The bamboos of the world: annotated nomenclature and literature of the species and the higher and lower taxa

578 Pages·1999·11.88 MB·English

by  D Ohrnberger

Preface
It is a pleasure for me to present the new completely revised
edition of ehT Bamboos of the World. It is the first publication
which combines and treats all currently known bamboos in a
single inventory.
The plan of a preliminary inventory study on bamboos arose
prior to 1981. The work was originally done by J. Goerrings and
myself, sometimes with the help of botanists, and later continued
by myself. From 1983 onwards, much of the work was published
in separate parts. For a survey of all these parts of ehT Bamboos
of the World, the reader is referred to the appendix of the current
work. However, all the previous parts are replaced by the present,
much enlarged and updated edition.
I am aware of the fact that the current edition may not be free
of errors and needs corrections, especially more checking than it
has received hitherto by specialists on bambusoid grasses. I would
be grateful for any hints and suggestions that could improve ehT
Bamboos of the World.
This work would not have been possible without the generous
help of numerous people. I like to express my sincere thanks to all
the unselfish people who contributed to the work up to the
present.
Berlin and Langweid a. L., July 1998
Dieter Ohrnberger
INTRODUCTION
2 THE BAMBOOS OF THE WORLD
General Aims of ehT Bamboos of the Worm
nI 1913, the French botanist and pharmacist, Ed- The dispersed and growing number of separate
mond Gustave Camus (1852-1915) Les Bambus~es sources of information, the increasing difficulties over
published which covered almost completely the then its availability of this information, and the still more
known bamboos. Included were not only the woody complicated general survey of this group of grasses,
bamboos, but also some herbaceous bamboos. Les make it necessary to sort the main data no the bam-
Bambus~es is hardly comparable with the scientifically boos and to make ti more easily accessible. The most
more important previous monographs on bamboos by important data era considered to eb the botanical
Munro (1868) and Gamble (1896), both of which are names of the genera dna species dna their geographi-
elaborated taxonomic revisions. Considering the huge cal distribution, along with their references. With the
number of species and genera of bamboos known at help of this information it will be possible for anyone to
the beginning of the 20th century, Camus' monograph obtain, from the different sources mentioned, a tol of
could not ,eb for most of its parts, a revision but merely additional facts: plant descriptions, diagnoses dna keys
a compilation of species and literature, albeit no a for plant identification, botanical drawings, photographs
world-wide basis. of the plants, precise plant localities, vernacular names
and many more.
Camus treated a total of 33 genera and 490 species.
More than 80 years have gone since the publication of With these considerations ni mind, the following
Les Bambus~es, and about one hundred generic general aims ni ehT Bamboos of eht World have been
names, and over 2,000 specific names (species and laid down:
new combinations) have been published since then.
(cid:12)9 to present a documentary review of the main data
Among these, about 1,000 represent new species not
(names, distribution) of lla the bamboo species,
included by Camus, meaning the number of species
generally accepted has been tripled since 1913. The (cid:12)9 to facilitate access to further data by citation dna a
numerical growth of generic names since that time is comprehensive bibliography.
considerable too, especially during the last two dec-
ades. tI should be noted that ehT Bamboos of eht World is
basically intended to be a documentary or reference
The state of classification of bambusoid grasses is work, with results based essentially no literature. oN
far behind that of lla the other groups of grasses. claims have been made to present a revision of the
Basically, the major reason rests ni the peculiarity ni bamboos.
the life of most of the bamboos themselves: The
flowering of the majority of the woody bamboos is a The work is directed to people and institutions inter-
rare event, and if flowers are available the vegetative ested ni the botany, horticulture, forestry dna ecology
parts of the plants often do not develop representative of bamboos, especially their morphology, taxonomy,
characteristics during flowering. This may prevent the distribution and cultivation.
collecting of enough of the flowering and vegetative
material, necessary to understand a species and es-
Mit der Publikation, ehT Bamboos of eht World, wird
tablish its relationships, allowing a stable classificatory
die Absicht verfolgt, nie umfassendes und aktuelles
system to be built. tI is therefore understandable that
Nachschlagewerk Uber die Gattungen und Arten der
the classification of bambusoid grasses has remained
Bambusgraser und ihre Verbreitung zu bieten. ehT
fluid, though ni the recent past much research has del
Bamboos of eht World basiert auf einer umfangreichen
towards a more stable classification ta suprageneric
Literaturauswertung. Schwerpunkt sind die derzeitige
level. The present relationships of many genera, how-
taxonomische Klassifizierung und Nomenklatur der ge-
ever, remain far from being sufficiently settled.
samten Bambus-Unterfamilie (insbesondere deren
The information about bambusoid grasses available Gattungen, Arten, Unterarten, Varietaten, Formen und
up to now is scattered through numerous monographs, Sorten), sowie die natUrliche geographische Verbrei-
Floras, and treatises ni journals. A remarkable number tung der Bambus-Arten und deren EinfUhrungen ni
of taxonomic, systematic and phytogeographic contri- den Westen. All diese Daten werden ausgewertet,
butions must also be expected to appear ni the near kompiliert und ni einer einheitlichen Form med Leser
future. All this makes it difficult to produce a general, prasentiert. Somit ist ehT Bamboos of eht World vor-
world-wide survey of the bamboos, or their classifica- wiegend yon Interesse fur Botaniker, Okologen, G~rt-
tion, even of a single genus. ner, Dendrologen und Pflanzensammler.
INTRODUCTION 3
Principals and General Notes on the Contents
To fix the size and scope of The Bamboos of the the World appear with their names in headings of bold
World, the general aims were followed as quickly as type, if not otherwise stated. Taxonomic groups not
possible, taking into consideration personal, financial accepted are printed in normal type.
and time restrictions. Apart from the main data pre-
sented, some additional information has been included.
The mapping of species distribution, included in the Codes of Nomenclature
first edition, has not been continued for the present
edition because of time and cost. Botanical names are ruled by the Botanical Code
(ICBN); cultivar names by the Cultivated Plant Code
The Bamboos of the World is aimed not only at
(ICNCP):
botanists, but also at a wider range of readers. It is
therefore intended to present the information in an (cid:12)9 International Code of Botanical Nomenclature
easily understandable arrangement. nI this section, ex- ICBN (Tokyo Code), Adopted by the Fifteenth In-
planations are given of some topics of general interest, ternational Botanical Congress, Yokohama, August-
and to some contents of the work. September 1993 / W. Greuter & .la (editors), 1994,
389 pp., K0nigstein: Koeltz, (Regnum Vegetabile,
Classification vol. 131).
(cid:12)9 International Code of Nomenclature for Cultivated
The suprageneric classification of bamboos has
Plants - 1995 (ICNCP or Cultivated Code) / oP Tre-
been considerably improved recently by the research
hane & .la (editors), 1995, 175 pp., Wimborne:
work by several botanists. A scheme of classification,
Quarterjack, (Regnum Vegetabile, vol. 133).
based essentially on these recent works, is presented
herein. For any further information on the current sys- The ICBN holds rules on the formation and applica-
tematics and classification of bamboos, the references tion of scientific names (botanical names) for taxo-
should be consulted. nomic groups (taxa pl., taxon sing.) of plants from the
wild or in cultivation. This provides an internationally
The generic classification is still in rather fluctuating
accepted system of reference. In botany, the names of
state. There are tendencies towards both a narrower
the taxa are displayed in a phylogenetic scheme of
and a wider definition of several bamboo genera by
classification. Each taxon therein has a particular
contemporary botanists, olden leading to different clas-
sifications and to name changes. The definition of circumscription, position, and rank in order to indicate
the degree of relationship. The basic unit of taxa is
many genera thus remains provisional.
called a species. Higher ranking taxa are, for instance,
Infrageneric classification has been proposed by a genus (a group of related species), and a tribe (a
botanists for some bamboo genera. If the genera con- group of related genera).
cerned were differently defined by various botanists
and have not undergone revision recently, infrageneric The ICNCP is a separate system of rules. It is con-
classification often seems to be of little help. No nected with the ICBN by the rule defining the botanical
attempt has been made in The Bamboos of the World name of a taxon in a certain rank (genus at least,
to assign all the species of a certain genus to an species as usual) to become part of the name for
infrageneric group, but in some cases established as- cultivated plants (ICNCP 1995, Art. 1.1). This implies
signations have been adopted. that a cultivar (cultivated variety, the basic unit of taxa
of cultivated plants) is always lower in rank than the
The classification of species in The Bamboos of the assigned botanical taxon (e.g. species, subspecies,
World generally follows classification proposed by con- variety, form). Cultivars are usually defined by char-
temporary botanists who have contributed to the revi- acters important to horticulture, agriculture and for-
sion of the whole group (genus). estry. Such characters (e.g. a differing colour of leaves
and culms) appear as variations within a species, and
For infraspecific classification, the same procedure
are inappropriate characters for the delimination of
as stated for species classification applies. Generally,
species.
variants of mere horticultural importance have been
classified as cultivars, or ranked down from variety to There is one basic difference in the method of
form. formation of names between the ICBN and ICNCP.
The application of a name of a botanical taxon is deter-
Genera and Species Accepted mined by a nomenclatural type (a certain herbarium
specimen), whereas the name of a taxon of a cultivar
Genera and species, and other taxonomic groups, need not be determined by a type; a valid description
accepted or provisionally accepted in The Bamboos of alone is sufficient to establish a cultivar name.
4 THE BAMBOOS OF THE WORLD
Botanical and cultivar names adopted in ehT Bam- Validly Published Names
boos of eht World have been checked for adherence to
the rules of the Codes wherever possible. Names not Names published in accordance with the rules of the
checked are those adopted from citation only, where Codes of nomenclature are called "valid" (for botanical
the original publications of the names concerned have names) and "established" (for cultivars). The Botanical
not been available. These names are especially those Code divides validly published names into legitimate
of forms and cultivars which have appeared in journals names (those in accordance with the rules), and illegiti-
or books of limited circulation. mate names (nom. illeg.). Illegitimate names are those
which are superfluous when published, and often later
homonyms. Homonyms are identical botanical names
Ranks of Taxa based on different nomenclatural types.
Every individual plant is treated as belonging to taxa Out of the legitimate names there is only one name
of consecutively higher rank. The following table dis- (for a taxon with a particular circumscription, position,
plays the ranks of the taxa (from family down). Basic and rank) that must be adopted under the rules of the
unit is the species. Botanical Code. This name is termed the "correct"
rank term (Latin) rank term rank term example
(English) (German)
familia (fam.) family Familie POACEAE
subfamilia (subfam.) subfamily Unterfamilie BAMBUSOIDEAE
supertribus supertribe Supertribus BAMBUSA TAE
tribus (trib.) tribe Tribus BAMBUSEAE
subtribus (subtrib.) subtribe Untertribus ARUNDINARIINAE
tribella tribella Tribella A R UNDINA AIR S TRAE
genus (gen.) genus Gattun9 Arundinaria
subgenus (subg.) subgenus Untergattung Pleioblastus subg. Nipponocalamus
sectio (sect.) section Sektion Far~esia sect. Sphaefi~emma
subsectio (subsect.) subsection Untersektion P/eioblastus sect. Nipponocalamus subsect. Nezasa
series (ser.) series Serie P/eioblastus sect. Medakea ser. Dolicho/igulae
species (sp.) species Art Indosasa triangulata
subspecies (subsp.) subspecies Unterart Otatea acuminata subsp, aztecorum
varietas (var.) variety Varietat Sinobambusa tootsik var. laeta
forma (f.) form Form Phyllostachys nigra .f henonis
('-.,9 cultivar Kultivar (Sorte) Bambusa multiplex 'Golden Goddess'
Formation and Format of Names name of a certain taxon governed by the Botanical
Code, and the "accepted" name if governed by the
Names at the rank of genus and higher consist of Cultivated Plant Code. tI appears typographically in
one word. The rank of a name higher than generic rank bold type. The other validly published names applied to
is discernible by its ending, e.g. "-aceae" for family, the same taxon are called synonyms.
"-oideae" for subfamily, "-eae" for tribe. The typo-
In ehT Bamboos of eht World the attempt has been
graphic Italic type and capitalising of letters serve to
distinguish names more easily and are not compulsory. made to include all the valid botanical names of the
bamboos ever published, from subfamily down to
Names at the rank of species consist of two parts forma. At species and higher level, the listings have
(binominal), a generic name and an epithet. Botanical achieved near-completeness, except for the latest pub-
names from genus downwards are preferably printed lications of names if they have not yet come to the
italicised. author's attention. The listings of names at infraspe-
cific and cultivar level are comprehensive but remain
Botanical names at the rank lower than species
incomplete.
consist of three (or more) parts, with a rank-denoting
term. The rank-denoting term is usually not italicised.
Invalidly Published Names
Names at cultivar rank consist of a botanical name
(usually species name but at least generic name) and Only validly published names in accordance with the
the cultivar epithet, the latter often termed as cultivar Botanical Code are of nomenclatural relevancy. Inval-
name or cultivar. The cultivar epithet is to be enclosed idly published names are of no importance. Neverthe-
by single quotation marks, and not printed italicised. less, they have been included, and invalid publication is
The former usage of the rank term "cultivar", abbrevi- always indicated, usually by adding "invalid", "as syn.",
ated "cv.", is no longer sanctioned by the current Code. "nom. nud." to the reference.
INTRODUCTION 5
Cultivar names not established in accordance with Culm Size and Flowering Record
the Cultivated Plant Code are also included and simi-
larly indicated. Data are given for culm size and flowering records if
available. These data are presented in a standardised
The major, and usually the only, reason for including form as follows: Culm height in m / Culm diameter in
invalid names is so that their nomenclatural status be- cm /Inflorescence (fl) known (+) or not known (-), e.g.:
comes evident.
5- 10 (12) m / 1 - 5 cm/fl(-)
Measurment enclosed in brackets indicate maximum
Abbreviations in Bibliographic Citation or minimum size. The plant size refers to plants in
natural environment, usually not under cultivated con-
Bibliographic citation is usually given abbreviated: ditions. A question mark is given if either culm height
name of author(s), year of publication, and page num- or diameter is not recorded, e.g.:
ber; e.g.
0.3 - 0.9 m / ? cm / fl(+)
Nakai, 1933: 20;
or given together with the abbreviated title and vol-
ume numbering of publication, e.g. Distinctive Characters
Nakai in J. Jap. Bot. 9 (1), 1933: 20. Distinctive characters of subspecies, varieties,
An asterisk indicates illustration. For full and non- forms, and cultivars are usually stated. For characters
abbreviated references the reader is referred to the of species and other taxa, and for descriptions and
Bibliography. keys, the reader is referred to literature cited.
Geographical Distribution
Taxonomic References
The geographical distribution of species and genera
Taxonomic references are those referring to a cer-
is stated, generally by political units: country, and in
tain botanical name of a taxon, not necessarily referring
most cases by the next lower unit, e.g. province.
to a certain taxon that might be differently defined by
Usually the natural geographical distribution is stated,
various authors. The nomenclatural type is the basic
naturaUsed and cultivated distribution is given only if
element that rules the application of names. For this
the natural distribution is unknown or obscurely known.
reason, it is cited (in certain cases only, not through-
Altitudinal distribution is stated if known, especially for
out); e.g.
montane bamboos. Data on habitat and ecology are
Fargesia declivis Yi in J. Bamb. often not available but if so, they will be summarised.
.seR 7 ,)2( :8891 101, fig. 31; type:
The geographical distribution should not be inter-
Yunnan, T.P. Yi 77315 (SCFS);
preted without considering the relevant classification.
The above example for the nomenclatural type of Especially for genera, it must be kept in mind whether
Fargesia declivis means: The herbarium specimen no. a certain genus had been recently revised or still awaits
77315 was collected in Yunnan by T.P. Yi, and is revision. In the latter case, which applies quite often,
deposited in the Herbarium of the Sichuan Forestry the geographical distribution is merely provisional. It
School. should also be taken into account whether a certain
genus or species is narrowly defined or not. Genera
and species may be differently circumscribed by
various authors, thus the pattern of distribution may
Selected References
vary.
Apart from taxonomic and nomenclatural references,
selected references are given in some cases. Such re-
Horticulture in Europe and the USA
ferences usually provide a modern treatment of the
taxon concerned, e.g. with plant descriptions, identifi-
Numerous species of bamboo, mostly their horticul-
cation key, illustrations.
tural varieties, have been introduced from their native
countries to Europe and the USA, to Europe as early
as the first half of the 19th century. If it is known that a
Non-Botanical Names certain bamboo is in cultivation in Europe and the USA,
the information is given, along with the year or decade
Common and vernacular names are occasionally
of its first introduction. The correct identification of
given, and if so, rather selectively. If available, Chinese,
bamboos in Western cultivation is often in doubt, and
Japanese, and English names are always adopted.
some records should therefore considered uncertain.
Common, local and vernacular names often do not
refer to a single bamboo species but may be applied to The frost tolerance of hardy species is given, prefer-
several species as well. Identification of bamboos ably when known from long-term observation of several
should therefore never be based on such names. established stands (without unusual environmental
6 THE BAMBOOS OF THE WORLD
factors, and without winter protection) in Germany. As its ornamental value. The range of temperature that
bamboos in Western cultivation are planted almost causes damage to leaves is therefore the more appro-
exclusively for ornament, damage to leaves in winter priate indicator for hardiness than rates for rhizome
caused by low temperatures and other meterological survival when culms have died from frost.
factors impairs the plant's appearance and diminishes
SUBFAMILY
BAMBUSOIDEAE
8 THE BAMBOOS OF THE WORLD
Tribes, Subtribes and Genera
in Systematic Order
The classification here is limited to grasses con- The scheme here includes 6 tribes, 10 subtribes
sidered bambusoid by modern agrostologists. Recent and 111 genera accepted, or provisionally accepted, in
phylogenetic research (L.G. Clark & al., 1995; L.G. ehT Bamboos of the World.
Clark & E.J. Judziewicz, 1996) has shown that some
groups hitherto considered bambusoid are not. These (cid:12)9 Selected taxonomic references:
genera require a position outside the bambusoids: C.E. Calderbn & Soderstrom in Smithson. Contr. Bot.
Anomochloa, Streptochaeta, Pharus, Leptaspis, Scro- no. 44, 1980: 1-27; W.D. Clayton in Ann. Missouri Bot.
tochloa and Suddia. Gard. 68 (1), 1981: 5-14; W.D. Clayton & Renvoize,
Gen. Gram., 1986: 1-375; Soderstrom & Ellis ni Soder-
Furthermore, the genus Streptogyna is now thought
strom & al., Grass Syst. Evol., 1987 1988: 225-238;
to be closer related to the oryzoid group (which is con-
Tsvelev in Komarov. Chten. Akad. NAUK Leningrad
sidered to represent a separate subfamily) than to the
37, 1987: 1-75; Tsvelev in Bot. Rev. 55 (3), 1989: 141-
bambusoids, and is therefore excluded from the latter.
204; .L Watson & Dallwitz, Grass Gen. World, 1992,
No recent phylogenetic research and re-evaluation is 1038 pp.; L.G. Clark & .la in Syst. Bot. 20 (4), 1995:
known for the African genera Pue/ia and Guadue//a, 436-460; L.G. Clark & E.J. Judziewicz in Taxon 45 (4),
and they are therefore kept within the bambusoids. 1996:641-645
Tentative Scheme of Related Groups Among the Bambusoid Grasses
With Approximate Number of Species in Each
family POACEAE (GRAM/NEAE), divided into 7 to 10 subfamilies, among them the
subfamily BAMBUSOIDEAE (comprising both woody and herbaceous bamboos with altogether 1575 species), divided
into 1 tribe of woody bamboos (BAMBUSEAE, this divided into 10 subtribes) and 5 tribes of herbaceous bamboos
as follows:
,sebirT sebirtbuS areneG
BAMBUSEAE (1447)
ARUNDINARIINAE (249) Acidosasa (22), Arundinaria (1), Bashania (6), Ferrocalamus (2), Gaoligong-
shania (1), Ge/idocalamus (12), Indocalamus (35), Menstruocalamus (1),
Metasasa (2), Oligostachyum (18), P/eioblastus (42), Polyanthus (1), Pseudo-
sasa (36), Sasa (58), Sasaella 11( ,) Vietnamocalamus (1)
THAMNOCALAMINAE (226) Ampe/oca/amus 11( ,) Borinda (8), Chimonocalamus (16), Drepanostachyum 11( ,)
Fargesia (83), Himalayaca/amus (9), Thamnocalamus (4), Yushania (84)
RACEMOBAMBOSINAE )72( Neomicrocalamus (7), Racemobambos (17), Vietnamosasa (3)
SHIBA TAEINAE (182) Brachystachyum (1), Chimonobambusa (38), Hibanobambusa (1),/ndosasa (27),
Phy//ostachys (76), Semiarundinana (6), Shibataea (9), Sinobambusa (24)
BAMBUSINAE (297) Bambusa (139), Bonia (5), Dendrocalamus (52), Dinoch/oa (27), Giganto-
chloa (37), Holttumochloa (3), Kinabaluchloa (2), Klemachloa (1), Mac/uro-
ch/oa (1), Me/ocalamus (9), Oreobambos (1), Oxytenanthera (1), Pseudo-
bambusa (1), Pseudoxytenanthera (12), Soejatmia (1), Sphaerobambos (3),
Thyrsostachys (2)
MEL OCANNINAE (87) Cephalostachyum (16), Davidsea (1), Dendrochloa (1), Melocanna (2), Neo-
houzeaua (7), Ochlandra 11( ,) Pseudostachyum (2), Schizostachyum (45),
Teinostachyum (2)
HICKELIINAE (37) Decaryochloa (1), Gres/ania (4), Hickelia (4), Hitchcockella (1), Nastus (24),
Perrierbambus (2), Temburongia (1)
GUADUINAE (40) Criciuma (1), Eremocau/on (1), Guadua (34), Olmeca (2), Otatea (2)
CHUSQUEINAE (156) Chusquea (135), Neuro/epis (21)
SUBFAMILY BAMBUSOIDEAE 9
BAMBUSEAE (continued)
ARTHROSTYLIDIINAE )641( mudalcon#cA (1), Alvimia (3), Apoclada (3), Arthrostylidium (28), Athroo-
stachys (1), Atractantha (5), aimeno/uA (32), ai/ehtna/oC (7), E/ytrostachys (2),
notyhpoiza/G (1), Merostachys (34), suda/coiryM (13), muda/codipihR (16)
LO YREAE (76) Agnesia (1), Arberella (7), aolhcotpyrC (9), Diandrolyra (2), Ekmanoch/oa (2),
aolhcoiseorF (1), Uthachne (4), ary/mu/caM (1), Mniochloa (1), O/yra (23),
ary/oidoraP (3), Piresia (4), alleiseriP (1), Raddia (5), alleiddaR (7), Rehia (1),
aiztieR (1), Sucrea (3)
PARIANEAE )04( Eremitis (1), Pariana (39)
BUERGERSlOCHL OEAE ( 1 ) Buergersiochloa )1(
PUELIEAE (5) aileuP (5)
GUADUELLEAE (6) Guaduella (6)
Tribes, Subtribes and Genera
in Alphabetical Order
The tribes, subtribes, and genera ni the previous Genera
chapter are given here ni alphabetical order. The most
important taxonomic synonyms are added ni brackets. asasodicA .C .D UHC & .C .S OAHC XE .P .C GNEK
mudalconJtcA ERULCCM XE MORTSREDOS
aisengA AGAOLUZ & ZCiWEiZDUJ
Tribes aimivlA .C .E NOREDLAC XE MORTSREDOS & ONODNOL
sumalacolepmA .S .L CriER, NEW & G. Y. GNEHS
BAMBUSEAE HTNUK XE SEEN sumalacolletaP( W. T. )NIL
BUERGERSIOCHLOEAE .S T. EKALB adalcopA ERULCCM
GUADUELLEAE MORTSREDOS & SILLE allerebrA MORTSREDOS & .C .E NOREDLAC
LO YREAE HTNUK XE RENNEPS muidilytsorhtrA THCERPUR
PARIANEAE .C .E DRABBUH airanidnurA XUAHCIM
PUELIEAE MORTSREDOS & SILLE A( -OTCART Athroostachys MAHTNEB
CARPEAE )SUMAC( XIL~IF-SEUQCAJ XE )VELEVST ahtnatcartA ERULCCM
aimenoluA TODUOG (Matudacalamus .F )AWAKEAM
Subtribes Bambusa REBERHCS (Dendrocalamopsis AILIC( &
.H .L )GNUF .P .C ,GNEK Ischurochloa ,ESUB abeleL
ARTHROS EANIIDILYT SWEB SUIHPMUR XE ,IAKAN Lingnania ,ERULCCM
ARUNDINARIINAE MAHTNEB (SASAINAE .P .C sumalaconisoeN .P .C ,GNEK sumalaconogarteT
)GREK )iAKAN
BAMBUSINAE .J .S LSERP (DENDROCALAMINAE ainahsaB .P .C GNEK & tY
)MARTREB Bonia ASNALAB (Monocladus CHIA, .H .L GNUF & Y. .L
CHUSQUEINAE SWEB (NEUROLEPIDINAE REDOS o )GNAY
MORTS & )SILLE adniroB NOTELPATS
GUADUINAE MORTSREDOS & SILLE Brachystachyum GNEK
EANIILEKCIH A. SUMAC (NASTINAE MORTSREDOS & aolhcoisregreuB REGLIP
)SILLE muyhcatsolahpeC ORNUM
MELOCANNINAE MAHTNEB (SCHIZO- asubmabonomihC Makino (Qiongzhuea (Wen & Ohm-
STACHYDINAE ORNUM XE MORTSREDOS & )SILLE berger) Hsueh & Yi, Oreocalamus Keng)
RACEMOBAMBOSlNAE NOTELPATS sumalaconomihC HEUSH & Yi
ABIHS TAEINAE )IAKAN( MORTSREDOS & SILLE aeuqsuhC HTNUK (Rettbergia ,IDDAR aolhconellawS
(PHYLLOSTACHYDINAE .P .C )GNEK )ERULCCM
THAMNOcALAMINAE .P .C GNEK ailehtnaloC ERULCCM & .E W. HTIMS