The Asteioinea of Fiji (Insecta, Diptera, Periscelididae, Asteiidae, Xenasteiidae)

2009·4.2 MB·English

by  Grimaldi, David A

A tamerican museum
Novitates
PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY
CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024
Number 3671, 59 pp., 32 figures, 2 tables November 30, 2009
The Asteioinea of Fiji (Insecta: Diptera:
Periscelididae, Asteiidae, Xenasteiidae)
DAVID A. GRIMALDI1
ABSTRACT
Three of the six families in the acalyptrate fly group Asteioinea are reported from Fiji,
comprising five genera and 23 species, with all except three of the species being new. The new
species in their respective families are the following: Stenomicra ariela. Si. brunnea. Si. castanea. Si.
distincta. Si. pallida. Si. sylpha. Si. tokotaai, and Si. xoutha; Cyamops femobrunneus, C.
femoctenidius; Stenocyamops luteus, Sy. pseudoluteus, Sy. robustus, and Sy. vittatus
(Periscelididae); Asteia pleurovitta, A. pleurovittata, A. rotundiscuta, A. vanuaensis, A. vitiensis
(Asteiidae); and Xenasteia fijiana (Xenasteiidae). This is the first report of Stenocyamops outside of
Southeast Asia. Stenomicra distinctipennis (Collin) is resurrected from synonymy with S.
fascipennis Malloch, based on study of male genitalia belonging to a complex of fascipennis-like
species from throughout the Pacific. Asteia nigriceps Bezzi is redescribed. Keys to genera and
species are provided. Some species were collected in the rolled leaves of bananas and wild gingers
(Zingerberales), which appears to be an important habitat for periscelidids in tropical forests.
INTRODUCTION the Fiji archipelago. Fiji consists of approxi¬
mately 322 islands and over 500 islets, lying
This is a taxonomic treatment of the species between 15°-20° S and 175° E-1750 W. Four
belonging to three families of small, obscure Fiji islands have sufficient size and relief
schizophoran flies from the South Pacific to support lush tropical forests, which are
nation of the Republic of the Fiji Islands. Viti Levu (10,388 km2), Vanua Levu (5587
The three families—Periscelididae, Asteiidae, km2), Taveuni (435 km2), and Kadavu
and Xenasteiidae—belong to the superfamily (411 km2). All of the insect sampling for this
Asteioinea (McAlpine, 1989), of which the and most other insect studies has focused on
taxonomy here is a small contribution towards these four islands, since smaller islands are
understanding the arthropod biodiversity of low, drier, and support less forest. Climate is
'Division of Invertebrate Zoology, American Museum of Natural History ([email protected]).
Copyright © American Museum of Natural History 2009 ISSN 0003-0082
2 AMERICAN MUSEUM NOVITATES NO. 3671
tropical, but because of the rugged, moun¬ has been described elsewhere for Stenomicra
tainous topography of the four largest islands and the closely related family Neurochaetidae
there are dramatic effects to local climate. (D.K. McAlpine, 1978, 1987a, 1987b, 1993),
Northern halves of these islands, for example, the Fiji periscelidids—and even the Asteia
have a pronounced dry season and the cohabiting with them—have a remarkable
southern halves support moist, evergreen way of moving. These flies run over the surface
forests; the higher peaks and ridges of the of a leaf in all directions with uniform effort,
central mountains are cool, misty, and wet. including backwards and sideways, which gives
Some islands and island groups in the them an appearance of floating over the
Pacific have had intensive surveys of the surface. Flies of the closely related family
insects, such as in New Caledonia (Tillier, Neurochaetidae always orient themselves with
1988, and additional volumes in the Zoolo- the head downward, the others with the head
gia Neocaledonica series), Micronesia (e.g., upward; all keep their wings folded while
Gressitt, 1954, and additional volumes in the running.
Bishop Museum Insects of Micronesia series), Prior to this study there were only two
and the Hawaiian Islands (Zimmerman, 1948, periscelidids known from Fiji: Cyamops fiji
and many subsequent authors). The Hawaiian Baptista and Mathis and Stenomicra distincti-
Islands are famous for spectacular insular pennis Collin, the latter of which was synon-
radiations of various plants and insects, ymized with S. fascipennis Malloch by Khoo
perhaps the best known being the 800 or so and Sabrosky (1989), but this is a misidenti-
species of endemic Drosophilidae (Hardy, fication and the status of S. distinctipennis is
1965). The study of Fiji arthropods, by reinstated in the present paper. Only one
comparison, has been relatively neglected Asteiidae was described from Fiji, Asteia
until recently (Evenhuis and Bickel, 2005). nigriceps Bezzi, to which four additional
As a result of funding from the Schlinger species are added here, along with a species
Foundation and the U.S. National Science of the small Indo-Pacific family Xenasteiidae.
Foundation, there has been an intensive effort
to survey the arthropods of Fiji using Malaise
METHODS
trapping and specialized hand collecting (e.g.,
ants: Sarnat, 2006). Though Fiji is not as Some specimens were collected in Malaise
isolated as the Hawaiian Islands, its tropical traps that were set up on Viti Levu, Vanua
climate, high plant endemism, volcanic geolo¬ Levu, Taveuni, and Kadavu by Evert Schlin¬
gy with mountainous terrain, and the fact that ger, Michael Irwin, and Moala Tokota’a in
it is an extensive archipelago, would seem to 2002-2003, and maintained by M. Tokota’a
foster significant diversification of native and villagers in the field for a year or more.
arthropods, which indeed some studies indi¬ All specimens were preserved in ethanol.
cate (Bickel, 2006). These were received at the AMNH in vials
In the process of preparing a treatment on of drosophilids extracted from the Malaise
the Drosophilidae of Fiji, samples of asteiid residues. Additional specimens were collected
and periscelidid flies were included in the by me in July 2008, by aspirating them from
Malaise trap samples sent to me (these are the young, rolled leaves of various understory
often confused with drosophilids because of a Zingerberalean plants on Viti Levu and Vanua
pectinate arista). Recent fieldwork by me in Levu.
2008 in addition uncovered an unexpected Specimens in alcohol were air dried by first
diversity of Periscelididae and a large series of dehydrating them in 100% ethanol, then
Asteia, which inspired this study. All of the blotting them on tissue paper, separating the
specimens collected by hand were aspirated appendages with fine forceps, and then point
from the young, rolled and tubular leaves of mounting. Though there is some collapse of
bananas {Musa) and understory gingers the cuticle, particularly of the smaller, softer
(Zingerberaceae), a habitat that I have found Stenomicra, this preparation is more efficient
in Southeast Asia and the neotropics to harbor than critical point drying. The terminalia of
diverse periscelidids, especially Stenomicra. As pointed specimens were dissected by first
2009 GRIMALDI: ASTEIOINEA FLIES OF FIJI 3
softening the insect for several hours in a closed Curiosimusca Rung, Mathis, and Papp com¬
jar containing vinegar-soaked paper towels on prises just three species from Southeast Asia.
the bottom. The dissected terminalia were then The genera Nemo D.K. McAlpine (Austa-
macerated in warm 10% KOH, rinsed with lasian, African) and Ningulus D.K. Mc¬
water and 70% EtOH, and transferred to Alpine (African) were formerly placed in the
glycerine, in which the genitalia were disartic¬ Aulacigastridae, but have now been placed in
ulated using minuten nadeln mounted on the family Neminidae with the genus Nemula
matchsticks. Genitalia were mounted in a 1% Freidberg (from Madagascar). Neurochaetidae
agarose glycerine jelly between two coverslips, are very distinctive, flat flies comprising 23
to orient the structures and prevent them from living species from Australia, Africa, Mada¬
being crushed in a slide mount. The double gascar, and Southeast Asia (D.K. McAlpine,
coverslip mount allows anterior (internal gen¬ 1978, 1987a, 1987b, 1993; Woodley, 1982),
italia) and posterior (surstyli, cerci) views classified in the genera Neurocytta D.K.
without needing to remount. Wings were McAlpine, Neurochaeta D.K. McAlpine, Neuro-
removed from select specimens with fine texis D.K. McAlpine, and Nothasteia Malloch.
forceps, washed in 70% ethanol, and mounted Teratomyzidae has three genera (Teratomy-
on a glass slide in glycerine for photomicro¬ za Malloch, Teratoptera Malloch, and Neo-
graphy and measurements. For the perisceli- geomyza Seguy), with approximately 12 species
dids, the following metrics were important in from New Zealand, Australia, southern South
vein proportions: ratio of greatest wing width America, Africa, Nepal, and the Phillipines.
to wing length (W/L, as measured from cross Based on the distributions of Neuro¬
vein h to the wing tip), and the ratio of length of chaetidae and Teratomyzidae, it would not be
cross vein br-m (L br-m) to the length of the surprising if they eventually are found in Fiji.
distance between this cross vein and dm-cu. Interestingly, there are no Aulacigastridae
Thorax length (ThL) was used as a reliable known from islands in the Pacific, which
metric for body size, which was measured with explains their absence in Fiji. Neurochaetidae
an ocular micrometer from the anterior surface have habits that are very similar to those of
of the mesoscutum to the posterior tip of the stenomicrine Periscelididae, specifically of run¬
mesoscutellum. ning over the broad surfaces of leaves that
Holotypes of new species will eventually be contain a wet substrate below. Aulacigas¬
deposited in the Fiji National Insect Collec¬ tridae are associated with tree trunks, partic¬
tion, Suva, but are temporarily deposited in ularly ones with wounds oozing sap or
the Bishop Museum in Honolulu (BPBM) decaying ones that are hanging over streams.
(most of the material collected in Malaise As currently defined (J.F. McAlpine, 1989),
traps) and American Museum of Natural monophyly of Asteioinea is only weakly
History, New York (AMNH) (material col¬ supported, and the position of Xenasteiidae
lected by D.A.G., and some from the Fiji in particular is controversial. That family, in
Malaise trap survey), kept in trust for Fiji for fact, may be within or closely related to the
when a new facility is completed. Milichioidea.
SYSTEMATICS FAMILY PERISCELIDIDAE
THE ASTEIOINEA
This is a global family of only approximate¬
Besides the three families treated here, ly 80 described species, currently including
another three placed in this group are the 10 extant genera and one extinct genus
Aulacigastridae, Neurochaetidae, and Tera- (Procyamops, known from Baltic amber).
tomyzidae (J.F. McAlpine, 1989). Aula¬ Three of the extant genera are cosmopolitan
cigastridae contains two genera with approxi¬ (Cyamops [ca. 24 spp.], Periscelis [ca. 15 spp],
mately 40 described species, virtually all in the Stenomicra [27 described species]), and the
genus Aulacigaster Macquart (in which there remaining genera are monotypic or contain
are also more than 40 undescribed Neotropical just a few species and are mostly restricted
species: W.N. Mathis, personal commun.). to the neotropics (Diopsosoma Malloch,
4 AMERICAN MUSEUM NOVITATES NO. 3671
Marbenia Malloch, Neoscutops Malloch, Paraf- Stenomicra ariela complex
scutops Mathis and Papp, Planinasus Cresson,
and Scutops Coquillett). Stenocyamops Papp This is a newly designated, taxonomically
was the most recently described genus, known informal group of seven small Stenomicra
until now from a species from Thailand. The species in Fiji that have a largely yellowish
genus Stenomicra contains more than 100 new body color, some of them with light brown
species from the neotropics alone (C.W. infuscation over the dorsum of the thorax and
Sabrosky, unpubl.). abdomen. The group is not necessarily mono¬
All of the Fiji periscelidids belong to three phyletic; such body coloration is common
closely related genera: Stenomicra, Cyamops, throughout the genus. In lieu of a worldwide
and the apparently intermediate Stenocya¬ revision of Stenomicra it would be difficult to
mops. A close relationship of the three genera identify even genitalic synapomorphies of
is based on the following synapomorphic these Fiji species. While subtle differences do
features: lower portion of face protrudent, occur in coloration and chaetotaxy, they are
with projecting pair of “pseudovibrissae,” loss difficult to separate because of considerable
of ocellar setae, loss of postocellar/postvertical overlap in external variation. The most
setae, loss to great reduction of postpronotal reliable features for identification are in the
setae, vein CuAi with an abrupt (vs. evanes¬ male terminalia, especially the structure of the
cent) distal end, tVII + sVIII fused in males,
aedeagus and surstyli. As a result, females (9
and females with syntergosternite VII. Cy¬ from Viti Levu, 1 from Vanua Levu, 3 from
amops is clearly a monophyletic group, but it Taveuni, which seem to represent three
is possible that Stenocyamops may be just a species) have only tentatively been assigned
basal grade to Stenomicra. A detailed phylo¬ to species (map, fig. 22). Because of the great
genetic analysis of these genera is needed
external similarity a detailed description is
at the species level, but in lieu of a mono¬
provided just for Stenomicra ariela, n. sp., and
graphic treatment of the world species of
S. tokotaai, n. sp.; diagnoses for the other
Stenomicra—of which there are hundreds—
species describe all relevant variation.
such an analysis would be premature at
present.
Stenomicra ariela, new species
Genus Stenomicra Coquillett Figures 5, 6
Stenomicra Coquillett, 1900: 262. Type species: S. Diagnosis: Easily distinguished from all
angustata Coquillett, by original designation. other Fijian Stenomicra by the completely
yellow body (only ocellar triangle dark brown),
Diagnosis: Small, slender, generally light-
colored flies with slender wings; possessing and by the male genitalia with the broad,
the following combination of features that incurved surstylus and the bulbous aedeagus.
distinguish the genus from Cyamops and/or Description: ThL = 0.54 mm. Differs
Stenocyamops: (maxillary) palps reduced to from S. distincta and S. tokotaai by the
minute papillae; inner vertical setae present, following features: Arista with four dorsal,
proclinate (absent in Cyamops)’, acrostichal three ventral branches. Basal flagellomere
setulae in one incomplete row, or lost (two with pointed apex, apical tuft of setulae nearly
or more rows in Cyamops)’, anal lobe of equal in length to flagellomere. Posterior
wing, alula, vein A!+CuA2 completely or fronto-orbital seta very far anteriad on frons,
virtually lost (all these features present in past dorsal level of antennae; both FO setae
Cyamops’, some features present in Steno¬ appear reclinate (possibly preservational, see
cyamops)’, anepisternum without silvery mi- below). Pseudovibrissae fairly long, length
crotomentum or row of setae on posterior nearly equal to that of posterior FO seta.
margin; paired structures of male genitalia Row of four long, fine acrostichals; dorsocen-
symmetrical (asymmetrical in Cyamops), trals in graded series, anteriormost shortest,
hypandrium asymmetrical (symmetrical in posteriormost dc longest (approximately 1.4X
Stenocyamops). length of preceding dc). Notopleuron with
2009 GRIMALDI: ASTEIOINEA FLIES OF FIJI 5
four setae; dorsal two about 0.5 X length of able. Arista with 5 dorsal, 2 ventral branches.
ventral ones. One katepisternal seta present, Besides coloration, also distinguished from
erect. Halter yellow, not white. Forefemur similar species by male terminalia: Surstylus
with two lateral rows long, erect, fine setae, relatively short, apical third curved upward,
plus two longer setae that project posteriad. apex acute; with dense setulae over much of
Midfemur with ventral row of long, stiff, fine surstylus. Aedeagus with straight trunk, apex
setae. Wing hyaline, rather slender and long brushy and with narrow tip.
(W/L = 0.38); tip slightly and bluntly pointed; Types: Holotype, male (dissected, no. 57),
dorsal edge of costal vein with row of ca. 5 FIJI: Taveuni: Cakaudrove Prov., 5.6 km
stout setae near junction of Ri. Cross veins r- SE Tavuki Vlg., Devo Peak, 1187 m, 30.VI-
m and m-cu close together (L r-m / L between 14. VIII. 2004, Malaise 1, Schlinger, M.
r-m and m-cu = 0.23). Alula, anal lobe, vein Tokota’a, 16.843°S, 179.966°W, FBA 50904.
A1+CuA2 essentially absent. Male genitalia: ThL = 1.22 mm; L r-m / L between r-m and m-
Epandrium shallow, lateroventral lobes with cu = 0.53. In BPBM. Specimen is missing the
ca. 7 large, stiff setae pointed downward; cerci scutellar and most dorsocentral setae. In addi¬
largely membranous, attached to epandrium, tion, there are 3 females that match the male
each lobe with ca. 10 minute, spinulelike setae, holotype in color and chaetotaxy, also from
one very long seta on ventral margin. Surstylus Cakaudrove Prov.: 3.2 km NW La vena Vlg.,
broad, curved inward and forming ventral Mt. Koronibuabua, 217 m, 24.11 11.111.04,
shelf. Hypandrium a simple, flat, bilobed plate, Malaise 3, Schlinger and Tokota’a, 16.855°S,
nearly symmetrical; aedeagus bulbous, ventral 179.89°W, FBA127415, 153258, 168376.
portion covered with microtrichia, lateral, Etymology: From the Latin, brunneus,
membranous lobe of aedeagus with microtri¬ for brown, in reference to the color of the
chia and sclerotized strip. Base of aedeagus scutum and tergites.
broad, trough shaped, with two lightly sclero¬
tized appendages (paraphyses?), left one dis¬
Stenomicra castanea, new species
placed laterally, right one pointed ventrad.
Type: Holotype, male: FIJI: Viti Levu, Figures 4, 8
VII.9.08, Nakobalevu Rd., 394 m., 18°03'31"S,
178°24'55"E, in rolled leaves, D. Grimaldi Diagnosis: Frons white; scutum and scu¬
(no. 39: terminalia dissected). In AMNH. tellum evenly light brown, without central
Known only from a unique male. The thorax longitudinal light stripe; light brown infusca¬
and appendages are in good condition, but the tion on dorsal portion of pleura. Wing
eyes, face, and portion of the frons are partly hyaline. Abdominal tergites I IV light brown.
collapsed. Besides the scutum coloration, best distin¬
Etymology: From Greek and Hebrew, guished on basis of male terminalia: surstyli
meaning an airy sprite, in reference to the straight, not curved, with blunt apex, without
light-colored body of this species and the microtrichia and few setulae; trunk of aedea¬
floating movements of flies in this genus. gus straight, apex brushy; syntergosternite
VII ventrally with median lobe on posterior
margin.
Stenomicra brunnea, new species
Type: Holotype, male (dissected, no. 59),
Figures 4, 7 Viti Levu: Naitasiri Prov., 4.8 km N Veisari
Stlmt, log rd to Waivudawa, 300 m, 12.XII-
Diagnosis: Frons whitish, pollinose; scu¬ 3.1.03. Malaise 1. Schlinger, Tokota’a, 18.075°S,
tum and scutellum dark brown, scutum with 178.362°E, FBA 178234. ThL = 1.20 mm; L r-
yellow longitudinal stripe that is narrower m / L between r-m and m-cu = 0.50. Type is in
than distance between dorsocentrals; pleura good condition, with most of head and thoracic
without brown infuscation. All abdominal setae intact. In BPBM.
tergites brown; wings hyaline. Scutellum Etymology: From the Latin castaneus,
narrow. Anterior fronto-orbital and post¬ for the color of chestnuts, in reference to the
vertical setae minute and light, barely detect¬ color of the scutum and scutellum.
6 AMERICAN MUSEUM NOVITATES NO. 3671
Stenomicra pallida, new species females, all Taveuni: Cakaudrove, 5.6 km SE
Tavuki Vlg., Mt. Devo, 892-1187 m, 30.VI-
Figure 9
14.VIII, 2004, Malaise 1, 3, 4; Schlinger and
Tokota’a, 16.843°S, 179.966°W, FBA 149851,
Diagnosis: An entirely whitish-yellow spe¬ 149860, 150903, 151943, 154918, 154919. In
cies, with barely any infuscation on scutum or
BPBM and AMNH. Genitalia of the holotype
tergites; frons creamy white; wing hyaline.
are well displayed, confirming that this specimen
Anterior fronto-orbital seta very fine, minute;
and the dissected paratype are the same species.
arista with 5 dorsal and 2 ventral branches; 5
Etymology: From the French, sylphe,
dorsocentrals in each row, row of 3 acrosti-
meaning a fairylike spirit of the air, in reference
chals. Male terminalia distinctive: surstyli
to the pale body coloration of this little fly.
slightly S-shaped, with only three apical/sub-
apical setulae (no microtrichia); aedeagus
spiculed, with central groove, without trunk, Stenomicra tokotaai, new species
tip narrow; strip of aedeagal sclerite not found.
Figures 1, 2, 3, 5 10
Type: Holotype, male (dissected, no. 68),
Viti Levu: Naitasiri Prov., 4 km WSW Colo-i-
Diagnosis: Scutum and scutellum light
Suva Village, Mt. Nakobalevu, 372 m, 17.III-
brown, with light bluish pruinescence; pleura
9.IV.03, Malaise 3, Schlinger, Tokota’a,
mostly light yellow with light brown stripe on
18.055°S, 178.424°E, FBA 144101. ThL =
dorsal margin of anepisternum and anatergite;
1.22 mm; Wing W/L = 0.30; L r-m / L
wing completely hyaline, long, slender, with
between r-m and m-cu = 0.40. Specimen is
cross veins very close together; most abdom¬
in good condition; deposited in BPBM.
inal tergites light brown, tV-VI in females and
Etymology: From the Latin, pallidus, for
tV VIII in males light yellow. Surstylus (male)
pale, in reference to the entirely whitish-yellow
simple, slender (length 6X greatest width),
body coloration.
slightly curved, tip acute.
Description: ThL = 0.60 mm. Body col¬
Stenomicra sylpha, new species oration: Frons entirely dull, whitish beige,
small area between ocelli dark brown; face and
Figures 4, 10
facial protuberance light brown; oral margin,
cheek, postgena, proboscis, antennal pedicel
Diagnosis: Body almost entirely light yel¬
and basal flagellomere light yellow. Eyes light
low, abdominal tergites I-IV slightly darker;
red to pink. Mesoscutum and scutellum with
frons white, pollinose. Wing hyaline. Pseu-
ground color a very light brown, having light
dovibrissae stout, laterally flattened, slightly
bluish pruinescence; lateral (notopleural)
scalelike. Arista with 5 dorsal and 2 ventral
branches. Best distinguished from similar edges diffuse, tip of scutellum light. Most of
yellow and light brown species by male pleuron light yellow, but with thin, light
terminalia: Each cercus with long, thick seta brown stripe on dorsal margin of anepister¬
at apex of ventral margin; cercus anteriorly num and anatergite. Legs entirely light yellow;
(underneath) with row of 7-8 short setae. wing entirely hyaline (no areas of infuscation
Surstylus long, curved and arched, apically or white); haltere entirely white. Abdominal
pointed, with very sparse setulae and micro¬ tergites V-VI in females and tVI-VIII in males
trichia. Aedeagus short (without visible light yellow; all others light brown; all
trunk), membranous, spiculed. sternites light brown. Setae on head and
Types: Holotype, male (not dissected), thorax dark copper; leg setae light.
FIJI: Taveuni, Cakaudrove Prov., 5.3 km SE Head: Slightly flattened, greatest length
Tavuki Vlg. [Village], Mt. Devo, 1064 m, oblique in profile. Ocellar setae lost; inner
3-20.XII.2002, Malaise 3. Coll. Schlinger, vertical setae present, proclinate; outer vertical
Tokota’a. 16.84US, 179.968°W, FBA 153797. setae reclinate. Two pairs fronto-orbital (FO)
ThL = 1.55 mm; Wing W/L =0.36; L r-m / L setae present, both pairs significantly anterior
between r-m and m-cu = 0.33. In BPBM. to level of ocellar triangle; anterior FO
Paratypes: 1 male (no. 55 [150902] dissected), 5 proclinate fine, minute, ca. 0.3 X (male) to
2009 GRIMALDI: ASTEIOINEA FLIES OF FIJI 7
Cyamops fiji
Fig. 1. Heads of select Fiji Periscelididae, oblique frontal views (not to same scale). A. Stenomicra
tokotaai, n. sp. B. Stenomicra distinctipennis Collin. C. Cyamops fiji Baptista and Mathis.
0.4X (females) length of posterior FO; poste¬ longer dorsally); no differentiation of facets;
rior FO reclinate. Eyes emarginate on face distance between eye margins on face slightly
around antennae and on postgenal margin; greater than distance between pseudovibris-
with dense, short interfacetal setulae (slightly sae. Antenna: Scape with fine dorsal seta
AMERICAN MUSEUM NOVITATES NO. 3671
Fig. 2. Thoraces of select Fiji Periscelididae showing coloration, left lateral views (to same scale). A.
Stenomicra tokotaai, n. sp. B. Stenomicra distincta, n. sp. C. Stenocyamops robustus, n. sp. D. Cyamops fiji
Baptista and Mathis.
2009 GRIMALDI: ASTEIOINEA FLIES OF FIJI 9
Fig. 3. Abdominal coloration of select Fiji Periscelididae, dorsal views (to same scale). A. Stenocyamops
vittatus, n. sp. B. Stenocyamops robustus. C. Stenocyamops luteus, n. sp. D, E: Stenomicra tokotaii, male (D)
and apex of female abdomen (E). F, G: Stenomicra distincta, male (F) and apex of female abdomen (G).
10 AMERICAN MUSEUM NOVITATES NO. 3671
Fig. 4. A-E: Terminal abdominal segments and syntergosternites of male Periscelididae (ventral views,
not to same scale). A. Stenocyamops robustus. B. Stenocyamops luteus/pseudoluteus. C. Stenomicra distincta.
D. Stenomicra distinctipennis. E. Stenomicra castanea. F-H: Thoraces (dorsal view) of select Fiji Stenomicra
(to the same scale). F. S. brunnea. G. S. xoutha. H. S. sylpha.