Temperate Fruit Crop Breeding James F. Hancock Editor Temperate Fruit Crop Breeding Germplasm to Genomics JamesF.Hancock MichiganStateUniversity Dept.Horticulture EastLansingMI48824 USA [email protected] ISBN978-1-4020-6906-2 e-ISBN978-1-4020-6907-9 LibraryofCongressControlNumber:2007939760 (cid:2)c 2008SpringerScience+BusinessMediaB.V. Nopartofthisworkmaybereproduced,storedinaretrievalsystem,ortransmitted inanyformorbyanymeans,electronic,mechanical,photocopying,microfilming,recording orotherwise,withoutwrittenpermissionfromthePublisher,withtheexception ofanymaterialsuppliedspecificallyforthepurposeofbeingentered andexecutedonacomputersystem,forexclusiveusebythepurchaserofthework. Coverphotographby:JosephD.PostmanattheUSNationalClonalGermplasmRepository,Corvallis, Oregon Printedonacid-freepaper 9 8 7 6 5 4 3 2 1 springer.com Preface Thisbookisintendedtobeabriefcompilationoftheinformationavailableonthe breedingoftemperatefruitcrops.Thegoalistoprovideoverviewsontheevolution ofeachcrop,thehistoryofdomestication,thebreedingmethodsemployedandthe underlying genetics. A serious effort is made to fully integrate conventional and biotechnologicalbreeding approaches. A discussion is also provided on licensing andpatenting. Itishopedthatthisbookcanbeusedasa springboardforbreedersdesiringan update,horticulturalistswhowonderwhatthefruitbreedersaredoingandgenomi- cistswhoaresearchingforawaytocontributetofruitbreedingefforts.Byfarthe fastestprogresscanbemadewhenwealltalkthesamelanguage. ThismanuscriptisinmanyregardsanupdateoftheinformationfoundinFruit Breeding,Volumes1and2,editedbyJ.JanickandJ.N.Moore(1996).Themajor differenceisthatmuchmoremolecularinformationisnowavailableonfruitcrops. Molecular linkage maps have been producedfor many of the commercialspecies and the first quantitative trait loci are being tagged and selected through marker assistedbreeding.Regenerationandtransformationsystemsareavailableformany ofthefruitcropsandpotentiallyusefulgeneshavebeenclonedandcharacterized. Fruitbreederswill soon haveall the tools in their toolboxthat the grainbreeders havehadforoveradecade. Theatmosphererevolvingaroundfruitbreedershasalsochangedinanothersig- nificant way since Janick and Moore’s book. While fruit breeding has long been conductedmostprominentlyinthepublicdomain,privatecompanieshavebecome moreactiveinthe breedingoffruitcrops,aslargemarketingorganizationsdecide thattheywanttobesureofhavinghighqualityvarietiesinthefuture.Infact,most publicbreedershavenowbecomeprivatebreeders,astheypatentandlicensetheir cultivarstothesemarketingagencies. Iwanttopubliclythanksomeofthepeoplewhomadethisbookpossible.Imust first acknowledgethe helpof Gustavo Lobos,whose willingnessto tackle the ref- erencesandfinalformatwas a hugehelp.I also needtothankRex Brennen,John Clark, Ross Ferguson,Craig Ledbetter,Amy Iezzoni,ChrisOwensand especially ChadFinn,fortheiragreeingtospearheadchapters,whenitbecamecleartomethat Iwouldneverfinishthisprojectwithouthelp.SusanBrown,JimLuby,PaulLyrene, BillOkie,RalphScorza,TomSjulinandNickVorsashouldalsobeacknowledgedas v vi Preface coauthorswhogavechapterscredibilitywhichtheywouldnothavehadotherwise. Veryspecialthanksalsoneedstogotomywife,AnnHancock,whocontinuestobe supportiveofallmyendeavors,saneorcrazy. EastLansing,MI,USA J.F.Hancock Contents 1 Apples ........................................................ 1 J.F.Hancock,J.J.Luby,S.K.BrownandG.A.Lobos 2 Apricots....................................................... 39 C.A.Ledbetter 3 Blackberries................................................... 83 C.E.Finn 4 BlueberriesandCranberries ....................................115 J.F.Hancock,P.Lyrene,C.E.Finn,N.VorsaandG.A.Lobos 5 Cherries ......................................................151 A.F.Iezzoni 6 CurrantsandGooseberries......................................177 R.M.Brennan 7 Grapes........................................................197 C.L.Owens 8 Kiwifruit......................................................235 A.R.FergusonandA.G.Seal 9 Peaches .......................................................265 J.F.Hancock,R.ScorzaandG.A.Lobos 10 Pears .........................................................299 J.F.HancockandG.A.Lobos 11 Plums.........................................................337 W.R.OkieandJ.F.Hancock vii viii Contents 12 Raspberries ...................................................359 C.E.FinnandJ.F.Hancock 13 Strawberries...................................................393 J.F.Hancock,T.M.SjulinandG.A.Lobos 14 IntellectualPropertyRightsforFruitCrops.......................439 J.R.ClarkandR.J.Jondle Contributors R.M.Brennan ScottishCropResearchInstitute,Invergowrie,DundeeDD25DA,Scotland,UK [email protected] S.K.Brown Department of Horticultural Sciences, 630 W. North Street, Cornell University, Geneva,NewYork14456,USA [email protected] J.R.Clark DepartmentofHorticulture,316PlantScience,UniversityofArkansas,Fayetteville, Arkansas72701,USA [email protected] A.R.Ferguson TheHorticultureandFoodResearchInstituteofNewZealandLtd(HortResearch), MtAlbertResearchCentre,PrivateBag92169,Auckland,NewZealand [email protected] C.E.Finn USDA-ARS,HorticulturalCropsResearchUnit,3420NWOrchardAvenue,Corvallis, Oregon97330,USA fi[email protected] J.F.Hancock DepartmentofHorticulture,342CPlantandSoilSciencesBuilding,MichiganState University,EastLansing,Michigan48824,USA [email protected] A.E.Iezzoni DepartmentofHorticulture,342BPlantandSoilSciencesBuilding,MichiganState University,EastLansing,Michigan48824,USA [email protected] ix x Contributors R.J.Jondle PatentandTrademarkAttorney,JondleandAssociates,P.C.,858HappyCanyonRoad, Suite230,CastleRock,Colorado80108,USA [email protected] C.A.Ledbetter USDA-ARS,9611S.RiverbendAvenue,Parlier,California93648,USA [email protected] G.A.Lobos LaboratoriodeEcofisiologiaVegetal,EscueladeAgronomia,UniversidaddeTalca, Chile [email protected] J.J.Luby DepartmentofHorticulturalSciences,329AldermanHall,UniversityofMinnesota, StPaul,Minnesota55413,USA [email protected] P.A.Lyrene HorticulturalSciences,2135FifieldHall,UniversityofFlorida,Gainesville,Florida 32606,USA lyrene@ufl.edu W.R.Okie USDA-ARS,SEFruitandTreeNutResearchLab,21DunbarRd,Byron,Georgia, 31008USDA [email protected] C.L.Owens USDA-ARS,GrapeGeneticsResearchUnit,308SturtervantHall,CornellUniversity, Geneva,NewYork14456,USA [email protected] R.Scorza USDA-ARS,45WiltshireRd,Kearneysville,WestVirginia25430,USA [email protected] A.G.Seal TheHorticultureandFoodResearchInstituteofNewZealandLtd(HortResearch), TePukeResearchCentre,RD2,TePuke,NewZealand [email protected] Contributors xi T.M.Sjulin DriscollAssociates,629CarpenteriaRoad,Aromas,California95004,USA [email protected] N.Vorsa ThePhilipE.MarucciCenterforBlueberryandCranberryResearchandExtension, 125ALakeOswegoRd,Chatsworth,NJ08019,USA [email protected]
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