Acta Zootaxonomica Sinica, 30 (3) : 590- 597 (July, 2005) ISSN 1000 0739 动物分类学报 TAXONOMY OF THE GENUS THORACOCHIRUS ( COLEOPTERA , STAPHYLINIDAE, OSORIINAE) FROM CHINA WU Jie1,2, ZHOU Hong Zhang1 1. Instituteof Zoology, ChineseAcademyof Sciences, Beijing100080, China 2. GraduateSchoolof ChineseAcademyof Sciences, Beijing100039, China Abstract The genus ThoracochirusBernhauer, 1903, is asuccessful rove beetle group andwell adapted to habitats associated with leaf litter layer and deadwood. Before ourstudy, onlytwospecies, Thoracochirus variolosus Fauvel, 1895, and T. for mosaeCameron, 1940, were recordedfrom Taiwan, China. In this paper, we describe three new species, T. yingjiangensis sp. nov. , T. protumidussp. nov. and T. arcuatussp. nov. , from Yunnan, China. T. variolosusis recordedfrom main land China for thefirst time. A key tothesefourspecies is presented. All thespecimens, including the types, are deposited in Institute of Zoology, ChineseAcademy of Science. Key words Coleoptera, Staphylinidae, Thoracochirus, newspecies. The tribeLeptochirini (Staphylinidae, Osoriinae) HW: head width; PW: pronotum width; EW: width is an interesting group withspecial bodyform and head of both elytra together. This study is based on the morphology and is successfully adapted to live under specimens in the collection of IOZ CAS (Institute of the bark of dead wood. There are four genera in this Zoology, Chinese Academy of Sciences). All types are tribe, namely Leptochirus Germar, Borolinus Bern deposited in the same institute. hauer, Priochirus Sharp, and Thoracochirus Bern Thoracochirus Bernhauer, 1903 New record to hauer, which are known to occur in tropical and sub mainland China tropical regions. Amongst them, Thoracochirus can be ThoracochirusBernhauer, 1903: 116, 155, 160; Bernhauer and Schu readily distinguished from other genera by distinctly bert, 1910: 18; Cameron, 1925: 6; Bernhauer, 1926: 254; having rugous andsetiferous surface. Before westarted Cameron, 1930:118; Scheerpeltz, 1933:1008; Shibata, 1973:27; this study, only 21 species were discovered in this Hammond, 1984: 199; Herman, 2001: 1142. Typespecies: Lep tochirus rugosus Fauvel, fixed by subsequent designation by Black genus worldwide. The species of this genus are usually welder, 1952: 388. known as leaf litter layer and/or under bark dwellers This genus is readily distinguishedfrom other gen and mainly distributed in tropical forests of South Asi era of the Leptochirini by strongly transverse prono a. As the north part of Oriental Region, South China tum, distinctly corser and rugouser surface. Head less is zoogeographically important for this genus. Howev armed, at most with a pair of triangular lateral teeth, er, since 1940s, nospecific taxonomic study have been frontal impression broadly impressed, mandible exter conducted for this genus. Before our study, only 2 nally sulcate throughout, maxillary papi 4 segmented, species, Thoracochirus variolosus Fauvel, 1895, and 3rd quadrate, 4th strongly elongate, about four times T. formosae Cameron, 1940, were recorded from as long as 3rd, inner lobe of maxilla denticulate, men Taiwan. In this paper, three new species, T. tum trapizeform, tridentate infront, pronotumstrong yingjiangensis sp. nov. , T. protumidus sp. nov. ly transverse, more or less denticulate, elytra rugous and T. arcuatus sp. nov. fro3m Yunnan are de and densely setiferous, abdomen cylindrical. scribed. One species, T. variolosus, is recorded from mainland China for the first time. Key to species of Thoracochirus from mainland China The following abbreviations are used in the text, 1. Laterallobeof headwithouttooth ………… T. variolosus Fauvel Laterallobeof headfurnishedfrontallywitha distincttooth ……2 HL: head length (from front of lateral lobe to hind 2. Frontal tooth auricular and broadly connected with lateral side of margin) ; PL: pronotum length; EL: elytron length; clypeus …………………………………… T. arcuatussp. nov. Thisstudy was supported by The State Key Basic Research and Development Plan of the Ministry of Science and Technology of China (G200046800) , NationalScience Fundfor Fostering TalentsinBasic Research (NFSCJ0030092) and KFBG (Kadoorie Farm &Botanic Garden) 2 BiodiversityStudentship of Hongkong. Correspondingauthor, Email: [email protected] Received13 May2005, accepted10June2005. 590 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 3 2 2 2 July, 2005 吴 捷等: 中国疣隐翅虫属 (鞘翅目,隐翅虫科,筒隐翅虫亚科) 分类研究 591 Frontaltoothtriangular, notconnectedwithlateralsideof clypeus setae and rugous punctures, along lateral and frontal …………………………………………………………………3 regionsparsely scattered with distinct verrucous denti 3. Median sulcus of head deep and wide, longer than the longitudinal cles. lengthof eyes; lateral region of pronotumscattered with distinct long verrucous denticles ………………… T. yingjiangensis sp. nov. Abdomen cylindrical, weakly broadened poste Mediansulcus of head indistinct, shorter than the longitudinal length riorly, segments except 7th and 8th densely covered of eyes; lateral region of pronotumscattered withindistinct verrucous with setae, each segment with an anterior and a poste ……………………… denticles T. protumidussp. nov. rior rows of setae, with the anterior row sparser and Thoracochirus yingjiangensis sp. nov. (Figs. 1 7) the posterior one denser on lateral side and vanished in the middle. Body length: 7 0 8 0 mm Male. Aedeagus (Fig. 2) with median lobeslight Measurement. HL: 0 51 mm; HW: 1 21 mm; ly bulbous at base, distinctly curved behind basal ori PL: 1 20 mm; PW: 1 99 mm; EL: 1 68 mm; EW: fice, posterior part slightly flat from lateral view, 1 95 mm. weaklysclerotized on the dorsal side and slightly con Body subconvex. Head black. Pronotum and ely tracted at apex; parameres short and rounded at apex, tra red brown. Abdomen black, with the terminal seg with the base protruding dorsally and connecting with ment reddish. Femora, tibiae and tarsi red yellow. each other. Ninth tergite (Fig. 5) sclerotized and left Head ( Fig. 1) transverse, frontal impression and right plates separated, each with 6setae of various broadly depressed, lateral lobe convex and furnished length. Tenth tergite almost membraneous except for frontally with a distinctly triangular tooth; clypeus sclerotized apical part, posterior margin truncate, each elongate, surface slightly crinkled, frontal part with side with 2 pairs of setae. Ninth sternite (Fig. 3) two convex tubercles; the median sulcus deep , longer composed of two plates, apical plate sclerotized except than the longitudinal diameter of eyes, vertex densely for membraneous apex, with two long setae and two punctate and setiferous, except for the smooth lateral pairs of short setae, basal plate less sclerotized, broad lobe, near posterior lateral margin scattered with some ened in middle. small verrucous denticles. Mandible protruding (Fig. 6). Mentum (Fig. 7) Female. Ninth sternite (Fig. 4) with hemister nite less sclerotized, second gonocoxite strongly sclero trapeziform and setaceous, with a curved groove, frontal margin medially tridentate, the median tooth tized except for membraneous apex, with onestout seta more protruding than lateral teeth and basally de and twoshort setae. pressed. Holotype ♂, Yunnan, Yingjiang (24 6°N, Antennae long, posteriorly reaching the middle of 97 9°E) , 1700m, 17 May 1980, SONGShi Mei leg. elytra, 1st segment extra elongate, about as long as Paratypes: 2 ♂♂, 1♀, same data as holotype. 2nd 5th segments together, 2nd longer than wide, 3rd Etymology. This specific epithet refers to the type baculiform, about 2 times as long as 2nd, 4th 6th locality“Yingjiang”. longer than wide, 7th quadrate, 8th 10th slightly Distribution. China (Yunnan). transverse, 11th coniform, bluntly pointed, about 2 Remarks. The new species is similar to T. verru times as long as 10th. Each segments with dense long cifer Fauvel from India and Myanmar, but can be ea setae. sily distinguished by distinctly smaller body size and Pronotum strongly transverse, frontal margin dis less convex lateral lobe. This species is also similar to tinctly bisinuate, lateralsides slightly convex outward, T. decanus Biswas from India, but differs from the median longitudinal sulcus obsolete in middle, anterior latter in having a more transverse pronotum and a less part more distinctly depressed, posterior end especially tridentate mentum. expanded as a“T”impression, disc densely covered Thoracochirus protumidus sp. nov. (Figs. 8 14) withsetae and umbilicatepunctures, close tolateral re gion evenly scattered with distinct long verrucous den Body length: 8 0 9 0 mm. ticles. In marginal area, lateral line continuous, Measurement. HL: 0 59 mm; HW: 1 21 mm; strongly bent upward at hind angle, and connecting PL: 1 17 mm; PW: 1 87 mm; EL: 1 64 mm; EW: with basal line of posterior constriction. 1 83 mm. Protibiae crenulate externally, with 12 14 denti Body subconvex. Mouthpart, clypeus and anten cles. nae reddish. Head, pronotum and elytra reddish Elytra slightly transverse, dorsal disc with dense black. Abdomen reddish black except for the yellowish 2 2 2 2 1 2 1 2 1 1 2 1 1 1 1 2 2 2 2 2 2 2 2 2 2 1 1 2 2 2 2 2 2 2 2 2 2 1 2 1 1 1 1 1 1 1 2 2 2 59 2 Acta Zootaxonomica Sinica动物分类学报 Vol. 30 No. 3 terminal segment. Femora and tibiae reddish, tarsi gate, surface smooth, frontal part with two small tu yellowish. bercles; the mediansulcusshorter than the longitudinal Head ( Fig. 8) transverse, frontal impression diameter of eyes, divergent forward and backward; broadly depressed, lateral lobe convex and frontally vertex densely punctate and setiferous except for the furnished with a small triangular tooth; clypeus elon lateral lobe smooth. Figs. 1 7. Thoracochirusyingjiangensissp. nov. 1. Headandpronotumin dorsal view. 2 Aedeagusin dorsal andlateral view. 3 9thsternite of male. 4 Left half of female 9thsternite. 5 Male 9th and 10th tergites. 6 Left mandible. 7 Mentum. Scale bars=0 5mm. Mandible protruding (Fig. 13). Mentum (Fig. necting with basal line of posterior constriction. 14) trapeziform and setaceous with a curved groove, Protibiae crenulate externally, with 12 or 13 den frontal margin medically tridentate, median tooth ticles. more protruding than lateral teeth and laterally de Elytra almost quadrate, dorsal disc with dense se pressed. tae and rugous punctures, lateral and frontal region Antennae long, posteriorly reaching the middle of sparsely scattered with indistinct verrucous denticles. elytra, 1st segment elongate, as long as 2nd 4th seg Abdomen cylindrical, weakly broadened poste ments together, 2nd longer than wide, 3rd baculi riorly, segments except 7th and 8th densely covered form, about 1 5 times as long as 2nd, 5th 7th longer with setae, each segment with an anterior and a poste than wide, 8th quadrate, 9th and 10th slightly trans rior rows of setae, with the anterior row sparser and verse, 11th coniform, bluntly pointed, about 2 times the posterior one denser on lateral side and vanished in as long as 10th Each segments with dense long setae. the middle. Pronotum transverse, frontal margin bisinuate, Male. Aedeagus (Fig. 9) with median lobeslight lateral sides parallel, median longitudinal sulcus nar ly bulbous at base, slightly curved behind basal ori row and evenly depressed, but the posterior end ex fice, posterior part slightlyflat from lateral view, very panded as inverse“Y”, disc densely covered with setae weakly sclerotized on the dorsal side, apical part and umbilicate punctures, near lateral region scattered slightly contracted, with a longitudinal carina; with indistinct verrucous denticles, but more distinct at parameres long and rounded at apex, with basal part the posterior angle. In marginal area, lateral line con protruding dorsally and connecting with each other to tinuous, slightly, bent upward at hind angle, and con form a triangular construction. Ninth tergite (Fig.2 2 2 1 1 1 1 1 1 1 2 2 2 2 2 2 2 1 2 2 2 1 2 2 2 2 2 2 July, 2005 吴 捷等: 中国疣隐翅虫属 (鞘翅目,隐翅虫科,筒隐翅虫亚科) 分类研究 593 12) sclerotized, left and right plates separated, each Yunnan, Xishuangbanna, Menglun, 650 m, 21 with 7 setae of various length. Tenth tergite almost Feb 2004, WU Jie and ZHANG JiaoLin leg. ; 1 ♀ membraneous except for sclerotized apical part, poste Yunnan, Xishuangbanna, Menglun, 690 m, 21 Feb. rior margin truncate, each side with 2 pairs of setae. 2004, WU Jie leg. ; 1 ♀, Yunnan, Xishuangbanna, Ninth sternite (Fig. 10) composed of two plates, api Mengla, 690m, 14 Feb. 2004, WU Jie leg. cal plate sclerotized with a membraneous median line, Etymology. Thespecies name is fromLatin adjec with two long setae and two pairs of short setae, basal tives protumidus to indicate the triangular protruding plate less sclerotized, broadened in middle. lateral teeth on head. Female. Ninth sternite (Fig. 11) with hemistern Distribution. China (Yunnan). ite less sclerotized, second gonocoxite strongly sclero Remarks. This species is similar to T. decanus tized except for slightly membraneous apex, with one from India, but could be distinguished by larger body stout seta and twoshort setae. size and less distinct verrucuous denticles of pronotum Holotype ♂, Yunnan, Xishuangbanna, Menglun and elytra. The mandible shape is also distinctly dif (21 9°N, 101 2°E) , 550 m, 8 Feb. 2004, WU Jie ferent. and BAI Dai Yuan leg. Paratypes: 4 ♂♂, 2 ♀♀, Figs. 8 14. Thoracochirusprotumidussp. nov. 8. Head andpronotumin dorsal view. 9. Aedeagusin dorsal andlateral view. 10. Male 9thsternite. 11. Left half of female 9th sternite. 12. Male 9th and 10th tergites. 13. Left mandible. 14. Mentum. Scale bars=0 5mm. Head (Fig. 15) transverse, frontal impression Thoracochirus arcuatus sp. nov. (Figs. 15 21) broadly depressed, lateral lobe moderately convex and Body length: 5 0 5 5 mm. furnished with an auricular tooth, basally depressed Measurement. HL: 0 37 mm; HW: 0 78 mm; and connected with lateral margin of clypeus; clypeus PL: 0 79 mm; PW: 1 23 mm; EL: 1 20 mm; EW: elongate, surface finely punctured, with two distinctly 1 32 mm. convex denticles, front part of clypeus evidently de Body subconvex. Head, pronotum and elytra pressed; the mediansulcus indistinct, weakly depressed black. Abdomen black, with the terminal segment red and as long as the longitudinal length of eyes; vertex brown. Antennae brown. Femora and tibiae red densely punctate and setiferous, except for the lateral brown, tarsi yellowish. lobe smooth. 1 2 2 2 2 2 2 2 1 1 2 2 1 2 1 2 1 1 1 1 1 1 1 2 2 2 59 4 Acta Zootaxonomica Sinica动物分类学报 Vol. 30 No. 3 Mandible protruding (Fig. 20). Mentum (Fig. Abdomen cylindrical, weakly broadened poste 21) trapeziform and setaceous with a curved groove, riorly, segments except 7th and 8th densely covered frontal margin tridentate, median tooth and lateral with setae, each segment with an anterior and a poste teeth equally protruding. rior rows of setae, both denser on lateral side and van Antennae long, posteriorly reaching the middle of ished in the middle. elytra, 1st segment elongate, almost as long as 2nd 4th Male. Aedeagus ( Fig. 16) with median lobe segments together, 2nd longer than wide, 3rd baculi slightly bulbous at base, slightly curved behind basal form, about 1 5 times as long as 2nd, 4th and 5th orifice, posterior part slightly flat from lateral view, longer than wide, 6th 10th transverse, 11th coniform, weakly sclerotized on the dorsal side, apical part bluntly pointed, about 3 times as long as 10th. Each slightly contracted, with a longitudinal carina; segments with dense long setae. parameres long and rounded at apex, with basal part Pronotum transverse, frontal margin slightly bi protruding dorsally and connecting with each other to sinuate, lateral sides parallel, median longitudinal sul form a triangular construction. Ninth tergite (Fig. cus narrow, more distinctly depressed infront, and ex 19) sclerotized, left and right plates separated, each panded at posterior end to form an inverse“Y”, disc with 6 setae of various length. Tenth tergite almost densely covered with setae and umbilicate punctures, membraneous except for sclerotized apical part, poste close to lateral region scattered with indistinct verru rior margin truncate, each side with 2 pairs of setae. cous denticles, and even more distinct at the posterior Ninth sternite (Fig. 17) composed of two plates, api angle. In marginal area, lateral line continuous, cal plate sclerotized except for membraneous apex, slightly bent upward at hind angle, and connecting with two long setae and two pairs of short setae, basal with basal line of posterior constriction. plate less sclerotized, broadened basally. Protibiae crenulate externally, with 11 14 denti Female. Ninth sternite (Fig. 18) with hemistern cles. ite less sclerotized, anterior margin not very round, Elytra almost quadrate, disc with dense setae and second gonocoxite strongly sclerotized except for mem rugous punctures, along lateral and frontal region braneous apex, with onestout seta and twoshort setae. sparsely scattered with indistinct verrucous denticles. Figs. 15 21. Thoracochirusarcuatussp. nov. 15. Headandpronotumin dorsal view. 16. Aedeagusin dorsalandlateral view. 17. Male 9thsternite. 18. Left half of female 9thsternite. 19. Male9th and10th tergites. 20. Left mandible. 21. Mentum. Scale bars=0 5mm. 2 2 2 2 2 1 2 2 2 2 2 2 2 2 2 2 2 2 1 July, 2005 吴 捷等: 中国疣隐翅虫属 (鞘翅目,隐翅虫科,筒隐翅虫亚科) 分类研究 595 Holotype ♂, Yunnan, Xishuangbanna, Menglun form, about 1 5 times as long as 2nd, 4th 9th longer (21 9°N, 101 2°E) , 860 m, 11 Feb. 2004, WU Jie than wide, 10th quadrate, 11th coniform, bluntly leg. Paratypes: 5 ♂♂, 1 ♀ Yunnan, Xishuangban pointed, about 2 times as long as 10th. Each segments na, Menglun, 855 m, 8 Feb. 2004, WU Jie and BAI with dense long setae. Dai Yuan leg. Pronotum transverse, frontal margin distinct bis Etymology. Thespecies name is fromLatin adjec inuate, lateral sides parallel, median longitudinal sul tives arcuatus to indicate the auricular lateral teeth on cus narrow, more distinctly depressed in front and ex head. panded at posterior end to form an inverse“Y”, disc Distribution: China (Yunnan). densely covered with setae and umbilicate punctures, Remarks. The lateral teeth of head in this species close to lateral region scattered with long verrucous basally connected with lateral mangin of clypeus are denticles. In marginal area, lateral line continuous, distinctly different from other species in this genus. slightly bent upward at hind angle, and connecting with basal line of posterior constriction. Thoracochirus variolosus Fauvel, 1895 New Protibiae crenulate externally, with 13 15 denti record for mainland China (Figs. 22 28) cles. Thoracochirus variolosus Fauvel, 1895: 183; Heller, 1898: 8; Bern hauer, 1903: 156; Bernhauer and Schubert, 1910: 19; Bernhauer, Elytra almost quadrate, dorsal disc with dense se 1922: 221; Bernhauer, 1926: 255; Wendeler, 1928: 119; tae and rugous punctures, along lateral and frontal re Cameron, 1930: 119; Scheerpeltz, 1933: 1009; Herman, 2001: 1 gion sparsely scattered with distinct verrucous denti 144. cles. Synoym: Thoracochirus forsteri Bernhauer, 1903: 156; Bernhauer and Schubert, 1910: 18; Bernhauer, 1926: 255; Cameron, 1928: 95; Abdomen cylindrical, weakly broadened posteri Scheerpeltz, 1933: 1009. orly, segments except 7th and 8th densely covered with Thoracochirus intermedius Bernhauer, 1926: 255; Wendeler, 1928: setae, each segment with an anterior and a posterior 119; Scheerpeltz, 1933: 1009; Herman, 2001: 1144. rows of setae, with the anterior row sparser and the Thoracochirus sublaevicollis Bernhauer, 1926: 255; Wendeler, 1928: posterior one denser on lateral side and vanished in the 119; Scheerpeltz, 1933: 1009; Herman, 2001: 1144. middle. Body length: 6 0 7 0 mm. Male. Aedeagus ( Fig. 23) with median lobe Measurement. HL: 0 47 mm; HW: 0 86 mm; slightly bulbous at base, curved behind basal orifice, PL: 0 90 mm; PW: 1 44 mm; EL: 1 40 mm; EW: posterior part baculiform, weakly sclerotized on the 1 52 mm. dorsal side and rounded at apex; parameres short and Body subconvex. Head, pronotum and elytra pointed, with basal part protruding dorsally and con black. Abdomen black, with the terminal segment necting with each other. Ninth tergite (Fig. 26) scle dark red. Antennae dark red. Femora and tibiae dark rotized and left and right plates separated, each with 7 red, tarsi yellowish. or 8setae of various length. Tenth tergite almost mem Head (Fig. 22) transverse, frontal impression braneous except for sclerotized apical part and posteri broadly depressed, lateral lobe moderately convex, not or margin truncate, each side with 2 pairs of setae. furnished with a tooth, just weakly protruding along Ninth sternite (Fig. 24) composed of two plates, api frontal margin; clypeus elongate, surface crinkled, cal plate sclerotized except for membraneous apex, frontally rounded; the median sulcus indistinct, shor with two long setae and two pairs of short setae, basal ter than the longitudinal length of eyes; vertex densely plate less sclerotized, broadened in the middle. punctate and setiferous except for the lateral lobe Female. Ninth sternite (Fig. 25) with hemistern smooth. ite less sclerotized, second gonocoxite strongly sclero Mandible protruding (Fig. 27) Mentum (Fig. tized except for membraneous apex, with onestout seta 28) trapeziform and setaceous with a curved groove, and twoshort setae. frontal margin tridentate, median toothprotruding and Materials examined. 3 ♂♂, 3 ♀♀ Yunnan, longer than lateral tooth. Xishuangbanna, Menglun, 810 m, 21 Feb. 2004, Antennae long, posteriorly reaching the middle of WU Jie and ZHANGJiaoLin leg. elytra, 1st segment elongate, almost as long as 2nd 4th Distribution. China (Yunnan, Taiwan) ; Myan segments together, 2nd longer than wide, 3rd baculi mar, Indonesia, Borneo, Philippines. 1 2 1 1 2 2 2 2 2 2 2 2 2 2 2 2 2 2 1 2 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 59 6 Acta Zootaxonomica Sinica动物分类学报 Vol. 30 No. 3 Figs. 22 28. Thoracochirus variolosus. 22. Headandpronotumin dorsal view. 23. Aedeagusin dorsal andlateral view. 24. Male 9thsternite. 25. Left half of female 9th sternite. 26. Male 9th and 10th tergites. 27. Left mandible. 28. Mentum. Scale bars=0 5mm. Acknowledgement We are grateful to Mr. YANG mologischeMitteilungen, 17 (2) : 90110. Song Hai and other colleagues of Xishuangbanna Na Cameron, M. 1930. The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 1. Taylor and Francis, ture Reserve, Yunnan Province, for their kindly assis London. pp. 89120. tance during our field investigations. Cameron, M. 1940. Newspeciesof OrientalStaphylinidae (Col.). The Entomologist’s MonthlyMagazine, 76: 249253. REFERENCES Fauvel,A. 1895. Staphylinidae nouveaux de l’Inde et de la Malaisie. Bernhauer, M. 1903. Die Staphyliniden Tribus Leptochirina nebst ana Revued’Entomologie, 14: 180286. lytischenBestimmungstabellender Gattungen. Deutsche Entomologis Hammond, P. M. 1984. An annotated checklist of Staphylinidae (In che Zeitschrift, 1903: 113160. secta: Coleoptera) recorded from Borneo. The Sarawak Museum Bernhauer, M. 1922. Sauter’s FormosaAusbeute: Staphylinidea. Journal, 33 (54) : 187218. Archivfür Naturgeschichte, (A) 88 (7) : 220237. Heller, K. M. 1898. Nr. 3. neue KfervonCelebes Ⅲ. Abhandlungen Bernhauer, M. 1926. Neue Staphyliniden aus Ostindien. Wiener Ento und Berichte K nigl. Zoologischen und Anthropologisch Ethno mologische Zeitung, 43 (1) : 1925. graphischen MuseumszuDresden, 7 (3) : 120. Bernhauer, M. and Schubert, K. 1910. Staphylinidae I. In: Schen Herman,L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleop kling, S. (ed.) , ColeopterorumCatalogus. 5 (19). Junk, Berlin. tera). 1758 to the end of the second millennium. Ⅲ. Oxyteline pp. 186. group. Bulletinof theAmericanMuseumof Natural History,265:1 Biswas, D. N. and Gupta, T. Sen 1982. Newspecies and new records 1081142. of Staphylinidae (Coleoptera) fromIndiaandSriLanka. RevueSu Scheerpeltz, O. 1933. Staphylinidae Ⅶ. In: Schenkling, S. (ed.) , isse Zoologie, 89 (1) : 135154. ColeopterorumCatalogus. 6 (129). Junk,Berlin. pp. 9891500. Blackwelder, R. E. 1952. The generic names of the beetle family Shibata, Y. 1973. Preliminarychecklist of thefamily Staphylinidae of Staphylinidae, with an essay on genotypy. United States National Taiwan (Insecta: Coleoptera). Annual Bulletin of the Nichidai MuseumBulletin, 200:iiv, 1483. Sanko, 16: 2188. Cameron, M. 1925. Catalogue of the Indian Insects. Part 6Staphylin Wendeler, H. 1928. Subtribus Leptochiri der Philippinen (Coleoptera, idae. Calcutta: Government of India. 126pp. Staphylinidae). Deutsche Entomologische Zeitschrift, 1928: 117 Cameron, M. 1928. Fauna sumatrensis. Staphylinidae (Col.). Ento 128. 2 2 2 1 2 2 2 2 2 2 2 2 2 2 2 (cid:190) 2 (cid:202) 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 July, 2005 吴 捷等: 中国疣隐翅虫属 (鞘翅目,隐翅虫科,筒隐翅虫亚科) 分类研究 597 中国疣隐翅虫属 (鞘翅目, 隐翅虫科, 筒隐翅虫亚科) 分类研究 吴 捷1,2, 周红章1 1. 中国科学院动物研究所 北京 100080 2. 中国科学院研究生院 北京 100039 摘 要 疣隐翅虫属 Thoracochirus种类多见于腐木树皮下, 突疣隐翅虫 T. protumidussp. nov. , 耳突疣隐翅虫 T. arcu 是重要的腐木甲虫。在本研究之前, 该属只有两个种在中国 atussp. nov. 及长疣隐翅虫 T. variolosus大陆新纪录。文中 台湾有记录: 长疣隐翅虫 Thoracochirus variolosus Fauvel, 提供了该4个种的检索表。所有标本 (包括所有模式标本) 1895和台湾疣隐翅虫 T. formosae Cameron, 1940. 记述了云 保存在中国科学院动物研究所。 南的3新种: 盈江疣隐翅虫 T. yingjiangensis sp. nov. , 尖 关键词 鞘翅目, 隐翅虫科, 疣隐翅虫属, 新种. 中图分类号 Q969 484 4 3 通讯作者. 2 1 1 3