The Asteioinea of Fiji (Insecta: Diptera: Periscelididae, Asteiidae, Xenasteiidae)

2009·28.4 MB·English

by  David A. Grimaldi

PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY
CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024
Number 3671, 59 pp., 32 figures, 2 tables November 30, 2009
The Asteioinea of Fiji (Insecta: Diptera:
Periscelididae, Asteiidae, Xenasteiidae)
DAVID A. GRIMALDI1
ABSTRACT
Three of the six families in the acalyptrate fly group Asteioinea are reported from Fiji,
comprising five genera and 23 species, with all except three of the species being new. The new
speciesintheirrespectivefamiliesarethefollowing:Stenomicraariela,Si.brunnea,Si.castanea,Si.
distincta, Si. pallida, Si. sylpha, Si. tokotaai, and Si. xoutha; Cyamops femobrunneus, C.
femoctenidius; Stenocyamops luteus, Sy. pseudoluteus, Sy. robustus, and Sy. vittatus
(Periscelididae); Asteia pleurovitta, A. pleurovittata, A. rotundiscuta, A. vanuaensis, A. vitiensis
(Asteiidae);andXenasteiafijiana(Xenasteiidae).ThisisthefirstreportofStenocyamopsoutsideof
Southeast Asia. Stenomicra distinctipennis (Collin) is resurrected from synonymy with S.
fascipennis Malloch, based on study of male genitalia belonging to a complex of fascipennis-like
species from throughout the Pacific. Asteia nigriceps Bezzi is redescribed. Keys to genera and
speciesareprovided.Somespecieswerecollectedintherolledleavesofbananasandwildgingers
(Zingerberales), which appearsto beanimportant habitat for periscelidids in tropical forests.
INTRODUCTION the Fiji archipelago. Fiji consists of approxi-
mately 322 islands and over 500 islets, lying
Thisisataxonomictreatmentofthespecies between 15u–20u S and 175u E–175u W. Four
belonging to three families of small, obscure Fiji islands have sufficient size and relief
schizophoran flies from the South Pacific to support lush tropical forests, which are
nation of the Republic of the Fiji Islands. Viti Levu (10,388 km2), Vanua Levu (5587
The three families—Periscelididae, Asteiidae, km2), Taveuni (435 km2), and Kadavu
and Xenasteiidae—belong to the superfamily (411 km2). All of the insect sampling for this
Asteioinea (McAlpine, 1989), of which the and most other insect studies has focused on
taxonomyhereisasmallcontributiontowards these four islands, since smaller islands are
understanding the arthropod biodiversity of low, drier, and support less forest. Climate is
1DivisionofInvertebrateZoology,AmericanMuseumofNaturalHistory([email protected]).
CopyrightEAmericanMuseumofNaturalHistory2009 ISSN0003-0082
2 AMERICAN MUSEUMNOVITATES NO. 3671
tropical, but because of the rugged, moun- has been described elsewhere for Stenomicra
tainous topography of the four largest islands and the closely related family Neurochaetidae
there are dramatic effects to local climate. (D.K. McAlpine, 1978, 1987a, 1987b, 1993),
Northern halves of these islands, for example, the Fiji periscelidids—and even the Asteia
have a pronounced dry season and the cohabiting with them—have a remarkable
southern halves support moist, evergreen wayofmoving.Thesefliesrunoverthesurface
forests; the higher peaks and ridges of the of a leaf in all directions with uniform effort,
central mountains are cool, misty, and wet. includingbackwardsandsideways,whichgives
Some islands and island groups in the them an appearance of floating over the
Pacific have had intensive surveys of the surface. Flies of the closely related family
insects, such as in New Caledonia (Tillier, Neurochaetidae always orient themselves with
1988, and additional volumes in the Zoolo- the head downward, the others with the head
gia Neocaledonica series), Micronesia (e.g., upward; all keep their wings folded while
Gressitt, 1954, and additional volumes in the running.
Bishop Museum Insects of Micronesia series), Prior to this study there were only two
and the Hawaiian Islands (Zimmerman, 1948, periscelidids known from Fiji: Cyamops fiji
andmanysubsequentauthors).TheHawaiian Baptista and Mathis and Stenomicra distincti-
Islands are famous for spectacular insular pennis Collin, the latter of which was synon-
radiations of various plants and insects, ymized with S. fascipennis Malloch by Khoo
perhaps the best known being the 800 or so and Sabrosky (1989), but this is a misidenti-
species of endemic Drosophilidae (Hardy, fication and the status of S. distinctipennis is
1965). The study of Fiji arthropods, by reinstated in the present paper. Only one
comparison, has been relatively neglected Asteiidae was described from Fiji, Asteia
until recently (Evenhuis and Bickel, 2005). nigriceps Bezzi, to which four additional
As a result of funding from the Schlinger species are added here, along with a species
Foundation and the U.S. National Science of the small Indo-Pacific family Xenasteiidae.
Foundation,therehasbeenanintensiveeffort
tosurvey thearthropodsofFijiusing Malaise METHODS
trapping and specialized hand collecting (e.g.,
ants: Sarnat, 2006). Though Fiji is not as Some specimens were collected in Malaise
isolated as the Hawaiian Islands, its tropical traps that were set up on Viti Levu, Vanua
climate, high plant endemism, volcanic geolo- Levu, Taveuni, and Kadavu by Evert Schlin-
gywithmountainousterrain,andthefactthat ger, Michael Irwin, and Moala Tokota’a in
it is an extensive archipelago, would seem to 2002–2003, and maintained by M. Tokota’a
foster significant diversification of native and villagers in the field for a year or more.
arthropods, which indeed some studies indi- All specimens were preserved in ethanol.
cate (Bickel, 2006). These were received at the AMNH in vials
In the process of preparing a treatment on of drosophilids extracted from the Malaise
the Drosophilidae of Fiji, samples of asteiid residues. Additional specimens were collected
and periscelidid flies were included in the by me in July 2008, by aspirating them from
Malaise trap samples sent to me (these are the young, rolled leaves of various understory
often confused with drosophilids because of a ZingerberaleanplantsonVitiLevuandVanua
pectinate arista). Recent fieldwork by me in Levu.
2008 in addition uncovered an unexpected Specimens in alcohol were air dried by first
diversityofPeriscelididaeandalargeseriesof dehydrating them in 100% ethanol, then
Asteia, which inspired this study. All of the blotting them on tissue paper, separating the
specimens collected by hand were aspirated appendages with fine forceps, and then point
from the young, rolled and tubular leaves of mounting. Though there is some collapse of
bananas (Musa) and understory gingers the cuticle, particularly of the smaller, softer
(Zingerberaceae), a habitat that I have found Stenomicra, this preparation is more efficient
inSoutheastAsiaandtheneotropicstoharbor than critical point drying. The terminalia of
diverse periscelidids, especially Stenomicra. As pointed specimens were dissected by first
2009 GRIMALDI: ASTEIOINEA FLIESOFFIJI 3
softeningtheinsectforseveralhoursinaclosed Curiosimusca Rung, Mathis, and Papp com-
jar containing vinegar-soaked paper towels on prises just three species from Southeast Asia.
thebottom.Thedissectedterminaliawerethen The genera Nemo D.K. McAlpine (Austa-
macerated in warm 10% KOH, rinsed with lasian, African) and Ningulus D.K. Mc-
water and 70% EtOH, and transferred to Alpine (African) were formerly placed in the
glycerine, in which the genitalia were disartic- Aulacigastridae, but have now been placed in
ulated using minuten nadeln mounted on the family Neminidae with the genus Nemula
matchsticks. Genitalia were mounted in a 1% Freidberg(fromMadagascar).Neurochaetidae
agarose glycerine jelly between two coverslips, are very distinctive, flat flies comprising 23
toorientthestructuresandpreventthemfrom living species from Australia, Africa, Mada-
being crushed in a slide mount. The double gascar, and Southeast Asia (D.K. McAlpine,
coverslip mount allows anterior (internal gen- 1978, 1987a, 1987b, 1993; Woodley, 1982),
italia) and posterior (surstyli, cerci) views classified in the genera Neurocytta D.K.
without needing to remount. Wings were McAlpine,NeurochaetaD.K.McAlpine,Neuro-
removed from select specimens with fine texis D.K. McAlpine,and NothasteiaMalloch.
forceps, washed in 70% ethanol, and mounted Teratomyzidae has three genera (Teratomy-
on a glass slide in glycerine for photomicro- za Malloch, Teratoptera Malloch, and Neo-
graphy and measurements. For the perisceli- geomyza Se´guy), with approximately 12species
dids, the following metrics were important in from New Zealand, Australia, southern South
vein proportions: ratio of greatest wing width America,Africa,Nepal,andthePhillipines.
to wing length (W/L, as measured from cross Based on the distributions of Neuro-
veinhtothewingtip),andtheratiooflengthof chaetidae and Teratomyzidae, it would not be
cross vein br-m (L br-m) to the length of the surprising if they eventually are found in Fiji.
distance between this cross vein and dm-cu. Interestingly, there are no Aulacigastridae
Thorax length (ThL) was used as a reliable known from islands in the Pacific, which
metricforbodysize,whichwasmeasuredwith explains their absence in Fiji. Neurochaetidae
anocularmicrometerfromtheanteriorsurface have habits that are very similar to those of
of the mesoscutum to the posterior tip of the stenomicrinePeriscelididae,specificallyofrun-
mesoscutellum. ning over the broad surfaces of leaves that
Holotypes of new species will eventually be contain a wet substrate below. Aulacigas-
deposited in the Fiji National Insect Collec- tridae are associated with tree trunks, partic-
tion, Suva, but are temporarily deposited in ularly ones with wounds oozing sap or
the Bishop Museum in Honolulu (BPBM) decaying ones that are hanging over streams.
(most of the material collected in Malaise As currently defined (J.F. McAlpine, 1989),
traps) and American Museum of Natural monophyly of Asteioinea is only weakly
History, New York (AMNH) (material col- supported, and the position of Xenasteiidae
lected by D.A.G., and some from the Fiji in particular is controversial. That family, in
Malaise trap survey), kept in trust for Fiji for fact, may be within or closely related to the
when a new facility is completed. Milichioidea.
SYSTEMATICS FAMILY PERISCELIDIDAE
THE ASTEIOINEA
Thisisaglobalfamilyofonlyapproximate-
Besides the three families treated here, ly 80 described species, currently including
another three placed in this group are the 10 extant genera and one extinct genus
Aulacigastridae, Neurochaetidae, and Tera- (Procyamops, known from Baltic amber).
tomyzidae (J.F. McAlpine, 1989). Aula- Three of the extant genera are cosmopolitan
cigastridae contains two genera with approxi- (Cyamops [ca. 24 spp.], Periscelis [ca. 15 spp],
mately40describedspecies,virtuallyallinthe Stenomicra [27 described species]), and the
genus Aulacigaster Macquart (in which there remaining genera are monotypic or contain
arealsomorethan40undescribedNeotropical just a few species and are mostly restricted
species: W.N. Mathis, personal commun.). to the neotropics (Diopsosoma Malloch,
4 AMERICAN MUSEUMNOVITATES NO. 3671
MarbeniaMalloch,NeoscutopsMalloch,Paraf- Stenomicra ariela complex
scutops Mathis and Papp, Planinasus Cresson,
and Scutops Coquillett). Stenocyamops Papp This is a newly designated, taxonomically
was the most recently described genus, known informal group of seven small Stenomicra
until now from a species from Thailand. The species in Fiji that have a largely yellowish
genus Stenomicracontains more than 100new body color, some of them with light brown
species from the neotropics alone (C.W. infuscationoverthedorsumofthethoraxand
Sabrosky,unpubl.). abdomen. The group is not necessarily mono-
All of the Fiji periscelidids belong to three phyletic; such body coloration is common
closely related genera: Stenomicra, Cyamops, throughout the genus. In lieu of a worldwide
and the apparently intermediate Stenocya- revision of Stenomicra it would be difficult to
mops. A close relationship of the three genera identify even genitalic synapomorphies of
is based on the following synapomorphic these Fiji species. While subtle differences do
features: lower portion of face protrudent, occur in coloration and chaetotaxy, they are
withprojectingpairof‘‘pseudovibrissae,’’loss difficult to separate because of considerable
ofocellarsetae,lossofpostocellar/postvertical overlap in external variation. The most
setae, loss to great reduction of postpronotal reliable features for identification are in the
setae, vein CuA1 with an abrupt (vs. evanes- maleterminalia,especiallythestructureofthe
cent) distal end, tVII + sVIII fused in males, aedeagus and surstyli. As a result, females (9
and females with syntergosternite VII. Cy-
from Viti Levu, 1 from Vanua Levu, 3 from
amops is clearly a monophyletic group, but it
Taveuni, which seem to represent three
is possible that Stenocyamops may be just a
species) have only tentatively been assigned
basal grade to Stenomicra. A detailed phylo-
to species (map, fig. 22). Because of the great
genetic analysis of these genera is needed
external similarity a detailed description is
at the species level, but in lieu of a mono-
providedjustforStenomicraariela,n.sp.,and
graphic treatment of the world species of
S. tokotaai, n. sp.; diagnoses for the other
Stenomicra—of which there are hundreds—
species describe all relevant variation.
such an analysis would be premature at
present.
Stenomicra ariela, new species
Genus Stenomicra Coquillett Figures 5, 6
Stenomicra Coquillett, 1900: 262. Type species: S. DIAGNOSIS: Easily distinguished from all
angustataCoquillett,byoriginaldesignation. other Fijian Stenomicra by the completely
yellowbody(onlyocellartriangledarkbrown),
DIAGNOSIS: Small, slender, generally light-
and by the male genitalia with the broad,
colored flies with slender wings; possessing
the following combination of features that incurvedsurstylusandthebulbousaedeagus.
distinguish the genus from Cyamops and/or DESCRIPTION: ThL 5 0.54 mm. Differs
Stenocyamops: (maxillary) palps reduced to from S. distincta and S. tokotaai by the
minute papillae; inner vertical setae present, following features: Arista with four dorsal,
proclinate (absent in Cyamops); acrostichal three ventral branches. Basal flagellomere
setulae in one incomplete row, or lost (two withpointedapex,apicaltuftofsetulaenearly
or more rows in Cyamops); anal lobe of equal in length to flagellomere. Posterior
wing, alula, vein A +CuA completely or fronto-orbital seta very far anteriad on frons,
1 2
virtually lost (all these features present in past dorsal level of antennae; both FO setae
Cyamops; some features present in Steno- appear reclinate (possibly preservational, see
cyamops); anepisternum without silvery mi- below). Pseudovibrissae fairly long, length
crotomentum or row of setae on posterior nearly equal to that of posterior FO seta.
margin; paired structures of male genitalia Row of four long, fine acrostichals; dorsocen-
symmetrical (asymmetrical in Cyamops), trals in graded series, anteriormost shortest,
hypandrium asymmetrical (symmetrical in posteriormost dclongest(approximately1.43
Stenocyamops). length of preceding dc). Notopleuron with
2009 GRIMALDI: ASTEIOINEA FLIESOFFIJI 5
four setae; dorsal two about 0.53 length of able. Arista with 5 dorsal, 2 ventral branches.
ventral ones. One katepisternal seta present, Besides coloration, also distinguished from
erect. Halter yellow, not white. Forefemur similar species by male terminalia: Surstylus
with two lateral rows long, erect, fine setae, relatively short, apical third curved upward,
plus two longer setae that project posteriad. apex acute; with dense setulae over much of
Midfemur with ventral row of long, stiff, fine surstylus. Aedeagus with straight trunk, apex
setae. Wing hyaline, rather slender and long brushyandwithnarrowtip.
(W/L50.38);tipslightlyandbluntlypointed; TYPES: Holotype, male (dissected, no. 57),
dorsal edge of costal vein with row of ca. 5 FIJI: Taveuni: Cakaudrove Prov., 5.6 km
stout setae near junction of R . Cross veins r- SE Tavuki Vlg., Devo Peak, 1187 m, 30.VI–
1
mandm-cuclosetogether(Lr-m/Lbetween 14.VIII. 2004, Malaise 1, Schlinger, M.
r-m and m-cu 5 0.23). Alula, anal lobe, vein Tokota’a, 16.843uS, 179.966uW, FBA 50904.
A +CuA essentially absent. Male genitalia: ThL51.22 mm;Lr-m/Lbetweenr-mandm-
1 2
Epandrium shallow, lateroventral lobes with cu 5 0.53. In BPBM. Specimen is missing the
ca.7large,stiffsetaepointeddownward;cerci scutellar and most dorsocentral setae. In addi-
largely membranous, attached to epandrium, tion, there are 3 females that match the male
eachlobewithca.10 minute,spinulelikesetae, holotype in color and chaetotaxy, also from
oneverylongsetaonventralmargin.Surstylus Cakaudrove Prov.: 3.2 km NW Lavena Vlg.,
broad, curved inward and forming ventral Mt. Koronibuabua, 217 m, 24.II–11.III.04,
shelf.Hypandriumasimple,flat,bilobedplate, Malaise 3, Schlinger and Tokota’a, 16.855uS,
nearly symmetrical; aedeagus bulbous, ventral 179.89uW,FBA127415,153258,168376.
portion covered with microtrichia, lateral, ETYMOLOGY: From the Latin, brunneus,
membranous lobe of aedeagus with microtri- for brown, in reference to the color of the
chia and sclerotized strip. Base of aedeagus scutum and tergites.
broad, trough shaped, with two lightly sclero-
tized appendages (paraphyses?), left one dis-
Stenomicra castanea, new species
placedlaterally,rightonepointedventrad.
TYPE: Holotype, male: FIJI: Viti Levu, Figures 4, 8
VII.9.08, Nakobalevu Rd., 394 m., 18u039310S,
178u249550E, in rolled leaves, D. Grimaldi DIAGNOSIS: Frons white; scutum and scu-
(no. 39: terminalia dissected). In AMNH. tellum evenly light brown, without central
Known only from a unique male. The thorax longitudinal light stripe; light brown infusca-
and appendages are in good condition, but the tion on dorsal portion of pleura. Wing
eyes, face, and portion of the frons are partly hyaline. Abdominal tergites I–IV light brown.
collapsed. Besides the scutum coloration, best distin-
ETYMOLOGY: From Greek and Hebrew, guished on basis of male terminalia: surstyli
meaning an airy sprite, in reference to the straight, not curved, with blunt apex, without
light-colored body of this species and the microtrichia and few setulae; trunk of aedea-
floating movements of flies in this genus. gus straight, apex brushy; syntergosternite
VII ventrally with median lobe on posterior
margin.
Stenomicra brunnea, new species
TYPE: Holotype, male (dissected, no. 59),
Figures 4, 7 Viti Levu: Naitasiri Prov., 4.8 km N Veisari
Stlmt, log rd to Waivudawa, 300 m, 12.XII–
DIAGNOSIS: Frons whitish, pollinose; scu- 3.I.03.Malaise1.Schlinger,Tokota’a,18.075uS,
tum and scutellum dark brown, scutum with 178.362uE,FBA178234.ThL51.20 mm;Lr-
yellow longitudinal stripe that is narrower m/Lbetweenr-mandm-cu50.50.Typeisin
than distance between dorsocentrals; pleura goodcondition,withmostofheadandthoracic
without brown infuscation. All abdominal setaeintact.InBPBM.
tergites brown; wings hyaline. Scutellum ETYMOLOGY: From the Latin castaneus,
narrow. Anterior fronto-orbital and post- for the color of chestnuts, in reference to the
vertical setae minute and light, barely detect- color of the scutum and scutellum.
6 AMERICAN MUSEUMNOVITATES NO. 3671
Stenomicra pallida, new species females, all Taveuni: Cakaudrove, 5.6 km SE
Tavuki Vlg., Mt. Devo, 892–1187 m, 30.VI–
Figure 9
14.VIII, 2004, Malaise 1, 3, 4; Schlinger and
Tokota’a, 16.843uS, 179.966uW, FBA 149851,
DIAGNOSIS: Anentirelywhitish-yellowspe- 149860, 150903, 151943, 154918, 154919. In
cies, withbarelyanyinfuscation onscutumor
BPBM and AMNH. Genitalia of the holotype
tergites; frons creamy white; wing hyaline.
arewelldisplayed,confirmingthatthisspecimen
Anterior fronto-orbital seta very fine, minute;
andthedissectedparatypearethesamespecies.
arista with 5 dorsal and 2 ventral branches; 5
ETYMOLOGY: From the French, sylphe,
dorsocentrals in each row, row of 3 acrosti-
meaningafairylikespiritoftheair,inreference
chals. Male terminalia distinctive: surstyli
tothepalebodycolorationofthislittlefly.
slightly S-shaped, with only three apical/sub-
apical setulae (no microtrichia); aedeagus
spiculed, with central groove, without trunk, Stenomicra tokotaai, new species
tipnarrow;stripofaedeagalscleritenotfound.
Figures 1, 2, 3, 5 10
TYPE: Holotype, male (dissected, no. 68),
VitiLevu:NaitasiriProv.,4 kmWSWColo-i-
SuvaVillage,Mt.Nakobalevu,372 m,17.III– DIAGNOSIS: Scutum and scutellum light
brown, with light bluish pruinescence; pleura
9.IV.03, Malaise 3, Schlinger, Tokota’a,
mostly light yellow with light brown stripe on
18.055uS, 178.424uE, FBA 144101. ThL 5
dorsalmarginofanepisternumandanatergite;
1.22 mm; Wing W/L 5 0.30; L r-m / L
wing completely hyaline, long, slender, with
between r-m and m-cu 5 0.40. Specimen is
cross veins very close together; most abdom-
in good condition; deposited in BPBM.
inaltergiteslightbrown,tV–VIinfemalesand
ETYMOLOGY: From the Latin, pallidus, for
tV–VIIIinmaleslightyellow.Surstylus(male)
pale,inreferencetotheentirelywhitish-yellow
simple, slender (length 63 greatest width),
body coloration.
slightly curved, tip acute.
DESCRIPTION: ThL 5 0.60 mm. Body col-
Stenomicra sylpha, new species
oration: Frons entirely dull, whitish beige,
smallareabetweenocellidarkbrown;faceand
Figures 4, 10
facial protuberance light brown; oral margin,
cheek, postgena, proboscis, antennal pedicel
DIAGNOSIS: Body almost entirely light yel-
and basal flagellomere light yellow. Eyes light
low, abdominal tergites I–IV slightly darker;
red to pink. Mesoscutum and scutellum with
frons white, pollinose. Wing hyaline. Pseu-
ground color a very light brown, having light
dovibrissae stout, laterally flattened, slightly
scalelike. Arista with 5 dorsal and 2 ventral bluish pruinescence; lateral (notopleural)
branches. Best distinguished from similar edges diffuse, tip of scutellum light. Most of
yellow and light brown species by male pleuron light yellow, but with thin, light
terminalia: Each cercus with long, thick seta brown stripe on dorsal margin of anepister-
at apex of ventral margin; cercus anteriorly numandanatergite.Legsentirelylightyellow;
(underneath) with row of 7–8 short setae. wing entirely hyaline (no areas of infuscation
Surstylus long, curved and arched, apically or white); haltere entirely white. Abdominal
pointed, with very sparse setulae and micro- tergitesV–VIinfemalesandtVI–VIIIinmales
trichia. Aedeagus short (without visible light yellow; all others light brown; all
trunk), membranous, spiculed. sternites light brown. Setae on head and
TYPES: Holotype, male (not dissected), thorax dark copper; leg setae light.
FIJI: Taveuni, Cakaudrove Prov., 5.3 km SE Head: Slightly flattened, greatest length
Tavuki Vlg. [Village], Mt. Devo, 1064 m, oblique in profile. Ocellar setae lost; inner
3–20.XII.2002, Malaise 3. Coll. Schlinger, verticalsetaepresent,proclinate;outervertical
Tokota’a. 16.841uS, 179.968uW, FBA 153797. setae reclinate. Two pairs fronto-orbital (FO)
ThL 5 1.55 mm; Wing W/L 50.36; L r-m / L setae present, both pairs significantly anterior
between r-m and m-cu 5 0.33. In BPBM. to level of ocellar triangle; anterior FO
Paratypes: 1 male (no. 55 [150902] dissected), 5 proclinate fine, minute, ca. 0.33 (male) to
2009 GRIMALDI: ASTEIOINEA FLIESOFFIJI 7
Fig. 1. Heads of select Fiji Periscelididae, oblique frontal views (not to same scale). A. Stenomicra
tokotaai, n.sp. B. Stenomicra distinctipennis Collin. C.Cyamopsfiji Baptista andMathis.
0.43 (females) length of posterior FO; poste- longer dorsally); no differentiation of facets;
rior FO reclinate. Eyes emarginate on face distance between eye margins on face slightly
around antennae and on postgenal margin; greater than distance between pseudovibris-
with dense, short interfacetal setulae (slightly sae. Antenna: Scape with fine dorsal seta
8 AMERICAN MUSEUMNOVITATES NO. 3671
Fig. 2. Thoraces of select Fiji Periscelididae showing coloration, left lateral views (to same scale). A.
Stenomicratokotaai,n.sp.B.Stenomicradistincta,n.sp.C.Stenocyamopsrobustus,n.sp.D.Cyamopsfiji
Baptista andMathis.
2009 GRIMALDI: ASTEIOINEA FLIESOFFIJI 9
Fig.3. AbdominalcolorationofselectFijiPeriscelididae,dorsalviews(tosamescale).A.Stenocyamops
vittatus,n.sp.B.Stenocyamopsrobustus.C.Stenocyamopsluteus,n.sp.D,E:Stenomicratokotaii,male(D)
andapexof female abdomen(E).F,G: Stenomicra distincta, male (F)and apexof female abdomen(G).
10 AMERICAN MUSEUMNOVITATES NO. 3671
Fig.4. A–E:TerminalabdominalsegmentsandsyntergosternitesofmalePeriscelididae(ventralviews,
nottosamescale).A.Stenocyamopsrobustus.B.Stenocyamopsluteus/pseudoluteus.C.Stenomicradistincta.
D.Stenomicradistinctipennis.E.Stenomicracastanea.F–H:Thoraces(dorsalview)ofselectFijiStenomicra
(tothe same scale). F.S.brunnea. G.S.xoutha.H. S.sylpha.