalso Nacimiento dam but are exposed during drought years
have been reported as transported remains in bathyal sub- (Squires & Saul, 1993). One can collect fossils along the
marine-fan deposits of Campanian age in the Chatsworth nearby ridge top, but these specimens are harder to find
Formation in the Simi Hills (Saul & Alderson, 1981). and are more poorly preserved than those along the lake
shore.
Etymology: The species is named for the state of Cali- The Dip Creek fauna contains some mollusks that re-
fornia. semble genera or species usually considered to indicate a
& Cretaceous age, as well as some indicative ofaPaleocene
Deusseniapacificana Squires Saul, sp. nov. age. Taliaferro (1944) reported an unidentified ammonite
(Figures 10-12) from the fauna, and Saul (1983) reported a fragment of
&
another ammonite, probably a Neophylloceras. Kirby
Deussenia? n. sp. Saul. 1986:figs. 54-55. Saul (1995) reported a fragment ofthe bivalve Roiidaria.
Diagnosis: A medium-shelled species ofDeussenia with These fossil remains suggest that at least the lower part
a subtabulate body whorl shoulder, bearing about 11 mod- ofthe section is ofvery Late Cretaceous age and that the
erately strong axial ribs, and with prominent, closely upper halfofthe section, where the new species was col-
spaced spiral ribs on body whorl. lected, is no younger than earliest Paleocene. Based on
specimens of Tiirritella peninsularis adelaidana (Merri-
Description: Shell medium in size, 31 mm high (incom- am, 1941) and T. webbi (Saul, 1983), Saul (1983) as-
plete); fusiform; spire ofmedium height, about two-fifths signed the Dip Creek mollusks to a latest Maastrichtian
oftotal height ofshell; spiral angle about 55°. Protoconch and possibly an earliest Paleocene age. This age assign-
not preserved, and spire sculpture not preserved. Whorls ment was followed by Saul (1986) and Squires & Saul
about 4 1/2 (estimated). Upper part of body whorl con-
(1993).
stricted somewhat to broad, slightly concave ramp. Body
whorl with about 11 (estimated) moderately strong and Etymology: The new species is named for the Pacific
widely spaced axial ribs; strongest on shoulder and be- Ocean.
coming obsolete toward base of whorl. Body whorl with
prominent, closely spaced spiral ribs persisting onto neck Acknowlegments. Richard A. Flory (CSUC) donated two spec-
area. Aperture elongate-lenticular. Columella smooth. imens ofD. sierrana to the LACMIP collection. James W. Hag-
gart (Geological Survey of Canada. Vancouver. British Colum-
Dimensions ofholotype: Height 31.5 mm, width 16 mm bia) facilitated our obtaining of specimens from the Pentz area.
(incomplete at both extremities, especially the anterior Lindsey T. Groves (LACMIP) provided access to the collections
and loaned specimens. Jean DeMouthe (CAS) loanedspecimens.
end).
Anton Oleinik (Department of Geology. Purdue University, In-
Holotype: CASG 61598.01 [ex Stanford University diana) kindly shared his knowledge ofbuccinid gastropods. Mi-
chael R. Cooper (Department ofGeology. University ofDurban-
30031].
Westville, Durban, South Africa) graciously provided informa-
Type locality: CASG loc. 61598, latitude 120°55'40"N, tion on the type section of the Mzamba Formation, Pondoland.
longitude 35°43'45"W. Transkei, South Africa. The manuscript benefited by reviews
from David T. Dockery III (Mississippi Office ofGeology) and
Distribution: Unnamed formation at Dip Creek, south an anonymous reviewer
—
Page 124 The Veliger, Vol. 43, No. 2
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ume and Guidebook: Los Angeles, California. Popenoe, W. p. 1973. Southern California Cretaceousformations
DDiaBvBiLeAGGCEseeaE,oonT,lroeaogg.xTgeatyM-.A.WPBl.aol,(rVoerokknelJv.r(fi&.sosore1uU1.dt9Onhbi9Twl2yhfi1.i/eFn2e.G)lEdLeC.qtooduPlF.maao:apdlgmroaLieaesocsnoingn)tdltm.ieooaolsn1npo,.9g7iL1o5oso.ff7ts1sTtTeAhpaerenprtng.itdTeairaloSyreptysaFundaCFeguoananuutannssatanyso,.df RennibSnabeuaono,nordmtknia:Jcf.&aMLuPVooAn.a.snalsieLEAoc..nnwagtiFoe1rtMl9lihoo3etug0ssei.nc,ssthpteaCNesaic(elnieaiswadfns.lo.dr)LPrn.aMpiemPi.afea.necler1ilre5fian-ilcbc2oer9gaS,ientsccoptthisiisto.hanFe,a1al-SSnl3iodcmFiiiiGneeatlHLysditloPGrlfsuoEipCacdoonoedl--d-a
California. Dibblee Geological Foundation Map DF-35. from the UpperCretaceous ofPondoland (with an appendix
Durhaanmd,BDr.adLl.ey19q6u8a.drGaenoglloesg.y MofonttheerTeiyerarandReSdaonndLauiMsouOnbtiasipno oofntshoemSeoustpheciAefsrifcraonmMtuhseeCurmeta2c8e(potu.s 1of21Z2u2l)u:l1a5n9d-)2.6A0n,napilss.
16-31.
counties, California. U.S. Geological Survey Bulletin 1255:
ElderC6,0ampWpp.a.,nPip.aisn.19A91n6-.4a.cBaicvhaolvLeismeasntdonega,stsrooutphodcsenftrraolmTtehxeas.miJdodulre- RRuossseePnlrbeles,rs:gJ.,NSe.G,.wS.Y1oL9r.9k2B..a2uT2hm4e&ppE.nR.cyWcaltokpiendisa. 1o9f86S.eaLsahteellsC.oniDaocrisaent
nal ofPaleontology 70(2):247-271, figs. 1-8. to early Campanian clastic shelfdeposits and molluscan as-
Eldero,f WC.oniP.ac&iaLn. Rt.hrSoauuglh.M1a9a9s6t.riTcahxtioannomAnycahnudrabio(sGtarsattriogproadpah:y snieam.blPpa.ges17o9f-1t9h6e ninorRtheLa.stAebrbnotStac(reda.m)e.nCtroetVaacleleoyu,sCSatlriaftoirg--
Aporrhaiidae) ofthe North American Pacific slope. Journal raphy Western North America. Pacific Section. Society of
ofPaleontology 70(3):381-399. figs. 1-6. Economic Paleontologists and Mineralogists, Book 46: Los
Fritsche, a. E. 1973. MapofCretaceous sedimentary exposures Angeles, California.
in the eastern Santa Monica Mountains. Map (in pocket) in Saul, L. R. 1983. Turritella zonation across the Cretaceous-Ter-
I. P. Colburn & A. E. Fritsche (eds.). Pacific Section,Society tiary boundary, California. University of California Publi-
of Economic Paleontologists and Mineralogists Fall Field cations in Geological Sciences 125:1-164, pis. 1-7.
Guidebook: Los Angeles, California. Saul, L. R. 1986. Mollusks of latest Cretaceous and Paleocene
GradsV.teViene,n,F.J.M.T,hiFerPr.yAg&teZb.eHruga,nJg..G.199O5g.g.AJT.riHaasrsidce,nJbuorlas,siPc. aKg.e.GrLoavkee&NaSc.imGireanhtao,m (Ceadlsi.f)o.rnGieao.loPpg.y2o5f-U3p1p,efrigCsr.et1a-c6e0ouisn
and Cretaceous time scale. Pp. 95-121 in W. A. Berggren, and Lower Tertiary Rocks Near Lake Nacimiento. Califor-
D. V. Kent, M. -P. Aubry & J. Hardenbol (eds.), Geochro- nia. Pacific Section, Society of Economic Paleontologists
nology Time Scales and Global Stratigraphic Correlation. and Mineralogists, Volume 49: Los Angeles, California.
SEPM (Society for Sedimentary Geology) Special Publica- Saul, L. R. 1988. New Late Cretaceous and Early Tertiary Per-
tion 54: Tulsa, Oklahoma. issityidae (Gastropoda) from the Pacific slope of North
Greyling, E. H.—1992. The Mzamba Formation (Cretaceous) of America. Natural History Museum ofLos Angeles County,
the Transkei stratigraphy, sedimentology and palaeontolo- Contributions in Science 400:1-25, figs. 1-128.
gy. Master of Science Thesis, University of Durban-West- Saul, L. R. & J. M. Alderson. 1981. Late Cretaceous Mollusca
ville. Durban, South Africa. ofthe Simi Hills: an introduction. Pp. 29-42, pis. 1-3 in M.
Grove, K. 1986. Field trip boatlog: depositional environments H. Link, R. L. Squires & \. P Colburn (eds.), Simi Hills
&
R. L. Squires L. R. Saul, 2000 Page 125
Cretaceous Turbidites, Southern California. Pacific Section, tion. Age: Late Cretaceous (latest Maastrichtian) or
SocietyofEconomic PaleontologistsandMineralogists,Vol- possibly earliest Paleocene. Collector: N. L. Taliaferro.
SOHL,muamNte.opaEonhd1o9rG6au4i.fdreNobemooogtkah;setLrRooipspoledAyan,,geOlOwepsil,stCChraoelbeirkfao,nrncaihnaid.aParnaidriBeaBsloumf-f CSeUasCt oPfN3H1iN.ghWAwtayel7e0v,at7i1o6n omf(5233050ft.f,t.)inSaansdma6ll9 mcan(y2o2n5
Eormations. U.S. GeologicalSurveyProfessionalPaper331- ft.) E of corner of section 31, T. 21 N, R. 4 E,
B:1-344, pis. 19-52. U.S. Geological SurveyCherokeequadrangle (7.5 min-
Squires, R. L. & L. R. Saul. 1993. A new species of Otostoma ute, 1970), Butte County, northern California. Chico
(Gastropoda: Neritidae) from near the Cretaceous/Tertiary Formation, Pentz Road member (informal) of Russell
boundary at Dip Creep, Lake Nacimiento, California. The et al. (1986). Age: Early Campanian. Collector: R.
Veliger 36(3):259-264, figs. 1-4. Watkins.
&
SQureES, R. L. L. R. Saul. 1997. Late Cretaceousoccurrences CSUC PN32. At elevation of 530 ft., in a small canyon
og1an9s3tt-rh2oe0p2oP,dacfigifgeisn.cus1s-lB1o7op.eggsofiaNoOrltsshonA,me1r9i2c9.a TohfethVeelmiegleran4o0p(s3i):d e(a3s5t0offt.)HWighowfaSyE70c,om7e5r4omf s(e2c4ti7o5nf3t.6),NT. a2n1dN,10R7.m3
Stephenson, L. W. 1941. The larger invertebrate fossils of the E, U.S. Geological Survey Cherokee quadrangle (7.5
Navarro GroupofTexas (exclusiveofcoralsandcrustaceans minute, 1970), Butte County, northern California. Chi-
andexclusiveofthefaunaoftheEscondidoEormation).The co Formation, Pentz Road member (informal) of Rus-
University ofTexas Publication 4101:1-641, pis. 1-95. sell et al. (1986). Age: Early Campanian. Collector: R.
Taliaferro, N. L. 1944. Cretaceous and Paleocene ofSantaLu- Watkins.
cia Range, California. Bulletin ofthe American Association CSUN 153. [= LACMIP 11975.] At elevation of 1450
Wardom,fagPRneettDri.o,cleaKu.nmoLmG.aeloVyleorg3io3ss-tu3s4b2i8&d(e4nJ)t.i:f4Wi4.e9d-Hi5an2g1tg.haerUtp.pe1r98C3r.etMaacreionues f(t1.,75i0n bfto.)ttSomanodf5s2o1utmh f(o1r7k1o0fftG.a)rEapioftothCereienkt,ers5e3c3tiomn
ofthe Great Valley Sequence ofCalifornia. Geology 11:90- of the San Bernardino base line and Los Angeles City
boundary, U.S. Geological Survey Topangaquadrangle
93.
Wenz, W. 1943. FamiliaGaleodidae. Pp. 1211-1224, figs. 3452- (7.5 minute, 1952, photorevised, 1967), Santa Monica
3479 in O. H. Schindewolf (ed.), Handbuch der Palazoolo- Mountains, Los Angeles County, southern California.
gie. Band 6, Prosobranchia, Teil 4. Gebriider Borntrager: Unnamed strata. Age: Late middle Campanian. Collec-
Berlin, [reprinted 1960-1961] tor: John Alderson, 1974-1987. [See Fritsche's (1973)
Woods, H. 1906. The CretaceousfaunaofPondoland. Annalsof map for aplot ofthis locality on adetailed topographic
Yerktehse,SRo.utEh&AfRr.icHa.nCMaumspebeulml.4(719)7:92.45S-t3r5a0t,igrpiasp.hi3c3-n4o4m,encla- LAbCasMeI.]P 10833. Fossiliferous layers cropping outinbeds
ture of the central Santa Monica Mountains, Los Angeles
County, California. U.S. Geological Survey Bulletin 1457E: of small gullies in field along Pentz Roamd [formerly
E1-E31. Durham-Pentz Road], approximately 290 (950 ft.) S
Yerkes, R. E & R. H. Campbell. 1980. Geologic map of east- and 107 m (350 ft.) E of NW corner of section 25, T.
central Santa Monica Mountains, Los Angeles County, Cal- 1 N, R. 3 E, U.S. Geological Survey Cherokee quad-
ifornia. U.S. Geological Survey Map 1-1146. rangle (7.5 minute, 1970), Butte County, northern Cal-
ifornia. Chico Formation, Pentz Road member (infor-
APPENDIX mal) of RusWsell et al. (1986). Age: Early Campanian.
Collector: P Popenoe, 1931.
LOCALITIES CITED LACMIP27002. South fork ofGarapito Creek, U.S. Geo-
logical Survey Topanga quadrangle (7.5 minute, 1952,
CASG 61598. Dip Creek, NE 1/4 of section 30, T. 25 S, photorevised, 1967), Santa Monica Mountains, Los
R. 10 E, U.S. Geological Survey Lime Mountainquad- Angeles County, southern California. Unnamed strata.
rangle (7.5 minute, 1948, photorevised 1979), San Luis Age: Late middle Campanian. Collector: Mike Ham-
Obispo County, central California. Unnamed forma- ilton, circa middle 1970s.