The American genera of Asilidae (Diptera): keys for identification with an atlas of female spermathecae and other morphological details. 6. Tribe Atomosiini Hermann (Laphriinae), with descriptions of two new genera and three new species, and a catalogue o

33 Pages·1991·9.4 MB·English

GayanaZool. 55(1): 53-85, 1991 1SSN0O16-531X
THE AMERICAN GENERA OF ASILIDAE (DÍPTERA): KEYS FOR
IDENTIFICATION WITH AN ATLAS OF FEMALE SPERMATHECAE
AND OTHER MORPHOLOGICAL DETAILS. VI. TRIBE ATOMOSIINI
HERMANN (LAPHRIINAE), WITH DESCRIPTIONS OF TWO NEW
GENERA AND THREE NEW SPECIES, AND A CATALOGUE OF THE
NEOTROPICAL SPECIES*
LOS GÉNEROS AMERICANOS DE ASILIDAE (DÍPTERA): CLA VES
PARA SU IDENTIFICACIÓN CON UN A TLAS DE LAS
ESPERMATECAS DE LAS HEMBRAS Y OTROS DETALLES
MORFOLÓGICOS TRIBU ATOMOSIINI HERMANN
VI.
(LAPHRIINAE), CON DESCRIPCIONES DE DOS NUEVOS GÉNEROS Y
TRES NUEVAS ESPECIES Y UN CATALOGO DE LAS ESPECIES
NEOTROPICALES
Jorge N. Artigas**, Nelson Papavero*** y Armando L. Serra'
ABSTRACT
spermathecaeandother morphologicaldetails, isgíven.Two
newgeneraaredescribed:Aphractia(forAtractialongicornis
A key for the identification of the 17 American genera of Hermann) and Cyphomyiactia (type-species, costai, sp. n.,
Atomosiini Hermann ILaphriinael, with illustrations of from Brazil,Goiás,Goiánia(Campiñas)).Twonewspeciesof
Dissmeryngodes are described: amapa (type-locality: Brazil,
Amapá, Serra do Navio) and iracema (type-locality: Brazil,
SaoPaulo, RibeiraoPreto(Fazenda Iracema).Thefollowing
newgenericsynonymiesareproposed:ParatracüaHull. 1862
synonym of Atomosia Macquart, 1838; Cyphotomyia
*This research was supported by the FundapTo de Williston, 1889 and Protichisma Hermann, synonyms of
Amparoá PesquisadoEstadodeSaoPaulo(Grants85/1772- CerotainiaSchiner, 1866;CentrolaphhaEnderlein, 1914and
5. 86/3327-1 and 87/3170-8). Catonomyia Hull, 1962 synonyms of Dissmeryngodes
Universidad de Concepción Facultad de Ciencias Hermann, 1912; Josmayala Kaletta, 1978 synonym of
Biológicas y de Recursos Naturales DepartamentodeZoolo- Eumecosoma Schiner, 1866; Lophoceraea Hermann, 1912
gía(Proyectos Dirección de Investigación. U. deConcepción synonym of Hybozelodes Hermann. 1912; Othoniomyia
N"s 20.38.12 y 20.38.17): Hermann. 1912. Sphagolestes Hull, 1962, Voluptarius
Museu de Zoología & Instituto de Estudos Kaletta, 1978 and lucundus Kaletta, 1978 synonyms of
Avanzados,UniversidadedeSaoPaulo. PesquisadordoCon- Oidardia Hermann, 1912. Rhatimomyia Lynch-Arribálzaga,
selhoNacionalde DesenvolvimentoCientíficoeTecnológico 1882isunrecognized. AcatalogueoftheNeotropicalspecies,
(CNPq. Proc. N" 30.0994/79. with several new combination, is added.
MuseudeZoología, UniversidaddeSaoPaulo. Bol-
sistadaCoordenadode Aperfei(¡oamentodo Pessoalde En Keywords Insecta. Taxonomy. America. Key. Asilidae.
sino Superior (CAPES). Atomosiini. Spermatheca. Morphology.
53
Cayana Zool. 55(1), 1991
RESUMEN ton, 1889yProtichismaHermann, 1912conCerotainiaSchi
ner, 1866: de Centrolaphria Enderlein, 1914 y Catonomyia
Sepresentaunaclaveparalaidentificacióndelos 17géneros Hull, 1962 con Dissmeryngodes Hermann. 1912; de Josma-
americanos de Atomosiini (Laphriinae), con ilustraciones de yalaKaletta, 1978conEumecosomaSchiner, 1866:de Olho-
espermatecas yotrosdetallesmorfológicos. Sondescritosdos niomyia Hermann, 1912.Sphagolestes Hull. 1962. Volupta-
nuevos géneros. Aphractia (para Aíractia longicornis Her rius Kaletta, 1978 e Iucundus Kaletta, 1978 con Oidardis
mann) yCyphomyiactia(especie-tipo,costal,sp. n.,deBrasil, Hermann, 1912. Rhatimomyia Lynch Arribálzaga, 1882 no
Goiás, Goiánia (Campiñas)). Son descritas dos nuevas espe puede ser reconocido. Se agrega un catálogo de las especies
cies de Dissmeryngodes: amapa (localidad-tipo: Brasil, Ama- neotropicales, con muchas nuevas combinaciones.
pá, Serra do Navio) e Iracema (localidad-tipo: Brasil. Sao
Paulo,RibeiraoPreto(FazendaIracema)).Seproponenlassi- Palabras claves Insecta. Taxonomía. América. Claves.
guientes nuevas sinonimias genéricas: de Paratractia Hull, Asilidae. Atomosiini. Espermateca. Morfología.
1962conAtomosiaMacquart. 1838;deCyphotomyiaWillis-
INTRODUCTION MIL Museo Cívico di Storia Naturale. Milán
MNHNP Muséum National d'Histoire Naturelle.
París
This is part VI ofa series of papers intended as a MUN Zoologische Sammlungen des Baye-
preliminary effort to define the American genera rischen Staates, Munich
of Asilidae, describing the new genera, MZUSP Museude Zoología, UniversidadedeSao
preparatory to the elaboration of Catalogue of Paulo,SaoPaulo
NAPLES Naples Museum, Italy
Neotropical species, for inclusión in the OXF Hope Department of Entomology. Ox-
forthcoming World Catalog of Flies, now being ford University, Oxford
prepared by the U.S. Department of Agriculture PHIL Academy of Natural Sciences, Phila
and U.S. National Museum of Natural History, delphia
SANT Museo Nacional de Historia Natural,
Washington, D.C.
Santiago
We have adopted a classification of the
ST Syntypes
Asilidae in 8 subfamilies. It follows, basically, the STOCKHOLM Naturhistoríska Riksmuseet.Stockholm
classification adapted by Papavero (1973), with TORO Istitutoe Museo di Zoología. Uníversítá
the elevation of the Stichopogoninae to di Toríno.Turin
subfamily rank, and the Apocleinae Papavero TP :Type(s)
UCV UniversidadCentraldeVenezuela.Cara-
are included within the Asilinae. The :
cas
Leptogastrinae are considered a subfamily of USNM UnitedStatesNationalMuseumofNatu-
:
Asilidae. Morphology and terminology we have ral History, Washington. D.C.
followed J.F. McAlpine, 1981. WASH :Washington State University. Pullman
WIEN Naturhistorisches Museum, Vienna
:
LiSTOFABBREVLATIONS
AMNH : American Museum of Natural History. The material used in this series belongs to
New York the Museu de Zoología da Universidade de Sao
BMNH : British Museum (Natural History), Lon- Paulo, Brasil and to the Departamento de
don
BUD Magyar Termésettudományi Múzeum. Zoología, Universidad de Concepción, Chile
:
Budapest (MZUC).
CRAC University ofCracovia, Poland
:
FAUCV Facultad de Agronomía, Universidad The methodology employed in the
:
Central de Venezuela. Maracay dissection and presentation of the male
FRAN :Natur-Museum und Forschungs-lnstitut terminalia, female spermathecae and other
MCZ (MSuesncekuenmbeorfg)C.omFpraarnkaftuírvteaZ.oMol.ogy. Har morphological details is the same employed by
:
vard University, Cambridge. Mass. Artigas (1971).
54
Amer. gen. Asilidae VI. Laphrünae: Atomosiini: Artigas,J.N. etai
TRIBE ATOMOSIINIHermann
Key tothe American genera
1. Anatergite with soft or coarse hairs, but never with spine-or spike-
like bristles 2
Anatergitewith characteristicspine-or spike-like bristles 12
2(1). Antenna with two flagellomeres (Fig. 2). Face extremely narrow; two very long and
stout ocellar bristles (Fig. 1). At least 4 pairsofstout, stiffbristleson upperocciput.
Mesonotum slightly bare, shining, with 1 notopleural, 1 supraalar and 1 postalar
bristles, all long and stout. Scutellum with a pair of exceptionally long and stout
marginal bristles. Abdomen parallel-sided, with fine and scattered punctures; lateral
bristles present on tergites 1-6 or 1-3. Hind fémur with (Fig. 3) or without
moderately long, tuberculate spines on apical half of ventral surface. Male
terminalia and aedeagus as in Figs. 4-8 and 11-15. Female spermathecae as in Figs.
9-10 and 16-17 (South America, but not inChile)
DissmeryngodesHermann, 1912.
Antenna witha single flagellomere. Othercombinationsofcharacters 3
3(2). Lateral bristles present on all tergites. Large (10-12 mm), robust flies. Face
entirely convex, no projection at the subcranial margin, the coarse bristles of the
mystax covering entire length of face. Abdomen, in dorsal view, with 6 tergites.
Frons with convergent slopes 4
Lateral bristles restricted to tergites 1-3 or only to tergite 1. Other combinations
ofcharacters 5
4(3). Scape twice as long as pedicel. Flagellomere three times length of scape and
pedicel together, trúncate at apex, with a dorsally placed spine. Body black. Male
terminalia as in Figs. 18-22. Female spermathecae as in Figs. 23-24 (México to
southern Brazil) AphestiaSchiner, 1866.
Scape subequal in length to pedicel. Flagellomere of variable length, but
apex prolonged intoa filiform process andspinesubapically placed. Groundcolorof
abdomen yellowish-brown, with or without dark spots and stripes (Perú, Brazil:
Bahia to RíoGrandedoSul) Aphractia, gen. n.
5(3). Frons with divergent slopes 6
Fronswithconvergentslopes 9
6(5). Vein M2 absent beyond cell d (i. e., only four posterior cells present). (USA,
México) Atomosiella Wilcox, 1937.
Vein M2 presentbeyondcelld(i. e.,five posteriorcellspresent) 7
7(6). Face and antenna with long and thick pile. Also the occiput very thickly pilóse.
Mesonotum entirely covered with semierect, thick pile. Tarsal claws thick at base
and empodial bristle developed, almost claw-like. Male terminalia as in Artigas
(1971: fig. 69). Femalespermathecaeas in Artigas(1971: fig. 70) andourFigs. 25-26
(Chile) Lamprozona Loew,1851.
Hairs of head and thorax relatively scarce. Tarsal claws and empodial
bristle not developed as above 8
55
1
GayanaZool. 5511). 1991
8(7). Eye, in lateral view, narrower on lower half, due to a recession of ¡ts postero-inferior
margin. Face slightly prominent at subcranial margin. Scutellum with smooth or
impressed rim, and with two long marginal bristles, or two long and two short
marginal bristles, or yet with several long and short marginal bristles intermingled.
Male terminaliaandaedeagusas in Figs. 27-31. Femalespermathecaeas in Figs. 32-
33 (Northand South America, but not inChile) EumecosomaSchiner, 1866.
Eye, in lateral view, as wide above asbelow (i.e., eye almost symmetrical in relation to its
equator). Face prominent or not at subcranial margin. Scutellum with short or long
marginal hairs, but never with differentiated long bristles. Male terminalia and
aedeagus as in Figs. 34-38. Female spermathecae as in Figs. 39-47 (Neotropical, but
not in Chile) Oidardis Hermann, 1912.
9(5). Minute (5 mm) flies, with globose head. Face not visible in lateral aspect.
Mystax composed ofabout 6 bristles restricted to subcranial margin offace. Thorax
and abdomen micropilose. Mesonotum with a few lateral bristles. Scutellum short
pilóse,with 2 marginalbristles (Perú) StrombocodiaHermann, 1912.
Largerflies. Head neverglobose. OtherCombinationsofcharacters 10
10(9). Face relatively wide and evenly convex. Flagellum prolonged into a filiform
process. Abdomen very wide and short, cup-shaped. Body with blue reflections.
Cyphomyia(Stratiomyidae) likeflies(Brazil: Goiás) Cyphomyiactia, gen. n.
Facefíatabove, prominentbelow. Abdomen neverasabove 1
1 1(10).Antennal flagellomere with acute apex, sometimes prolonged into a more or
less filiform process and with a subapically or dorsally placed spine (if dorsal, spine
always placed distally to middle of flagellum). Male terminalia and aedeagus as in
Figs. 48-52. Female spermathecaeasin Figs. 53-56 (CentralandSouth America,but
not in Chile) Hybozelodes Hermann, 1912.
Antennal flagellomere trúncate at apex and spine always placed dorsally and
alwayssituated before middle offlagellum. Male terminalia andaedeagusas in Figs.
57-61. Female spermathecae as in Figs. 62-65 (Brazil: Amazonia and southern
states) Lyeosimyia Hull, 1958.
12(1). Antenna with two flagellomeres 13
Antenna with one flagellomere 14
13(12).Scape and pedicel subequal in length. Frons with divergent slopes. Male
terminalia and aedeagus as in Figs. 66-70. Female spermathecae as in Figs. 71-72
(México tosouthern Brazil) Atoniomyia Hermann, 1912.
Scape about three times as long as pedicel. Frons with (?) convergent slopes
(Panamá, Venezuela, Perú) Bathropsis Hermann, 1912.*
14(12).Frons relatively narrow, with convergent slopes. Scape never several times
longer than pedicel 15
Frons extremely widened, with slopes definitely divergent (Fig. 89). Scape
several times longer than pedicel (Fig. 90). Male terminalia and aedeagus as in Figs.
Not seen.
56
1
Amer. gen. Asilidae VI. Laphriinae: Atomosiini: Artigas, J.N. etal.
91-95, 98-102. Female spermathecae as in Figs. 96-97, 103-104 (North and South
America, but not in Chile) CerotainiaSchiner, 1866
15U4).Flagellum with acute apex prolonged into a filiform process, and spine
subapically placed (Southern Brazil) Atractia Macquart, 1838
Flagellum trúncate at apex, with spinedefinitely dorsal in position 16
16(15).Frons extremely shallow. Pleura, mesonotum and abdomen very coarsely punctate.
Scutellum with only a pair of fine, stiff, marginal hairs. Lateral bristlesconfined to
first two tergites and replaced by spiky pile on tergites 3-6. Apex of abdomen
strongly cupped. Pronotum with a collar ofspikelike bristles. Occipital bristles very
weak. Female spermathecae as in Figs. 73-74 (Brazil: Distrito Federal).
HoditesHull, 1962
Frons deeply excavated. Body punctures variable. Scutellum with several
marginal hairs, or with 2 to 6 or more spikelike bristles. Lateral bristles variable:
from confined to tergite 1 to present on all tergites. Pronotum with orwithout long
spikelike bristles. Occipital bristles variable. Male terminalia and aedeagus as in
Figs. 75-79 and 82-86. Female spermathecae as in Figs. 80-81 and 87-88 (Americas,
but not in Chile) Atomosia Macquart, 1838.
GenusAphestiaSchiner Materialexamined
Aphestia Schiner, 1866:663 (key), 673. Type- Brazii RondóNIa Pimenta Bueno.x¡. 1960(Alvarengal. 19
species, brasiliensis Schiner (orig. des.) . Amapa Mazagao. Jaú ao Vila Nova, 1958 (Damascenol. 1
= annulipes (Macquart). 9. Mato Grosso Utianti, Rio Papagaio. xi.1966 (Lenko&
Pereiral. I9. MnasGeraiS: Macaúba,nodate(Pereiral. 1 9
; Pouso Alegre, xii.1952 (Pereiral. 1 9 . Goias: Goiánia
(Campiñas), i.1936 (Spitzi. 1 9 . RiodeJaneiro Muri,Nova
Aphestiaannulipes(Macquart) Friburgo, i.1966 (Guimaraesl. 2d. 29 (one withspermathe
cae preserved on vial on same pin); Itatiaia, 700 m, ii.1955
Atomosia annulipes Macquart, 1838:74 (Zikán), 1 9 ; Magé. iii.1940 (Shannon), 1 9 . Sao Paulo
(1839:190). Type-locality: "Brazil". TP Arafatuba, RioJacaretinga. x.1961 (Lañe & Rabello), 1 9 ;
MNHNP. Praia Grande, Ilha de Santo Amaro, xi.1969 (Valí. I 9 ;
PraiaGrande(Fazenda Rondónial. ii.1945(Carreral. 3rf,Ju
Atomosia affinis Macquart, 1850:379 (1850:75). quiá, no date (Lañe). I 9 ; Alto da Serra. xii.1925. xii.I927
Type-locality: "Brazil". TP MNHNP (Spitzi.29 ; RioClaro, xii.1930lBorgmeier). I 9 ;SaoPaulo
N. SYN. (Cantareira.Chapadaol. xi.1946(Barreltol. 39 ;do., xi.195
Aphestia brasiliensis Schiner, 1866:378. Type- (Carrera & d'Andretta). I 9 ; Sao Paulo (Horto Florestall.
locality: "Brasil". TP WIEN. bxeil.l1o9).46I (9Car;rdeor.a,).111.119934; (SSapiotzPi.aulleoí'(dlop.i,rain.g1a9)4.81(11D.e1n9t4e8).(RIa<S
Aphestia calceata Schiner, 1867:379. Type- Itermmaha in vial on same pin);Sao Paulo, v. 1923. iv.1926
locality: "Brazil". TP WIEN. Ref.- Her- (Barbiellinil, 39 ,2&Paraná Curitiba. x.1936.1.1940(Cía
mann, 1912:123 (Synonymy). retianol. 2d\ I 9 ; Rio Negro, vn.1929, co collector. 1 9
Aphestia calcarata Williston, 1901:317, error. TSeaunttóaniaC,at¡ia.1r9i5n4aiPBllauummeannna)u.. 11.9196.4All(CainrrMerZaUl,SP|.9 ; Nova
There exists a considerable variation in the
color of legs and in the shape of cell r5 (from wi-
dely open to closed and petiolate). We interpret AphestianigraBigot
thisspeciesascontaining those representativesof
Aphestia with mixed black and white bristles in Aphestia nigra Bigot, 1878:235. Type-locality:
the mystax. "México". TP OXF.
57
Cayana Zool. 55(1), 1991
Aphestia mexicana Williston, 1901:317, pl. 6, vivax (Hermann), 1912:177, fig. 67 (Atractia).
fig. 2. Type-locality: México, Guerrero: Type-locality: Perú, Umuhuankiali,
Amula; Veracruz: Atoyac. ST BMNH. Urubamba R., 500 m, and "Perú". ST ?
N. SYN. N. COMB.
Atomosiaandrenoides Bromley, 934:339.Type-
1
locality: Guyana, Bartica, Kartabo. TP
AMNH. N. SYN. GenusAtomosia Macquart
Asoccurs with A. annulipes, there is a great
variation in the coloroflegsand theshape ofcell Atomosia Macquart, 1838:73 (1839:189). Type-
r$. We are considering as this species specimens species, incisuralis Macquart (Coquillett,
1910:512) = puella (Wiedemann).
with entirely white mystax.
Paratractia Hull, 1962:384. Type-species,
Laphria dasvpus Wiedemann (orig.
Materialexamined: des.). N. SYN.
BRAZ1L. Amazonas Manaus, xi.1957(Elias& Ropa), 1 9 . anonyma Williston, 1901:316. Type-locality:
Distrito Federal Corumbá(FazendaMonjolinho),ii.1945 México, Guerrero, Chilpancingo. TP
(Barreno), 11 specimens. GoiÁs Goiania (Campiñas), AMNH, BMNH.
xii.1935 (Spitz), lcf. RiodeJaneiro Itatiaia,700m,ii.1948,
ii.1950(Zikán),29 .SaoPaulo NovaEuropa(Fazendalta- argyrophora Schiner, 1868:169. Type-locality:
queré), v.1964 (Lenko). 1 9 ; Barueri. n.1955 (Lenko). 1 9 ; "Brazil". TP WIEN.
Sao Paulo (Cantareira, Chapadao), xi.1946 (Barretto), leí. armata Hermann, 1912:152, fig. 58. Type-
APaRrGaEnNáTIMaNtAi.nhosC,óriid.o19b4a5, nDoepctool.lecSt.or,Malre?t.in, i.1950. no locality: "Brazil". TP MUN, MIL.
cAolllleicntoMr,ZU1ScfP.. harbiellinSiiaoCuPrarualno,. 1T9P35:A7M.NTyHpe.-locality: Brazil,
beckeriJaennicke, 1867:359. Type-locality: "Mé-
xico". TP MUN.
Aphractia, gen. n. bequaerti Bromley, 1934:339. Type-locality: Bra-
zil, Para, Santarém. TP USNM.
Very similar to Aphestia, differing however in bigoti Bellardi, 1861:120 (20). Type-locality:
the following characters: scape subequal in "México". TP TORO.
length to pedicel; flagellomere of variable length, cen'erai Bromley, 1929:282, fig. 8. Type-locality:
but apex prolonged into a filiform process and Cuba, Soledad. TP USNM.
spine subapically placed; abdomen more or less coxalis Curran, 1930:18. Type-locality: Brazil,
parallel-sided,moreslender than inAphestia,and Mato Grosso, Corumbá. TP AMNH.
yellowish-brown in ground color, with or cyanescens Rondani, 1848:92. Type-locality:
without dark spotsand stripes. Wingswith cell r_<¡ "Brazil". TP NAPLES.
open (in the very few specimens examined; there danforthi Curran, 1935:8. Type-locality: Puerto
may be a strong variation, as in Aphestia). Rico, Coamo Springs. TP AMNH.
Type-species: Atractia longicomis Her- dasypus (Wiedemann), 1828:527 (misprinted as
mann. 257) {Laphria). Type-locality: "Brazil".
N
TP COMB.
?
fredericoi Carrera, 1952:209. Type-locality: Bra-
Listofspecies zil, Sao Paulo, Sao Paulo (Ipiranga). TP
MZUSP.
longicomis (Hermann), 1912:190, fig. 74 (Atrac- frontalis Curran, 1930:17. Type-locality: Brazil.
tia).Type-locality: Brazil, Bahia. TP? N. Mato Grosso, Chapada dos Guimaraes.
COMB. TP AMNH.
rubida (Hermann), 1912:179 (/lrrac<7fl).Type- geniculata (Wiedemann), 1821:241 (Laphria).
locality: Brazil, Rio Grande do Sul. TP Type-locality: "Brazil". TP ?WIEN.
WIEN. N. COMB. hondurana James, 1953:53. Type-locality: Hon-
58
Amer. gen. Asilidae VI. Laphriinae: Atomosiini: Artigas, J.N.etal.
duras, Zamorano, Escuela Agrícola Pa- bo, Maruiara. TP UCV.
namericana. TP MICH. rufipes Macquart, 1847:55 (1847:39). Type-
limbiventris Thomson, 1869:466. Type-locality: locality: USA, Pennsylvania. Distr.-
Uruguay, Montevideo. TP USA, México. TP MNHNP.
STOCKHOLM. eupoda Bigot, 1878:234 [Cormansis). Type-
limbativentris Lynch Arribálzaga, 1880:50, locality: "México". TP OXF.
emend. sóror Bigot, 1878:236. Type-locality: "Méxi-
lineata Curran, 1930:19. Type-locality: Brazil, co". TP OXF.
Mato Grosso do Sul, Corumbá. TP punctifera Hermann, 1921:161 (as rufipes
AMNH. var.). Type-locality: "México". TP
macquarti Bellardi, 1861:120 (20). Type-locality: MUN.
"México". TP TORO. ?scoriacea (Wiedemann), 1828:529 (Laphria).
maestrae Bromley, 1929:287, fig. 7. Type- Type-locality: "Brazil". TP?
locality: Cuba, Sierra Maestra. TP setene Curran, 1935:7. Type-locality: Brazil, Sao
USNM. Paulo. TP AMNH.
melanopogon Hermann, 1912:144, fig. 53. Type- setosa Hermann, 1912:145, fig. 54. Type-
locality: USA, Texas. Distr.- USA, Mé- locality: "Brazil". TP MIL, MUN.
xico, Honduras. TP ?WIEN. tenuis Curran, 1930:19. Type-locality: Brazil,
metallescens Hermann, 1912:150, fig. 57. Type- Mato Grosso, Chapada dos Guimaraes.
locality: Argentina, Mendoza. TP TP AMNH.
MUN, WIEN. tibialis Macquart, 1846:204 (1846:76). Type-
metallica Bromley, 1929:283. Type-locality: Cu- locality: México, Yucatán, Mérida. TP
ba, Sierra Maestra, Palma Mocha. TP ?lost.
USNM. unicolor Macquart, 1838:74 (1839:190), pl. 7,
mucida Osten Sacken, 1887:184. Type-locality: figs. 2-3.Type-locality: Brazil, RioGran-
México, Sinaloa, Presidio. TP BMNH. de do Sul. TP MNHNP.
nigroaenea Walker, 1851:154. Type-locality: Co- venustula Lynch Arribálzaga, 1880:50. Type-
lombia, Bogotá (in label of type). TP locality: Argentina, Buenos Aires, Las
BMNH. Conchas and Gran Chaco. TP lost.
nuda Hermann, 1912:147, fig. 55. Type-locality: xanthopus (Wiedemann), 1828:529 [Laphria).
Trinidad, Belmont (in label of type). TP Type-locality: "Brazil". TP?
MUN.
panamensis Curran, 1930:17. Type-locality: Pa-
namá, Canal Zone, Barro Colorado I. GenusAtomosiellaWilcox
TP AMNH.
pilipes Thomson, 1869:465. Type-locality: Ar- Atomosiella Wilcox, 1937:40. Type-species,
gentina, Buenos Aires. TP Atomosia antennata Banks (orig. des.).
STOCKHOLM. antennata (Banks), 1920:66 [Atomosia). Type-
pilosipes Lynch Arribálzaga, 1880:52, locality: USA, Arizona. Distr.- USA,
emend. México (Baja California). TP ?MCZ.
pubescens Bromley, 1929:281, pl. 1, fig. 3. Type-
locality: Cuba, Soledad. TP MCZ.
puella (Wiedemann), 1828:531 {Laphria). Type- GenusAtoniomyiaHermann
locality: unknown. TP WIEN.
incisuralis Macquart, 1838:76 (1839:192), pl. Atonía Williston, 1889:257 (preocc. Gistel,
7, fig. 1. Type-locality: "Cuba". TP 1848). Type-species, Atomosia mikii
MNHNP.
Williston (Williston, 1901:316).
rica Curran, 1935:9. Type-locality: Puerto Rico, Atoniomyia Hermann, 1912:81 (nom. nov. for
Mayaguez. TP AMNH. Atonía Williston). Type-species,
rosalesi Carrera & Machado-Allison, 1963:242, Atomosia mikii Williston (aut.).
fig. 1. Type-locality: Venezuela,Carabo- Neatonia Bromley, 1935:130. Unavailable ñame
59
Gayana Zool. 55(1), 1991
(no type-species designated). transferred to the genera Hybozelodes, Lycosím-
albifacies (Hermann), 1912:85 (key), 97 (descrip- yia and Aphractia. Only two species remain in
tion, as albiceps (sic)), fig. 38 (Atonía). Atractia:
Type-locality: Brazil, Mato Grosso, psilogaster (Wiedemann), 1828:456 (Asilus).
Cuiabá. TP BUD. Type-locality: "Brazil". TP WIEN.
ancylocera (Schiner),1868:170 (Atomosia). Type- coronata Schiner, 1867:412. Type-locality:
locality: "Venezuela". TP WIEN. "Brazil". TP WIEN.
brevistylata (Williston), 1901:316, pl. 6, fig 1 pulverulenta Schiner, 1867:412. Type-locality:
(Atonía). Type-locality: México, Tabas- "Brazil". TP WIEN.
co, Teapa. TP BMNH.
fulvipes Carrera, 1946:122, figs. 7, 10, 20. Type-
locality: Brazil, Mato Grosso doSul, Sa- GenusBathropsisHermann
lobra. TP MZUSP.
grossa Carrera, 1946:125, figs. 9, 3, 12. Type- Bathropsis Hermann, 1912:68. Type-species, pe-
locality: Brazil, Sao Paulo, Severinia. TP ruviana Hermann (orig. des.).
MZUSP.
hispidella (Hermann), 1912:95, fig. 37 (Atonía). basalis Curran, 1930:6, fig. 1. Type-locality: Pa-
Type-locality: Brazil, Sao Paulo. TP namá, Canal Zone, Barro Colorado I.
MIL, MUN. TP AMNH.
laterepunctata (Hermann), 1912:93, fig. 36 (Ato- delgadoi Kaletta, 1978:56. Type-locality: Vene-
nía). Type-locality: Perú, mouth of zuela, Aragua, Hacienda El Periquito,
Pachitea River, Umuhuankiali at R. Ocumare de la Costa. Type, FAUCV.
Urubamba and Puerto Yessup; Bolivia, peruviana Hermann, 1912:69, fig. 23, pl. 2, figs.
Mapiri, Sarampiuní; Perú, Vilcanota. 13-14. Type-locality: Perú, Vilcanota.
ST WIEN, MIL, BUD, BMNH. TP MUN.
mikii (Williston), 1886:290 (Atomosia). Type-
locality: "San Domingo". TP ?lost.
mollis (Hermann), 1912:88, fig. 33 (Atonía). Genus CerotainiaSchiner
Type- locality: Perú, Unini at Ucayali R.
and Vilcanota. ST BUD, MUN. Cerotainia Schiner, 1866:662 (key), 673
pinguis (Hermann), 1912:90, fig. 34 (Atonía). (1868:170, second erection of genus).
Type-locality: Perú, mouth of Pachitea Type-species, Laphria xanthoptera
R. TP MUN. Wiedemann (orig. des.).
scalarata (Hermann), 1912:92, fig. 35 (Atonía). Ceratotaenia Lynch Arribálzaga, 1880:52,
Type-locality: Perú, Rosalina at Uru emend.
bamba R.; Bolivia, Mapiri, Chimate. ST Cyphotomyia Williston, 1889:257. Type-species,
WIEN, MUN, BMNH. lynchii Williston (orig. des.). N. SYN.
setigera (Hermann), 1912:86, pl. 3, figs. 24-25 Protichisma Hermann, 1912:35. Type-species,
(Atonía). Type-locality: "Brazil". TP longimanus Hermann (orig. des.). N.
WIEN. SYN.
viduata (Wiedemann), 1819:50 (Laphria). Type-
locality: "Brazil". TP WIEN. albibarbis (Curran), 1930:7 (Protichisma). Type-
locality: Panamá, Canal Zone, France
Field. TP AMNH. N. COMB.
GenusAtractia Macquart argyropasta Hermann, 1912:61. Type-locality:
Perú, Ucayali R. TP?
Atractia Macquart, 1838:151 (1839:267). Type-
species, Asilus psilogaster Wiedemann argyropulsoSmcbhiian"er(,in1e8r6r8o:r;17a0c.tuTaylpley-lVoceanleiztuy:el"aCoi-n
(mon.). type label). TP WIEN.
argyropyga Hermann, 1912:52, fig. 16. Type-
Most of the species included here are now locality: "Perú". TP MUN.
60
Amer. gen. Asilidae VI. Laphriinae: Atomosüni: Artigas, J.N. etal.
aurataSchiner, 1868:171. Type-locality: "Colom- Type-locality: "México". TP TORO.
bia". TP WIEN. ornatipes James, 1953:54. Type-locality: Hondu-
bella Schiner, 1867:380. Type-locality: "Brazil" ras, Zamorano. TP WASH.
(probably Rio de Janeiro State, as it was propinguaSchiner, 1868:171. Type-locality: "Co-
collected by Bescke). TP WIEN. lombia" (in error; actually Venezuela in
brasiliensis Schiner, 1867:379. Type-locality: type's label). TP WIEN.
"Brazil" (As the preceding species, this rhopalocera Lynch Arribálzaga, 1882:189 (as
one was also collected by Bescke, accor- Ceratotaenia). Type-locality: Argentina,
ding to the label ofthe type). TPWIEN. Buenos Aires, Chacabuco. TP lost.
camposiCurran, 1934:3. Type-locality: Ecuador, unicolor Hermann, 1912:64, fig. 22. Type-
Posorja. TP AMNH. locality: "Perú". TP MUN.
dasythrix Hermann, 1912:48, fig. 13. Type- violaceithorax Lynch Arribálzaga, 1880:52 (as
locality: Perú, mouth of Pachitea R.; Ceratotaenia). Type-locality: Argentina,
Bolivia, Mapiri, Sarampiuní, 700 m. ST Buenos Aires. TP lost.
MUN. willistoni Curran, 1930:14. Type-locality: Brazil,
debilis Hermann, 1912:59, fig. 21. Type-locality: Mato Grosso, Chapada dos Guimaraes.
Perú, mouth of Pachitea R., 150 m; TP AMNH.
Chanchamayo, 800 m; Bolivia, Saram- xanthoptera (Wiedemann), 1828:530 (Laphria).
MUN
piuní; Perú, Vilcanota. ST Type-locality: unknown ("America" on
WIEN, BMNH. type label). TP WIEN.
dubia Bigot, 1878:238. Type-locality: "México".
TP OXF.
femínea Curran, 1930:14. Type-locality: Pana Genus Cyphomyiactia,gen. n.
ma. Canal Zone, Corozal. TP AMNH.
This beautiful fly, with bluish reflections on the
flavipes Hermann, 1912: 57, fig. 20. Type- body, somewhat resembles the stratiomyid genus
locality: Perú, Chanchamayo, 800 m;
Cyphomyia.
Paraguay, Asunción. ST MUN, BUD.
jamaicensis Johnson, 1919:430 (as macrocera Head wider than high. Face relatively wide,
var.). Type-locality: "Jamaica". TP nearly 3/5 máximum width of an eye, evenly
MCZ. convex, densely tomentose; bristles of mystax
laticeps Bromley, 1929:279, pl. 1, fig. 2. covering entire face, longer on ventral half.
Type-locality: Cuba, Zapata, Santo To- Proboscis relatively short, robust, subcylindrical,
más. TP USNM. N. SYN. obliquely directed downwards, reaching apex of
leonina Hermann, 1912:49, fig 14. Type-locality: bristles of mystax. Palpus short, with several
Argentina, Mendoza. TP MUN. apical bristles. Antenna attached above middleof
longimana (Hermann), 1912:37, pl. 1, figs. 7-8 head, long and slender; scape about 1.5 times
(Protichisma). Type-locality: Perú, length of pedicel, with 2 bristles on ventral
Callanga. TP BUD. N. COMB. surface; pedicel with 1 more or less short bristle
lynchii (Williston) 1889:258 (Cyphotomyia). dorsally; flagellum elongate, thickened, a Hule
Type-locality: Brazil. Mato Grosso, compressed laterally, stronglyacuminate atapex,
Chapada dos Guimaraes. Distr. Brazil over 3 times the combined length of scape and
(MatoGrosso, Sao Paulo), Perú, Bolivia. pedicel, prolonged into a filiform process; a
TP ?AMNH. N. COMB. definite spine dorsally placed on the second third
marginata Hermann, 1912:62. Type-locality: Pe- of the flagellum. Frons short, as wide as face,
rú, Pachitea R. TP? with convergent slopes; 3 more or less strong
mínimaCurran, 1930:12.Type-locality: Panamá, orbital bristles. Ocellar tubercle with 2 divergent
Canal Zone, Barro Colorado I. TP bristles. Occiput tomentose, with strong bristles
AMNH. above, some weaker ones laterally, and ventrally
nigra Bigot, 1878:238. Type-locality: "México". with sparse, long, fine pile.
TP OXF. Mesonotum arched, moderately pilóse, the
nigripennis (Bellardi), 1861:119 (19) {Atomosia). pilesemierect. Prothorax with severalerect hairs.
61
Cayana Zool. 55(11. I1)1»I
Proepisternum with several hairs. Postpronotal áreas. Antepronotum predominantly golden
lobe (humerus) with semierect pile. tomentose, with a brownish-black stripe on the
Anepislernum with sparse long hairs and 3 long anterior borderand yellow hairs. Lateral margins
and strong bristles. Anatergite with short hairs. of antepronotum silverywhite tomentose.
Katatergite with long reclínate bristles. 1 Postpronotum brownish-golden tomentose, hairs
notopleural, 1 supraalar and 1 postalar bristles brown. Proepisternum silverywhite tomentose,
present. Posterior callus with 3 bristles. as well as the anepisternum, which shows a
Scutellum dorsally with fine, recumbent hairs polished brown spot and brown hairs.
and several marginal bristles. Anepimeron brown, with white pollinosity.
Legs: All femora equally swollen; hind tibia Anatergite goldish-silvery-white pollinose, with a
a little swollen on the distal half and slightly fringe of yellowish hairs behind posterior callus.
arcuate; all tibiae with long bristles dorsally, Meron brown, white pollinose, with afew brown
anteriorly and posteriorly and ventrally with a hairs. Katatergite brown, silverywhite
brush of short pile extending to tarsomeres. tomentose, with brown bristles. Posterior callus
Abdomen characteristically short, broad brown.
and robust, wider than thorax, strongly cupped, Wings hyaline, sligtly fumóse along veins.
shining blue. First segment slightly wider than Halteres yellow.
second. The entire abdomen strongly arched and Abdomen blackish in ground color, with
curved downwards, punctured, with modérate blue shine. Hairs of abdomen white. Tergites 3-4
pile. Tergite 1 with 10-12 strong lateral bristles; with posterolateral golden and silverywhite
the remaining tergites only with pile laterally. tomentose spots.
Type-species, Cyphomyiactia costai, sp. n. Holotype 9. BRAZIL, Goiás: Goiánia
(Campiñas), 1935 (Borgmeier & Souza Lopes), in
MZUSP.
the
Cyphomyiactiacostai,sp. n.
This species is dedicated to Prof. Dr.
Face densely silverywhite tomentose. Hairs of Newton C. A. da Costa (Instituto de Estudos
mystax white. Occiput golden tomentose at Avancados, Universidade de Sao Paulo).
margins, brownish-black tomentose around
foramen, with long, sparse, fine white pile GenusDissmeryngodesHermann
bellow; dorsal occipital bristles yellowish. Scape
brown, pedicel brownish-yellow, flagellum red- Dissmeryngodes Hermann, 1912:75. Type-
brown and white pollinose. Frons golden species, Laphria antica Wiedemann
tomentose, with a slender black stripe running (orig. des.).
from anteriorocellusto baseofantennae. Vértex Centrolaphria Enderlein, 1914:241. Type-
black. species, columbiana Enderlein (orig.
Thorax blackish in ground color, with des.) = nigripes (Macquart). N. SYN.
bluish reflections. Pleura with mixed silvery- Catonomyia Hull. 1962:380. Type-species.
white and golden-brown tomentum in some spiculata Hull (orig. des.). N. SYN.
KEY TO SPECIES:
1. Hind fémur only with short white hairs on apical half of ventral surface.
Wing hyaline. Legs yellow, except for hind fémur and tibia, which are black, with
yellow at base and apex. Tergites 1-6 with lateralbristles(Brazil: southern states)
anticus (Wiedemann)
Hind fémur with 4-5 strong, tuberculate, black or white spines on apical half
of ventral surface
62