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植物研究雑誌 J. Jpn. Bot. Originals 75: 127-136 (2000) Taxooomic Notes Ophiopogon of South Asia VI 00 Noriyuki TANAKA 。 Deptmentof Education,Sc hool of Liberal Arts,Te ikyo University, 訂 tsuka359,Hac hioji,To kyo,19 2-0395 JAPAN (Received on October 12,1 998) Ophiopogon chingii F.T.Wang & Ts.Tang,O. hainanensis Masam.,O. platyphyllus Merr. & Chun and O. latifolius L.Rodr. are taxonomically reinvestigated. Ophiopogon chingii var. glaucifolius F.T.Wang & L.K.Dai is not clearly demarcated from v chingii. 紅. This species is exceedingly variable,o ccurring in China and Vietnam. Some specimens from Vietnam formerly recorded by Rodriguez (1934) as O. reptans Hook.f. eidenti- 紅 fied here as O. chingii. Ophiopogon reptans recorded from Hainan Island (China) is also highly likely to be O. chingii. Ophiopogon hainanensis is regarded as conspecific with O. chingii. Ophiopogon chingii,O. plaηphyllus and O. latifolius differ at least in the apical form of the ovy. (Continued from 1. Jpn. Bot 75: 69-79) 訂 Key words: Ophiopogon chingii,O phiopogon hainanensis,O phiopogon latifolius, Ophiopogon platyphyllus,t axonomy (11) Ophiopogon chingii and O.・hai- E,P ),2 9181 (Tonkin,E ,P ),C lements 4308 nanenSIS (Annam,K ),P oilane 7029 (Annam,P ) and Ophiopogon chingii was described by Balansa 4155 (fig. 1),4 155-bis (both from Wang and Tang (1937) from Guangxi, Tonkin,K ) are assigned to var. chingii,wh ile China. Later,v glaucifolius was distin- specimens such as Tsang 22479 (type local- 紅 . guished under this species (Dai and Chen ity of var. glaucifolius in Guangxi,P; Fig. 2), 1978). Currently,v ar. glaucifolius is re- 27262 (Tonkin,P) ,2 7338 (Tonkin,E ,P) and corded from Guangdong, Guangxi and 30407 (Tonkin,C ,P ) are assigned to var. Sichuan (Dai and Chen 1978,Z eng 1991), glaucifolius. However,th e gap between the and var. chingii from Hainan,G uangdong, two varieties is rather indistinct. Guangxi, Yunnan, Sichuan and Guizhou Dai and Chen (1978) described var. (Dai and Chen 1978,Z eng 1991). The pre- chingii as having roots always more or less sent survey confirmed that this species,in - ligneous aIld somewhat stiff,l eaves 2.5- cluding the form orresponding to var. 8m mw ide,pe rianth lobes about 5m ml ong glaucifolius,o ccurs not only in China but and pedicels 6-9 mml ong. According to also in Vietnam. them,v ar. glaucifolius is distinguishable The typical forms of the two varieties of from v chingii by having ligneous stiff 訂. Ophiopogon chingii look. quite distinct from roots 3-4 mmi n diameter,g enerally wider each other (Figs. 1,2 ),a nd many of the leaves 0.5-2 cm wide,a nd pedicels nearly specimens available are actually assignable twice longer than perianths. However,m ost to either variety: e.g. specimens such as of these diagnostic characters overlap in the Tsang 22431 (Guangxi,P ),2 7171 (Tonkin, range of variation between the two varieties. -127- 128 植物研究雑誌第75巻第3号 平成12年6月 Fig. 1. Af orm of Ophiopogon chingii from northern Vietnam (B.Balansa 4155,K) . This form corresponds to var. chingii. Actually,in the specimens at hand,an y of variation; viz. the roots vary continuously these characters is rather continuous in from 0.8 to 3.5 mmi n diameter,st iffness of June 2000 Journal of Japanese Botany Vo .I75 No. 3 129 Fig.2. A form of Ophiopogon chingii from southem China (type locality of var. glaucifolius in Guangxi,W. T.Tsang 22479,p) . This form corresponds to var. glaucifolius. the root increases as its thickness increases, mmin width,a nd the proportion in length of the leaves vycontinuously from 2.5 to 21 the pedicel to the tepal (perianth lobe) (ab- 紅 130 植物研究雑誌第75巻第3号 平成12年6月 breviated here as p,汀)varies from 0.8 to 2.3. In Flora Hainanica (1977; abbreviated The leaves of the specimens assignable to hereafter as ‘FH'), Ophiopogon reptans var. chingii tend to be arranged consecu- Hook.f. is recorded. This species is recorded tively along the stem (Fig. 1),wh ile those of from eastem India and Myanm (Tanaka 紅 the specimens assignable to v glaucifolius 1999). In general appe anceO. repωns 訂. 紅 tend to be fasciculate on the apical part of closely resembles af orm of O. chingii (var. the stem (Fig. 2). However,t here are some chingii). Ophiopogon chingii is not recorded specimens showing the consecutive leaf ar- in FH,bu t its occuenceon Hainan Island is 町 rangement,de spite the fact that they 'econ- recorded in Iconographia Cormophytorum 訂 sistent with var. glaucifolius in other Sinicorum,v ol. 5 (1976),a nd by Dai and respects; e.g. in Morse 184 (2 sheets in fruit Chen (1978,as var. chingii). In the descrip- from Guangxi,K ),l eaves are aangedcon- tion of O. reptans in FH,t here esome 町 紅 secutively along the stem,to 9.7 mmw ide, points inconsistent with O. reptans of prop roots stiff and thick,to 3.5 mmi n di- Hooker. For instance,i n FH it is described ameter,pe rsistent (but withering) tepals 5.6 that the leaves of O. reptans are somewhat -5.8 mml ong,a nd pedicels to 13 mml ong obtuse at the apex,t he bracts eusually 訂 (P,汀=ca. 2.2-2.3). Further,th ere訂ealso shorter than the pedicels at anthesis,a nd the some specimens showing the intermediate pedicels e5-8 mml ong. These features 訂 leaf aangementbetween the two (e.g., agree with O. chingii [cf. also Anonymous 町 Tsang 27171,E ,P) . Therefore,e ven by the (1976) and Dai and Chen (1978) for O. leaf rangementpattem we cannot clearly chingiiJ rather than with O. repωns of 紅 demarcate the two varieties. From all these Hooker,s ince in the latter species the leaves observations,t he two varieties seem not to are acute,th e bracts are usually longer than be clearly delimited. the pedicels,a nd the pedicels are 1.8-3.5 Ophiopogon chingii var. chingii is re- mml ong at anthesis (to 5.5 mml ong in the ported to occur on Hainan Island (southem fruiting stage; cf. Tanaka 1999). Therefore,it China),( Dai and Chen 1978). Meanwhile, seems highly likely that O. reptans in FH is Masamune (1939) described O. hainanensis O. chingii. from this island. Masamune' sd escription of Rodriguez (1934) recorded Ophiopogon O. hainanensis agrees quite well with af orm reptans from Vietnam. Ie xamined some of O. chingii (corresponding to var. chingii). specimens [Balansa 4155 (Fig. 1),4155-bis, In Flora Hainanica (1977),h owever, O. both P; Poilane 7029,pJ formerly identified chingii is not recorded,a nd O. hαinanensis is as O. reptans by him. These eno doubt a 紅 treated as as ynonym of O. plaかphyllus(for form of O. chingii (v chingii). 1h ave 訂. this species see chapter 12). The following never seen as pecimen of true O. reptαns (of , passage in the protologue of O. hainanensis, Hooker) from Vietnam. “.. vaginis saepe cristatis; ..antherae 2 mm As mentioned earlier,O phiopogon reptans 10時ae",seems to conform to O. chingii [see and af orm of O. chingii (var. chingii) resem- also Dai and Chen (1978) for the description ble each other,bu t actually these two species of O. chingiiJ rather than to O. plaηphyllus, differ in several respects. One notable since in the latter species the leaf vagina is difference is that in O. reptans the leaves are not cristate (or not rugulose) and the anthers tufted at intervals on the stem (Tanaka are more than ca.3.3 mml ong. As far as the 1999),w hile in the form of O. chingii (v 低 original description is surveyed,it is highly chingii) the leaves tend to be arranged con- likely that O. hainanensis is conspecific with secutively along the stem (Fig. 1). O. chingii. Ophiopogon stenophyllus (Me町.)L.Rodr. June 2000 Journal of Japanese Botany Vol. 75 No. 3 131 described from Guangdong bears some re- less declinate,t o ca.23 cm long. Inf1ore- semblance with O. chingii,bu t the anthers of scence racemose. Pedicels (incl. basal stalky the former are coherent (e.g. Hance 10086, p tofperianth) 3.5-9.5 mml ong (to 13 mm 紅 K) (Anonymous 1976,D ai and Chen 1978), long in fruit),jo inted usually around middle while those of the latter are separate. (articulation site variable). Bracts ovate to The ov紅Y of Ophiopogon chingii is lanceolate, acute, acuminate or caudate, slightly inf1ated at the apex as illustrated in scarious, entire, usually shorter than Fig. 6A,b ut the degree of the inf1ation ap- pedicels,to ca.9 mml ong (rarely to 16 mm pears to be slightly weaker than in O. pierrei long). Flowers secund,c emuous,1 -3 in axils L.Rodr.; in the latter the same part is nearly of bracts. Perianth lobes 6,o vate-oblong,4. dome-shaped (Tanaka 2000). 3-5.8 mml ong,1. 5-2.1 mmw ide. Stamens 6. Anthers lanceolate,2 .3-3.5 mm long. Ophiopogon chingii F. T. Wang & Filaments very short,f ree. Pistil 1. Style Ts.Tang in Bull. Fan Mem. Inst. Biol. 7: 282 subulate,3 -5 mml ong. Ovary slightly con- (1937); Anonym.,I con. Corm. Sin. 5: 526, vex at apex. Seeds globul toellipsoidal. 訂 f.7881 (1976); L.K.Dai and S.C.Chen [see Distribution: Vietnam and SWC hina. Chen et al. (1993), for authorship] in Representative specimens examihed: F.T.Wang and Ts.Tang,F l. Reipub. Pop. Sin. Vietnam. Annam,B ana,p res Tourane,J ul. 10, 15: 146, t.48, f.1,2 (1978); W.Z.Zeng in 1923,fl. ,Po ilane 7029 (P); Tourane,M ay-Jul. 1927,J. & M.S.Clements 4308 (K); Tonkin,T u-phap,J un. J.M.Xu,F l. Sichuan. 7: 308,t .l03,f .1-3 1887,fl .,B .Balansa 4155 (K); ibid.,J an. 1887,f ,. r (1991). [Figs. 1,2 ,6 A] B.Balansa 4155-bis (K); Tonkin,Ta ai Wong MoS han O. reptans auct. non Hook.f.: L.Rodr. in and vicinity,Ch uk-phai,Ha coi,No v. 18-Dec. 2,1 936, Lecomte,F l. Indo-Chine 6: 660 (1934), fr.,W. T.Tsang 27262 (E,P) ; ibid.,No v. 1-9,1 936,fr. , saltem quoad specim. Poilane & Balansa; W.T.Tsang 27171 (E,P) ; ibid.,日.white,fr agrant,M ay 3-Jun. 22,19 39,fl .,W .T.Tsang 29181 (E,P ); Taai Anonym.,F l. Hainan. 4: 110 (1977). Wong MoS han and vicinity,S hui Mei village,N Eo f O. hαinanensis Masam. in Trans. Nat. Chuk-phai,Ha coi,fl. white,fr agrant,Ju n. 23-Aug. 31, Hist. Soc. Formosa 29: 28 (1939); Fl. Kai- 1939,日., W.T.Tsang 29409 (E,P ); Sai Wong Mo nan. 404 (1943); syn. nov. Shan,L omg Ngong village,D am-ha,J ul. 18-Sept. 9, O. chingii F.T.Wang & Ts.Tang v 1940,(l ate)日.,W.T.Tsang 30407 (C,K ,L ,P) ; Ouonbi, 紅. glaucifolius F.T.Wang & L.K.Dai in L.K.Dai Nov. 1885,fr .,B .Balahsa 274 (K); ~au Nga Shan and vicinity,T ien-yen,D ec. 13-22,1 936,f ,.r W.T.Tsang and S.C.Chen in F.T.Wang and Ts.Tang,F l. 27338 (E,P) . Reipub. Pop. Sin. 15: 146 (sine descr. China. Guangxi (Kwangsi),S hap Man Taai Shan, Latinよt.49,f. 1ムp.252(1978); W.Z.Zeng near Hoh Lung village,S E of Shang-sze,K wangtung in J.M.Xu,F l. Sichuan. 7: 309 (1991); syn. border (Shan'g-sze Distrょfl.odorless,w hite,Ju n. 17, nov. 1933,fl. ,W .T.Tsang 22479 (P); ibid.,A pr. 18,1 933, fr.,W .T.Tsang 22135 (P); Shap Man Taai Shan,nea r Glabrous perennial herb. Stem decumbent Iu Shan village,S E of Shang-sze,fl. white,M ay 19, to semi-erect,to more than 43 cm long,to 1933,fl. (bud),W .T.Tsang 22330 (K); Shap Man Taai ca.6 mmin diameter,oc casionally branched, Shan,n ear Iu Shan village,S E of Shang-sze,fl. light emitting prop roots to ca.3.5 mmin diameter. blue,J un. 2-7, 1933,日.W.T.Tsang 22431 (P); Leaves fasciculate on apex of stem or bome Guangxi,L ungchow,s outh hill,F eb.,fr .,H .B all along stem,ac ute or obtuse at apex,ta per- 1凶84(K); Guangxi,L aohutiao,N apo Pref.,c a.1000 m, Oct. 20,19 97,H . Akiyama et al. 1652 (MAK); ing to base which is vaginate with-broad, Guangxi,P ingmen,N apo Pref.,8 00-1000 m,O ct. 17, scarious and often transversely rugulose 1997,fr. ,H. Akiyama et al. 135 (MAK). wings,g laucous on abaxial surface,5 -21ve Ohiopogon stenophyllus quoted in the text: ・ ined (veins often indistinct),7 -28 cm long, China. Guangdong (Prov. Cantonensis), West' 2.5-21 mmw ide. Scape complanate,mo re or River,1 872,fl. ,H ance 10086 (K). 132 植物研究雑誌第75巻第3号 平成12年6月 (12) Ophiopogon pla砂']Jhyllus by Merrill and Chun (1935) from Hainan' Ophiopogon plaか'Phylluswas described Island,s outhern China. In general appear- Fig.3. Ophiopogon platyphyllus from Hainan Island (S China,F. C.How 70701,K) . The collector and collection number of this specimen,F. C.How 70701,co incides with that of the type of this species,bu t the exact locality within the island differs between the type and this specimen. June 2000 Journal of Japanese Botany Vol. 75 No. 3 133 ance this species (Fig. 3) closely resembles prop roots to 3 mm in diameter. Leaves the wide-leaf form of O. chingii (correspond- tufted on apical p tof stem,l inear or 訂 ing to var. glaucifolius) (e.g.,T sang 27262, broadly linear,a cute or obtuse at apex,a t- P). But,t hey differ at least in the following tenuate to base which is vaginate with floral character; in O. platyphyllus the cen- sciouswings,st riate on abaxial surface,to 紅 tral part of the apex of the ovary is concave 19veined,to 30 cm long,to 15.5 mmw ide. ・ as illustrated in Fig. 6B,w hile in O. chingii Scape complanate,d eclinate,t o 11.5 cm the same part is slightly inflated (.or convex) long. Inflorescence racemose. Pedicels (incl. above (Fig. 6A). On the other hand,th e n basal stalky part of perianth) to 9.5 mml ong 紅- row-leaf form of O. plaかphyllus(McClure (true pedicels excl. perianth part to 3.5 mm 9334,K ) much resembles O. stenophyllus long),a rticulate in lower p紅t.Bracts entire, (Me町.)L.Rodr. described from Guangdong, sciousat margin. Flowers secund,c emu- 紅 China. But,t hese two species differ also in ous,1 -3 in axils of bracts. Perianth lobes 6, the apical form of the ovary; viz. in O. lanceolate,ca .5.3-6.5 mml ong,1 .6-2.2 mm stenophyllus the same p tis inflated above wide. Stamens 6. Anthers lanceolate,3 .3-4 訂 (e.g.,H ance 10086,K ; Merrill 10757,L ) mml ong. Pistil 1. Style subulate,5 .2-5.5 somewhat like that in O. chingii (Fig. 6A), mml ong. Ovary inferior,c oncave at apex. while in O. pla砂phyllusit is concave (Fig. Distribution: China (Hainan and Guang- 6B),as stated above. Further,th e anthers of dong). O. stenophyllus are coherent laterally (Hance Specimens examined: 10086,K ),wh ile those of O. platyphyllus are China. Hainan,Y aichow,1 800 ft,fl. white,an thers pale green,fr uits blue,M ar.-Jul. 1933,日., F.C.How free. Ophiopogon platyphyllus also resem- 70701 (K); Hainan,Fi ve Finger M ,.tfls. light blue,la v- bles O. caulescens. But in O. plaかphyllus ender and white,f rs.,b lue,A pr. 23, 1922, 乱f.,l the filaments are separate, while in O. F.A caulescens they訂emore or less united later- Ophiωopogon stenophyllus quot胞edin the text仁: ally (Tanaka 2000). China. Guangdong,W est River,1 872,fl .,H ance 10086 (K); Guangdong,L oh Fau Mountain (Lofau- shan),M errill 10757 (fragment of type specimen of Ophiopogon platyphyllus Me & Chun 町. Peliosanthes stenophylla Merr.,L) . in J. Bot. Inst. Col. Agr. Sun Yatsen Univ. 2: 211,t. 38 (1935); Anonym.,F l. Hainan. 4: (13) Ophiopogon latifolius 110 (1977),p. p.,e xcl. syn. O. hainanensis; Ophiopogon latifolius was described by L.K.Dai and S.C.Chen [cf. Chen et al. Rodriguez (1928) from northem Vietnam. In (1993) for authorshipJ in F.T.Wang and general appearance this species (Figs. 4,5) Ts.Tang,F l. Reipub. Pop. Sin. 15: 146,t. 49, resembles O. chingii (cf. chapter 11),O . f.5-7 (1978),e xcl. syn. O. hainanensis. platyphyllus (cf. chapter 12) and O. [Figs. 3,6 BJ caulescens (Tanaka 2000). But,O. latifolius Peliosαnthes stenophyllus auct. non民.fe汀. is distinguishable from them by the relatively [in Philip. 1. Sci. Boi. 13: 134 (1918)J: long inflorescence with many flowers,t he Me in Lingnan Sci. 1. 5: 47 (1927). 町. pedicels articulate near the apex,a nd the Ophiopogon stenophyllus auct. non ovary which is rather flat at the apex as illus- (Me町.)L.Rodr. [iri Bull. Mus. Hist. Nat. trated in Fig. 6C. In O. chingii the apical part Paris ser. 2,6 : 95 (1934)J: L.Rodr. in of the ovyis slightly inflated above (Fig. 紅 Lecomte,F l. Indo-Chine 6: 663 (1934), 6A),w hile in O. pla砂phyllus(Fig. 6B) and quoad specim. McClure 9334. O. caulescens (Tanaka 2000-Figs. 6A,6 B) Glabrous perennial herb. Stem to more the same p tis concave. The filaments of O. 訂 than 9c m,u p to 8m min diameter,em itting 134 植物研究雑誌第75巻第3号 平成12年6月 Fig.4. Type specimen of Ophiopogon latifolius (N Vietnam,B. Balansa 2156,p) . latifolius are separate,wh ile in O. caulescens scribed from Guangxi,Ch ina,al so somewhat they are more or less united laterally (Tanaka resembles O. 1αtifolius,bu t the anthers of the , 2000). Ophiopogon multiflorus Y.Wan de- former are coherent (Wan 1988),wh ile those June 2000 Journal of Japanese Botany Vol. 75 No. 3 135 of the latter are free. Ophiopogon latifolius L.Rodr. in Bull. Soc. Bot. Fr. 75: 998 (1928); in Lecomte,F l. Indo-Chine 6: 660 (1934). [Figs. 4,5 ,6 C] Glabrous perennial herb. Root stiff,2 .3 mmi n diameter. Leaves many,t ufted on apical ptof stem broadly linear,a cute,at - 訂 tenuate to base which is vaginate with sc紅白uswings,to 58.5 cm long,to ca.1.5c m wide,ve ins ca.22 incl. rather indistinct ones, midrib prominent beneath. Scape ca.32 cm long,c omplanate. Inflorescence racemose, 16.5 cm long. Flowers secund,m any,2 -6 in axils of bracts. Pedicels (incl. basal stalky p ofperianth) 7.5-10 mm long (true 制 pedicels excl. perianth p t6.7-9 mml ong), 紅 thin,j ointed near apex. Bracts lanceolate, acute to acuminate,sc iousat m gin,to 1.6 紅 紅 cm long. Perianth lobes 6,o vate-lanceolate, 4-5 mml ong,2 -2.4 mmw ide. Stamens 6, free. Anthers ovate-lanceolate,2 .7-2.8 mm long,1 .4-1.5 mmw ide at base. Filaments very short,0 .3-0.4 mml ong. Pistil 1. Style subulate,4 .5-5 mml ong. Ovary inferior, rather flat at apex. Distribution: N Vietnam. Also recorded from SWC hina (SE Yunnan) by Dai and Fig. 5. Close-up of a part of the inflorescence of Chen (1978). Ophiopogon latifolius (N Vietnam,B. Balansa Specimen examined: 2156,ty pe,p) .Pe dicels are制 iculatenear the Vietnam. Tonkin. Valee de Lankok,M ont. Bavi, apex (indicated by the arrow) . Oct. 1887,fl .,B .Balansa 2156 (type,P ). A B C Fig. 6. Diagrammatic sketches of the floral structure of three species of 附.c. Ophiopogon (lateral view). A. O. chingii. B. O. plαat砂y帥yll O. latifolius. 136 植物研究雑誌第75巻第3号 平成12年6月 1e xpress my sincere thanks to the curators Chen S. c., Li J. L., Zhu X. Y. and Zhang Z. Y. 1993. of the following herbaria for the loan of Bibliography of Chinese Systematic Botany specimens; C,E ,K ,L and P. Warm thanks (1949-1990). Guangdong Science & Technology Press,G uangzhou. ealso extended to Professor Jin Murata of 紅 Dai L. K. and Chen S. C. 1978. Ophiopogon latifolius, Tokyo Metropolitan University (now at the p. 149. O. stenophyllus,p. 150. In: Wang F. T. and University of Tokyo) for access to the speci- Tang Ts. (edsよFloraReipublicae Popularis mens (MAK) collected by the team of Sino- Sinicae 15. Science Press,B eijing. Japanese botanical expedition to Yunnan and Tanaka N. 1999. Taxonomic notes on Ophiopogon of South Asia 11. J. Jpn. Bot. 74: 25-33. Guangxi in 1997. 一一一2000.Taxonomic notes on Ophiopogon of South Asia V. J. Jpn. Bo .t75: 69-79. References Wan Y. 1988. A new species of the genus Ophiopogon Anonymous 1976. Ophiopogon stenophyllus. In: from Guangxi. Guihaia 8( 3): 235-236. Iconographia Cormophytorum Sinicorum 5: 526. See also those cited in the text. Science Press,B e討ing. 田中教之:南アジア産ジャノヒゲ属の分類学的検 討VI ジャノヒゲ属の 4種 (Ophiopogonchingii, O. これもその記載(海南植物志第 4巻, 1977)から hainanensis,O . platyphyllus,O . latifolius)について 判断すると,O. chingiiの一型 (v低 chingiiに相当) 検討した• O. chingiiには 2変種, var. glaucifolius と一致する.ベトナムからも O. repωnsが とvar.chingii,があるが,変種間の境界が不明瞭 Rodriguez (1934)によって記録されているが,彼 なことから,これらの変種を分類群として区別せ が引用している標本の少なくともいくつかは,今 ず、に扱った• O. chingiiは変異が大変幅広い種であ 回筆者が検したところ, O. chingiiの一型 (v訂. り,変異の極端型を比べるとそれらは別種のよう chingiiに相当)であった.0. chingiiのベトナムに に見えるが,変異は概して連続する • O. おける分布は本報告が初めての記録となる • O. hainanensisは中国の海南島から記載された種であ chingii,O . platyphyllus" O. latifoliusの 3種は子 るが,その原記載は O.chingiiのー型 (var.chingii 房上部(もしくは花柱下部)の形態 (cf.図6)等に に相当する)とよく一致することから, O. おいて差異が見られた.これらの種と類似性が高 hainanensisを O.chingiiと同種として扱った.ま い種 (0.caulescensなど)との相違点についても た,同じ海南島から,インド東部とミャンマーか 触れた(帝京大学文学部教育学科) ら知られている O.repωnsが記録されているが,

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