Taxonomic notes, new records, and a key to the adults of North American Byrrhidae (Coleoptera)

11 Pages·1991·5 MB·

by  P J Johnson

PROC. ENTOMOL. SOC. WASH.
93(2), 1991, pp. 322-332
TAXONOMIC NOTES, NEW RECORDS, AND A KEY TO THE
ADULTS OF NORTH AMERICAN BYRRHIDAE (COLEOPTERA)
Paul Johnson
J.
Department ofEntomology, University ofWisconsin, Madison, Wisconsin 53706.
Abstract.—Synonyms, lectotypedesignations, holotype recognitionsandtaxonomicnotes
are provided for North American Byrrhidae not treated elsewhere. Twenty-one names
are reduced to junior synonym status, three junior synonyms are recombined, and one
name is moved from subspecies to species status. Lectotypes are designated for 1 1 species
described by T. L. Casey, G. H. Horn, J. L. LeConte, and W. F. Erichson. Holotype data
is given for all species not previously reviewed. Byrrhuspilula L. is newly recorded from
Canada, and Sierraclava cooperi Johnson is newly recorded from Mexico. A key is pre-
sented for the identification ofthe subfamilies, genera, and species ofByrrhidae known
to occur in North America.
Key Words: Coleoptera, Byrrhidae, North America, taxonomy
The taxonomy of the North American to the known and recognized taxa in North
Byrrhidae has not been fully treated since America is provided. These data are pro-
Casey (1912) monographed the family. Ca- vided here due to delays in preparing an
sey described 46 species and subspecies, of adequate monograph on the North Amer-
which I consider 3 to be valid. In contrast, ican byrrhids, and the need to complete no-
hedescribed 6 generaandall areconsidered menclatorial clarifications for final prepa-
valid. Since Casey's monograph, Sierracla- ration of the Byrrhidae fascicle for the
va (Johnson 1982) has been the only genus United States Department of Agriculture
added, 5 new species have been described Coleoptera Catalog (R. D. Gordon, editor),
(Johnson 1985, 1986, 1991), 2 have been as well as clarifying names for use by other
species recognized and reported as immi- workers.
grants from Europe(Johnson 1990), and one Generally, only new synonymical data is
additional species is herein newly reported presented here unless inclusion ofprevious
asamemberoftheNorthAmericanbyrrhid synonymies is felt pertinent forclarity. Fur-
fauna. ther, only the more important taxonomic
During the course of continuing taxo- andfaunisticreferencesarecitedundereach
nomic and ecological studies on the Byr- synonymy. Synonymical data and com-
rhidae, a number ofsynonymies, lectotype ments in Simplocaria are presented else-
designations, and name corrections were where (Johnson, submitted).
found to be necessary, and which could not Types mentioned are reposited in the fol-
be incorporated into revisionary studies in lowing institutions: Museum of Compara-
atimelymanner. Inaddition, Byrrhuspilula tive Zoology, Cambridge (MCZ); U.S. Na-
L. is now recorded from North America for tional Museum of Natural History,
the first time, Sierraclava cooperi Johnson Washington, D.C. (USNM); Zoologisches
is newly reported from Mexico, and a key Museum Humboldt-Universitat, Berlin,
VOLUME NUMBER
93, 2 323
DDR (ZMHU); The Natural History Mu- Amphicyrta chrysomelina (pars) of Hatch
seum, London (BMNH); Canadian Nation- 1961: 301.
al Collection, Ottawa (CNC); and the Car- Amphicyrta elongata Casey 1912: 65 New
negie Museum of Natural History, Synonym. Holotype, female: "Cal; Casey
Pittsburgh (CMNH). All specimens person- bequest 1925; Type USNM 48380; elon-
ally examined have my own handwritten gata Csy" (USNM).
designation or determination labels at- Amphicyrta gentilis Casey 1912: 66 New
tached. Types ofall species have been ex- Synonym.Lectotypeheredesignated, sex
amined except where noted. notconfirmed: "Cal; Caseybequest 1925;
USNM
Type 48381; gentilis Csy"
Amphicyrta chrysomelina Erichson (USNM). Paralectotypes: 4, same data
(USNM).
Amphicyrta chrysomelina Erichson 1843: Amphicyrta gentilis ventricosa Casey 1912:
40, ofCasey 1912: 64, Hatch 1961: 301 66 New Synonym. Holotype, female:
(pars). Holotype, female: "9438; TYPUS; "Cal; Casey bequest 1925; Type USNM
chrysomelina, Koching Mont., Oregon, 48382; ventricosa Csy" (USNM).
Willcox; Zool. Mus. Berlin" (ZMHU). AmphicyrtanevadensisCasey 1912: 66New
Amphicyrta chrysomelina oblonga Casey Synonym. Holotype, female: "Nev.; Cas-
1912: 64 New Synonym. Holotype, fe- ey bequest 1925; Type USNM 48383;
male: "Cal; Casey bequest 1925; Type nevadensis Csy" (USNM).
USNM 48378; oblonga Csy" (USNM).
AmphicyrtachrysomelinaparvulicepsCasey Amphicyrta dentipes is a highly variable
1912: 65 NewSynonym. Holotype, male: speciesin sizeandcoloration,andthesewere
"Cal; Casey bequest 1925; Type USNM theprimarytraitsconsideredbyCasey. None
48379; parvuliceps Csy" (USNM). ofthecharacterstatesinvolvingsizeorcolor
are useful for unequivocal delimitation of
Casey's specimens ofA. oblonga and A. species or subspecies. Variations examined
parvulicepsaretypical examplesofv4. chrys- do suggest somegeographicregionalization,
omelina. Several hundred specimens o{A.
but there is extensive intergradation.
chrysomelina have been examined from
throughout its range, and I can find no mor- Lioligus nitidus (Motschulsky)
phological or bionomical evidence for seg-
regating local populations into several spe- Simplocaria nitida Motschulsky 1845: 362,
cies or subspecies. Coloration differences ofMannerheim 1852: 341, LeConte 1854:
noted by Casey (1912) are the same seen by 1 16. Type not seen; probably lost (Kelen-
specimens discolored by decomposing fatty ikova, in Htt.)
tissues orspecimens subjected to prolonged Lioligus nitidus (Motschulsky), of Casey
exposure to killing agents such as sodium 1912: 62, Hatch 1961: 300.
cyanide. Lioligus keeni Casey 1912: 61, of Hatch
96 30 NewSynonym. Lectotype here
1 1: 1
designated, male: "Metlakatla, B. Col.,
Amphicyrta dentipes Erichson Keen; Casey bequest 1925; Type USNM
Amphicyrta dentipes Erichson 1843: 40, 48385; keeni Csy." Paralectotypes: 3,
Casey 1912: 65. Lectotype, sex not con- same data (USNM).
firmed: "9437;TYPUS;Amphicyrtaden- LioligusstriolatusCasey 1912: 61, ofHatch
tipes Esch., Califom. Esch.; Amphicyrta 1961: 300 New Synonym. Lectotype here
Esch. Er.; Zool. Mus. Berlin 1984" designated, female: "Metlakatla, B. Col.,
(ZMHU). Paralectotype: "California Keen; Casey bequest 1925; Type USNM
Eschsch. Nr. 9437; TYPUS; Zool. Mus. 48384; striolatusCsy." Paralectotypes: 11,
Berlin" (ZMHU). same data (USNM).
324 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETYOFWASHINGTON
Lioligus aequabilis Casey 1912: 62, syn- (USNM). Paralectotypes: 10, same data
onymy by Hatch 1961: 301. Holotype, (USNM).
fUeSmaNleM: "Br. C; Caseybequest 1925; Type Two series of specimens with identical
48386; aequabilis Csy." (USNM).
labelling, less the bequestand catalogingla-
Specimens attributable to L. nitidushave bels, are reposited in the CNC and the
been examined from throughout the com- BMNH. These may be syntypical with the
posite range ofits synonyms and I can find lectotype series, but are not so treated here
no morphological orecologicaljustification due to a lack ofconfirming evidence.
for recognizing more than one species.
Morychus oblongus (LeConte)
Characters given by Casey in separating his
"species"arehighlyvariable, andareminor Pedilophorus oblongus LeConte 1857: 39,
variations in sculpture, body dimensions, ofWickham 1903: 181.
and coloration. Morychus oblongus (LeConte), of Casey
Although the type has apparently been 1912: 8, Hatch 1961: 295. Lectotypehere
lost, a neotype is not designated due to this designated, male: "medium blue disc in-
species being readily identified on morpho- dicatingthe Oregon Territory; Type 2293;
logical and distributional factors in con- PedilophorusoblongusLEC,acuminatus
junctionwithMotschulsky'sdescriptionand 7^ LEC."(MCZ). Paralectotype: samedata
denoted locality ofprovenance. There is no (MCZ).
available evidence suggestingnomenclatur- Morychus insulsus Casey 1912: 8, synony-
al confusion. my by Hatch 1961: 295. Holotype, sex
not confirmed: "Vernon, B.C., VI, Ven-
USNM
Lioliguspallidus Casey ables; Casey bequest 1925; Type
48324; insulsus Csy" (USNM).
Lioliguspallidus Casey 1912: 62.
Pedilophorus subcupreous Fall 1907: 225.
Lioligus nitidus, of Hatch 1961: 301. Lec-
Morychussubcupreous(FaW), ofCasey 1912:
totype here designated, sex not con-
firmed: "ID.; Casey bequest 1925; Type 9, synonymy by Hatch 1961: 295. Ho-
USNM lotype, male: "Aberdeen, Wash.; subcu-
48387; pallidus Csy." Paralecto- preous TYPE; MCZ Type 24474; H.C.
type: "Coeur d'Alene, Idaho" (USNM).
Fall Collection; Pedilophorus subcu-
The pallid coloration of the type is due preous Fall" (MCZ).
to its teneral condition. Typical L. pallidus Morychus insulsusrepresentsa blue-green
specimens are piceous with an olivaceous
color variant of the typical M. oblongus
sheen dorsally, whereas L. nitidus is bril- which is rarely seen in living material, but
liantly viridescent or aeneo-viridescent. is commonly induced by prolongedMexpo-
Othertraits for separatingthese two species sureto sodium cyanide. Thetypeof sub-
are in the key below.
cupreousis onlya slightlylargerand slighMtly
lessbrilliantly aeneous representative of
Exomellapleuralis Casey
oblongus.
ExomapleuralisCasey 1908: 282, ofCasey
Morychus aeneolus (LeConte)
1912: 37.
Exomellapleuralis (Casey), ofCasey 1914: PedilophorusaeneolusLeConte 1863: 74, of
378, Hatch 1961: 299,Johnson & Russell Wickham 1903: 181.
1978: 159, Johnson 1985: 155. Lectotype Morychus aeneolus (LeConte), of Casey
here designated, sex not confirmed: 1912: 8. Holotype, female: "Neb.; Pedi-
"Metlakatla, B. Co., Keen; Caseybequest lophorusaeneolusLEC; HenryUlkeColl.
1925; TypeUSNM48357; pleuraUsCsy." CMNH Ace. No. 1645" (CMNH).
VOLUME 93, NUMBER 2 325
PedilophorussubsetosusFall 1907: 225 New NorthAmericanByrrhusdeterminedassuch
Synonym. by El Moursy (1970) and previous catalog-
Morychus subsetosus (Fall), ofCasey 1912: ers (Hamilton 1894a, b; Dalla Torre 1911;
14, Hatch 1961: 295. Holotype, female: Leng 1920). Interestingly,neitherofthemo-
"Kalispell, Mont., June 13, Wickham; nographers of North American Byrrhidae,
subsetosusTYPE;TypeMCZ24475;H.C. LeConte(1854)orCasey(1912),appliedthe
Fall Collection; Pedilophorus subsetosus name B.fasciatus to any native species. El-
Fall" (MCZ). Moursy had incorrectly recognized B. fas-
Pedilophorus lateralis Fall 1907: 225 New ciatus as occurring in North America, for
Synonym. reasons unknown to me. My examination
Morychus lateralis (Fall), ofCasey 1912: 9. ofthetypeof5. cyclophorusandconspecific
Holotype,female: "N.M., LasVegas, head specimens from throughout North Ameri-
ofDailyCan.; 6.26.0 T.D.A.C;lateralis ca, typical B. fasciatus from Europe, and
TYPE; Type MCZ 214;473; Pedilophorus referencetotaxonomic treatmentsbyJohn-
lateralis Fall" (MCZ). son (1966), Paulus (1979), and earlier au-
Pedilophorus hesperus Wickham 1903: 182 thors, clearly shows the regional misappli-
New Synonym. cation ofthis name.
Morychus hesperus (Wickham), of Casey
Byrrhus geminatus LeConte
1912: 9. Holotype, female: "Leadville,
Col., H.F. Wickham, July 7-14 96, Byrrhus geminatus LeConie 1854: 114, of
10,000-11,000 ft.; Wickham Collection El Moursy 1970: 330, Fiori 1982: 437.
1933; TYPE; Pedilophorus hesperus Holotype, female: "Light blue disc with
Wickham" (USNM). two converging tangential cuts indicating
Morychus albertanus Brown 1932: 8 New thenorthernshoreofLakeSuperior; Type
Synonym. Holotype, sex not confirmed: 2297;B.geminatusLEC, L. Sup."(MCZ).
"Crow'sNestPass,Alberta,June7, 1930, Byrrhus pettiti Horn 1870: 76, synonymy
J.H. Pepper; No. 3246" (CNC). by ElMoursy 1970: 330, Fiori 1982: 438.
Lectotypeheredesignated: Female; "Can.;
Thevarioussynonymsforthis species re- B. pettiti Horn; Lectotype 3260" (MCZ).
flect slight variations of integument color Paralectotype: same data (MCZ).
and patterns of pubescence. Minor sculp-
tural variations occur, but none correlate Neither El Moursy (1970) or Fiori (1982)
with discreet populations. Slight variations examinedthetypematerialof5.pettiti, and
anddistributionalpatternsofgenitalicchar- did not designate a type; however, they did
acters and pubescence support the synon- correctly synonymize it with B. geminatus.
ymies,butalsosuggestthepresenceofclines The label reading "Lectotype 3260" is ap-
over large geographic areas. parently a cataloging label and has no tax-
onomic standing, and was apparently placed
Byrrhus cyclophorus Kirby on the specimen while the Horn Collection
was held by the Academy of Natural Sci-
Byrrhus cyclophorus Kirby 1837: 1 17, ofEl ences, Philadelphia (A. Newton, pers.
Moursy 1970: 329. Holotype, female:
comm).
"Type; N.Amer.; 5828a; Byrrhus cy-
clophorus Kirby, N. Amer., 5828, Rev. Byrrhus kirbyi LeConte
Wm. Kirby (BMNH).
Byrrhus kirbyi LeConte 1854: 114, of El
Byrrhusfasciatus, ofElMoursy 1970: 330 Moursy 1970: 331. Holotype, female:
New Combination.
"light blue disc with two converging tan-
The application ofthe name B.fasciatus gential cuts indicating the northern shore
is here restricted in its application only to ofLake Superior" (MCZ).
326 PROCEEDINGSOFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON
ByrrhnsfulvovestitusCasey 1912: 27, ofEl Until now, B. pilula has been considered
Moursy 1970: 329. New Combination. only a Eurasian species (Dalla Torre 1911,
Neither LeConte nor any subsequent Winkler 1926, Fiori 1951, Horion 1955,
worker labelled the above designated spec- Bonadona 1975). The lack ofprevious rec-
imen ofB. kirbyi in a manner indicating its ognition of this species in North America
identity or status. The entire byrrhid series is ascribed to inadequate series ofmaterial,
intheLeContecollectionwasexaminedand especially males, and to the failure of the
only one specimen was found which fit most previous byrrhid workers to appreci-
LeConte's description and possessed the ate the potential of Holarctic distributions
correct label for the type locality. The spec- in the family (see also Johnson, submitted).
imen had been placed next to specimens of Further complication ofdetermination can
B. concolor Kirby and Porcinolus undatus be ascribed to a close relationship ofB. pi-
(Melsheimer) in an unlabelled series, pos- lula and B. americanus, as indicated by ae-
sibly for comparative purposes. This spec- deagal morphology, and a general difficulty
imenishereinconsidered LeConte'sunique ofcorrelating unassociated Byrrhusfemales
type ofB. kirbyi, and has been so labelled. with males; this latter difficulty may also
El Moursy (1970) provided a lectotype help explain occasional confounding of B.
designation, but incorrectly syonymized B. pilula with B. cyclophorus by previous
fulvovestituswithB. cyclophorus, apparently workers.
under the impression that Byrrhus females
Porcinolus undatus (Melsheimer)
are not determinable to species. Casey's
specimens are badly rubbed and slightly Byrrhus undatus Melsheimer 1844: 1 17.
narrowin form, butotherwisearetypicalB. Byrrhus murinus Fabricius 1794: 437, of
kirbyi. LeConte 1854: 115.
Porcinolus undatus (Melsheimer), ofCasey
Byrrhuspilula Linnaeus
1912: 33. Holotype, female: "Melsh.; un-
datus M.; murinus; [red torn paper]"
Recently, a series of Byrrhidae from
(MCZ).
nfoorrtdheetrenrCmainnaatdiaonwabsygiJf.tedP,ilwniyt,hWaarteeqruleosot ByrCrahsuesygl1a9b1e2l:lu3s3.MeHloslhoetiympeer, 1s8e4x4:no1t17c,ono-f
University,Ontario. Includedwere78 spec-
firmed: "Melsh.; glabellus *Melsh.; [red
imens ofa Byrrhus which could not be ac-
torn paper]" (MCZ).
curately assigned to any described or re- PorcinoluscrescentiferCasey 1912:32 New
corded North American species; however, Synonym. Holotype, sex not confirmed:
its close affinity with B. americanus Le-
"Baldwin, Kansas; Casey bequest 1925;
Contewasevident. Furtherinvestigationhas USNM
Type 48354; crescentifer Csy"
shown that these specimens are conspecific
(USNM).
with examples of B. pilula, of British and Porcinolus hystrix Casey 1912: 33 New
Austrian provenance.
Synonym.Lectotypeheredesignated, sex
NorthAmericanspecimensascribedtoB.
not confirmed: "Aweme, Manitoba, S.
pilula have been seen from the following
Criddle, 23.VL10; Casey bequest 1925;
locality: Canada, Northwest Territories, USNM
Type 48355; hystrix Csy." Para-
Keewatin, Lat +62.41 Long +97.03, What-
lectotype: same data (USNM).
ever Lake, June 1989. All specimens were
collected by pitfall traps in tundra and ri- All ofthe Melsheimer and Casey names
parian habitats. Specimens are in my col- represent simple color pattern variations of
lection and representatives will be depos- the pubescence. Porcinolus undatus is dis-
ited in the USNM and CNC. tributed from the Atlantic coast to the east-
VOLUME 93, NUMBER 2 327
era slopes of the Rocky Mountains, with USNM 48330; longulus Csy" (USNM).
the westera specimens being slightly small- Paralectotypes: 10, same data (USNM).
er. The transelytral crescentiform pattern is Cytilusmimicusdiffersfrom C alternatus
highly variable from well colored and con- by its larger size and more elongate form,
tiguous to indistinctly colored and broken slight differences in aedeagal structure, and
in outline. habitat. This proposed new status for C
mimicus is an attempt at better recognition
Cytilus alternatus (Say)
of two ecologically segregated, but geo-
Byrrhus alternatus Say 1825: 186, of Le- graphically sympatric forms of Cytilus in
Conte 1854: 115. North America, which are difficult to di-
Cytilus alternatus (Say, of LeConte 1870: agnose morphologically. Considerable ef-
398,Casey 1912: 18.Typenotfound, pre- fort is still required for a full evaluation of
sumed lost (LeConte 1859, Lindroth & both species and their relationship to Eur-
Freitag 1969). asian species.
Byrrhus trivittatus Melsheimer 1844: 117,
original synonymybyLeConte 1854: 115, Curimopsis echinata (LeConte)
LeConte 1877: 108, Casey 1912: 18, Syncalypta echinata LeConte 1850: 224, of
Hatch 1961: 297. Holotype, sex not con- LeConte 1854: 114.
firmed: "Melsh.; varius; red torn paper; Curimopsis echinata (LeConte), of Casey
trivittatus 'Melsh.'" (MCZ). 1912: 34, Johnson 1986: 42. Holotype,
Cytilus nigrans Casey 1912: 19 New female: lightbluediscindicatingthe Lake
Synonym.Holotype, female: "LittleRiv- Superior region (MCZ).
er, Colroy, NFLD, July 10-18 '07; Casey Curimopsis brevicollis Casey 1912: 35, of
USNM
bequest 1925; Type 48331; ni- Hatch 1961: 299, synonymy noted by
grans Csy" (USNM). Johnson 1986:42. Holotype,male:"W.T.;
USNM
MyinterpretationofC. alternatusisbased Caseybequest 1925; Type 48356;
on LeConte's understanding ofthis species brevicollis Csy." (USNM).
asrepresentedbyhisspecimensattheMCZ. Morphological differences between C
The type ofCytilus nigrans is badly rubbed brevicollis and C echinata used by Casey
of pubescence, as noted by Casey (1912), (1912) seem to be allometric and sexual.
while that ofC. trivittatus lacks aeneous or The identical aedeagi and female gonocox-
viridescent elytral patches. Otherwise these ites ofthe types and other specimens sup-
specimens are typical examples ofC. alter- port this synonymy.
natus.
Curimopsis albonotata (LeConte)
Cytilus mimicus Casey
Syncalypta albonotata LeConte 1861: 344.
Cytilus mimicus Casey 1912: 18 New Curimopsis albonotata (LeConte), ofCasey
Status. 1912: 34, Hatch 1961: 299,Johnson 1986:
Cytilus alternatus mimicus CdiSty 1912: 18. 42. Holotype, female: "W.T.; S. albono-
Lectotype here designated, sex not con- tata LLC; Type 2302" (MCZ).
firmed: "Cal.; Casey bequest 1925; Type Syncalypta grisea LeConte 1879: 514.
USNM
48329; mimicus Csy" (USNM). Curimopsisgrisea(LeConte), ofCasey 1912:
Paralectotypes: 2, same data (USNM). 35, synonymy noted by Johnson 1986:
Cytilus alternatus longulus Casey 1912: 18, 42.
Hatch 1961: 297 New Combination. Curimopsis brevicollis. Hatch 1961: 299.
Lectotype here designated, sex not con- Holotype, female: "Garland, Col., 19.6;
firmed: "W.T.;Caseybequest 1925; Type 376; S. grisea LEC; Type 2300" (MCZ).
328 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON
It is difficult to understand why LeConte temisia scrub, Adenostoma chaparral, or
didnotobservethequiteobvioussimilarity mixed sarcophyllous scrub, while Sierran
ofthese two species. His own descriptions specimens were found in Quercus-Pinus
indicate only minor color shading and size woodlands.Whetherthemorphologicaland
differences between the specimens, and di- habitatdifferencescanberefinedtoindicate
rect comparison reveals these same differ- two allopatric species is still being investi-
ences. Conspecificityissupportedbygenital gated.
morphology.
Key TO THE Adults of
North American Byrrhidae
Sierraclava cooperiJohnson
This species was originally recorded la. Antennae filiform, compressed; palps with
ultimatesegmentsecuriform;appendagesnot
(Johnson 1982)fromthevicinityofSequoia retractile;tarsomere3withlargefleshylobe;
National Park, in the southern Sierra Ne- integument appearing glabrous, rufocasta-
vada, Cahfomia, U.S.A. Since then, addi- neus to piceous Amphicyrtinae 2
tionalmaterialhasaccumulatedfrom 8 new lb. Antennae short, clavate or capitate; palps
with ultimate segment fusiform or pyri-
disparate locations through central and
form 3
southern California, as well as a new coun- 2a. Usually larger, 8-12 mm, elongate; integu-
tryrecordfromBajaCaliforniaNorte, Mex- mentusuallyrufocastaneuswithmetalliccu-
ico. These new records [abbreviated data] preous sheen, occasionally with viridescent
are: U.S.A., California, Amador Co., Elec- reflections;coastalconiferousforestsofOr-
tra, Mokelumne River; Calaveras Co., 3 mi egon and northern California
Amphicyrta chrysomelina Erichson
south ofMokelumne Hill; Fresno Co., 3 mi 2b. Usuallysmaller,5-10mm,ovoid,occasion-
northeastofAuberry, 8 misouthwestofAu- ally inflated posteriorly; integument rufo-
berry, and 9 mi east ofCoalinga; Riverside piceous to piceous, without metallic sheen;
Co., Lamb Canyon, 2 mi northwest ofGil- prairies and oak-pine woodlands ofcentral
man Hot Springs; San Benito Co., 18.4 mi Californiaand southwestern Oregon
Amphicyrtadentipes Erichson
northwest ofNew Idria, and 1.8 mi south- 3a. Antennaecapitate;bodysmall(0.9-2.6mm),
westof[New] Idria; Mexico,BajaCalifornia shallowlytomoderatelyconvex;appendages
Norte, 10.7 km east of El Rosario, Lat closely retractile; dorsum with clavate or
+30.04.35 Long +1 15.38.25, 1 1.7 km east truncatebristles Syncalyptinae 4
of El Rosario, Lat +30.04.30 Long 3b. Antennae clavate; body various (1.9-8.7
+ 115.37.55, and 14.7 km east ofEl Rosa- tmom)el,onmgoadtee;raatpepleyndtoagsetsronnogltyocropnavretxi;alolyvartee-
rio, Lat +30.04.1 Long + 1 15.36.00. Most ceived intofossae;dorsum withdecumbent
specimens were collected in pitfall traps to erect fine setae Byrrhinae 10
during late winter and spring months. Cal- 4a. Length2.4-3.2mm;ovoid,dorsumwithap-
itfhoernsitaafsfpoefcitmheensCawleifroerncioalleDcetpeadrltamrgeenlty boyf 4b. eLplreyentsgrsatelhds0ts.rc9ia-ale2e.-pl1uinkmcemtas;teteoa;veadtaoenr,dssuetrmreocnwtgiltbyhricsbotrnlievsstelxe;s. 5
Food and Agriculture, Sacramento, and the only; northern U.S. and southern Canada,
Mexico specimens by William H. Clark, Europe Chaetophoraspinosa (Rossi)
College ofIdaho, Caldwell. 5a. Ovatetooval,shallowlytomoderatelycon-
The apical portion of the penis of the vex, lateral margins arcuate; striae on disc
shallowly impressed, or serially punctate 6
Mexican specimens is much less broadened 5b. Elongate,stronglyconvex,parallel-sided;el-.
and spatulate than of northern specimens ytra with striae deeply punctate and im-
from Sierran localities, and specimens from pressed, sulciform; central and southern
CoastRange sitesareintermediatein penial California, BajaCalifornia Norte
Sierraclava cooperiJohnson
form. Coast Range and Mexican specimens
6a. Length 2.1-2.8 mm; subparallel laterally;
have been found in desert or semidesert appressed squamae linear; median lobe of
communities dominated by Juniperus-Ar- aedeagus slender, narrowly acuminate api-
.
VOLUME 93, NUMBER 2 329
cally; Alaska tessellate; elytral striaecompleteorbecom-
Curimopsissetulosa (Mannerheim) ingevanescent preapically 13
6b. Length 2.8-3.2 mm; oval in lateral outline; 13a. Length3.5-^.5 mm;pronotalpuncturesfine
aedeagus not as above 7 and sparse on disc; elytral striae becoming
7a. Dorsalsquamaeround,setintoroundpunc- evanescent towards apex; pubescence de-
tures; 3.0-3.2 mm; subparallel laterally; cumbent;northemCanada,Greenland,Eur-
western and northern U.S., Canada asia Simplocaria elongataJ. Sahlberg
Curimopsisalbonotata(LeConte) 13b. Length 2.8-3.5 mm; pronotal punctures
7b. Dorsal squamae more-or-less linearto nar- moderateon disc; elytral striaecompleteto
rowly subtriangular, emergent from small apex; pubescence recumbent to suberect;
usually barely evident punctures; usually northem U.S., Canada, Greenland, Eurasia
smaller 8 Simplocaria metallica (Sturm)
8a. Bodyoval, rounded laterally; striaenot im- l^a. Pubescence simple, slender, fine; epipleura
pressedaslinesorshallowsulci,usuallyrep- flat; integument shiningto metallic 16
resentedbyrowsofpunctures;south-central 14b. Pubescencestout,recurved;epipleuradeep-
U.S Curimopsisstrigosa (Melsheimer) ly emarginate to receive femoral knee; in-
8b. Body elongate, subparallel laterally; striae tegument brunneous 15
narrowly,shallowlyimpressed,notevident- •5a. Length 1.8-2.3 mm; lateral elytral margin
ly punctate 9 crenate anteriorly; coastal Oregon, Wash-
9a. Length 2.9-3.2 mm; appressed squamae ington, British Columbia
short, subtriangular; elytral apical declivity Exomellapleuralis(Casey)
long, gradually sloping; sutural stria at de- 15b. Length 2.3-2.4 mm; lateral elytral margin
clivity shallowly impressed; northeastern smooth throughout; northem Idaho
U.S., Canada, Alaska Exomella merickeliJohnson
Curimopsis moosilaukeJohnson 16a. Epipleura broad, extending length of ely-
9b. Length 2.8-2.9 mm; appressed squamae tron; dorsal integument piceous to oliva-
longer, linear; elytral apical declivity short- ceous, shining 17
er,abruptlysloping;suturalstriaatdeclivity 16b. Epipleura narrow, attenuating and termi-
distinctly and deeply impressed, sulcate; natingbeforesecondvisibleabdominalster-
northern U.S., Canada, Alaska nite; dorsal integument viridescent 18
Curimopsisechinata (LeConte) 17a. Larger,3.2-4.1 mm,lateralmarginsinflated;
10a. Frontal margin beaded, occasionally re- integument piceous, shining, occasionally
flexed and carinate; mesostemum strongly withsubmetallicreflections;pubescentsparse
reducedexceptformedianfossa;palpswith to moderate, short to long; north-coastal
ultimatesegment fusiform to narrowlypyr- Califomia tosoutheastem Alaska, and west
iform;metacoxaesmall,distantfromelytral ofCascade Range crest in Oregon, Wash-
margin; elytra often connate 11 ington, British Columbia
10b. Frontal margin rounded or truncate, never Lioonsimplicipes(Mannerheim)
reflexed, beaded or carinate; mesostemum 17b. Smaller,2.9-3.6mm,lateralmarginsweakly
distinct laterad ofmedian fossa; palps with rounded; integument piceous, olivaceous,
ultimate segment compressed or cylindri- without submetallic reflections; pubescence
cally pyriform; metacoxae flattened, nearly moderately dense, long; northem Idaho
. .
reachingelytral margin; elytra not connate 19 Lioon nezperceJohnson
11a. Elytraseparateandmetathoracicwingspres- 18a. Length2.4-2.6mm;dorsalintegumentbright
ent; integument rufopiceous to piceous, oc- viridescent; west ofCascade Range crest in
casionally with submetallic reflection; me- Oregon, Washington, British Columbia . .
socoxae moderately separated 12 Lioligus nitidus(Motschulsky)
lib. Elytra connate, metathorax apterous; me- 18b. Length 2.2-2.4 mm; dorsal integument ol-
socoxae widely separated 14 ivaceouswithevanescentviridescentreflec-
12a. Form short, ovate, slightly depressed dor- tions; northem Idaho . . LioliguspallidusCasey
sally; elytral pubescence long, moderately 19a. Frontal margin tmncate and thickened; ab-
dense, evenly distributed; elytral stria be- dominal stemite 1 without crural modifi-
comingevanescentatmidlength; northeast- cations 20
em U.S., southeastem and southwestern 19b. Frontal margin obtusely rounded; abdomi-
Canada Simplocariasemistriata(F.) nalstemite 1withdistinctcmraldepressions
12b. Form elongate, ovoid; elytral pubescence or fossae 22
330 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OFWASHINGTON
20a. Form subparallel, moderately convex dor- 26a. Integument piceous, frequently with viri-
sally; piceous, without metallic patches or descent reflection; punctures moderate to
reflections; pubescence long; decumbent, dense;pubescencecinereouswithdarkbrun-
with erect black setae; northern U.S., Can- neouspatches; montanewestern U.S.,Can-
ada, Greenland . . . Tylicussubcanus(LeConte) ada Morychusaeneolus(LeConte)
20b. Formovoidtosubparallellaterally,strongly 26b. Integument aeneous to cupreous, occasion-
convexdorsally;piceous,frequentlywithcu- ally with viridescent reflections; punctures
preous or viridescent patches or intervals; fine to moderately dense, moderately to
pubescence short, appressed, without erect sparsely distributed; pubescence mixed ru-
setae 21 fobrunneousandcinereous;westernU.S.and
21a. Form ovoid, short; integument rarely with Canada Morychusoblongus(LeConte)
cupreousorviridescentpatches;pubescence
27a. Form oval to subparallel, strongly convex
black and cinereous, tessellate on alternate
dorsally; pubescence simple, appressed ... 28
elytral intervals; transcontinental
27b. Form ovate, shallowlyconvexdorsally; pu-
Cytilus alternatus(Say)
21b. Formelongatetosubparallel; elytra usually bescence including erect, bristle-like setae;
with viridescent patches or intervals; pu- central and eastern U.S., southern Canada
Porcinolus undatus(Melsheimer)
bescence pale cinereous, rarely tessellate;
montane western U.S. and Canada 28a. Pronotalandelytral integumentshiningbe-
Cytilus mimicusCasey tweenmoderatesizedpunctures;sparselyand
22a. Tarsomere4withmembranousventrallobe; finelymicroreticulate 29
dorsal integument shining, usually virides- 28b. Pronotal and elytral integument dulled due
centand/orcupreous 23 todense, finetocoarsemicroreticulationor
22b. Tarsi simple; dorsal integument opaque or micorugosities;puncturessmall,oftenoblit-
shining, piceous to black 27 erated 32
23a. Form ovate; aeneous to viridescent; punc- 29a. Pronotal punctures smaller, well separated
turesobsolescentorcoarseand formingru- on disc by interspacesgreaterthan own di-
gose patches on elytra; pubescence minute ameter 30
ormoderatelylongandpatchy,usuallymixed 29b. Pronotal punctureslarger, separatedby less
cinerousand rufobrunneous 24 than own diameter; northern U.S., Canada
23b. Form subparallel; piceous with viridescent Byrrhusgeminatus LeConte
reflections to cupreous; punctures fine to 30a. Puncturesofheadcoarse,confluent,rugose;
moderate; pubescence moderately long, similarbut shalloweron elytra 31
evenly distributed 26 30b. Puncturesofheadlargeandshallowonfrons;
24a. Length6.5-10.0mm;dorsalpuncturesfine; deepandmoderatelydenseonelytra;north-
subglabrous,pubescenceminute;aeneousto em U.S., Canada ByrrhuscyclophorusKirby
viridescent, with elytra bearingdistinct ae- 31a. Punctures on pronot. a.l disc small, subequal
neous and viridiaeneous vittae; northern oronlyslightlylargerindiameterthanbase
Idaho, northeastern Washington, western
ofseta;northeasternU.S.,southeasternCan-
Montana, southeastern British Columbia
ada ByrrhusamericanusLeConte
Eusomalia lecontei(Wickham)
24b. Length 3.2-4.0 mm; punctures coarse, pu- 31b. Punctures on pronotal disc larger, 2-3x
wider than seta; northern Canada, Eurasia
bescence usuallyevident, moderately long 25
Byrrhuspilula L.
25a. Dorsalpuncturessmalltomoderatelycoarse,
simple, forming rugose patches with little 32a. Punctures on elytra larger, distinct; sculp-
suggestionofstrialformation,orsmoothwith turingweak 33
finely punctate striae on elytra; aeneous to 32b. Punctures on elytra minute, often obliter-
viridiaeneous,oftenvittateonelytra;north- ated; sculpturing transverse, reticulate, mi-
coastal California to Sitka, Alaska, west of crorugose,orundulating;northeasternU.S.,
Cascade Range crest in Oregon, Washing- Canada ByrrhusconcolorKirby
ton, British Columbia 33a. Length >6.5 mm;elytralpuncturessmaller,
Listemusacuminatus(Mannerheim) dense but separated, rarely rugulose; north-
25b. Dorsal punctures large, coarse, umbilicate, em and western U.S., Canada
not condensing into rugose patches on ely- Byrrhus kirbyiLeConte
tra; piceous, with only faint aeneous reflec- 33b. Length <6.0 mm; elytral punctures dense,
tions; northern Idaho, northeastern Wash- usuallycoalesced,mgulose;westernU.S.and
ington ListemuskootenaiJohnson Canada Byrrhuseximius LeConte
VOLUME 93, NUMBER 2 331
Acknowledgments Europei: II Contribute alia conoscenza della fa-
miglia Byrrhidae(Coleoptera). BoUetinade Insti-
My thanks are extended to J. M. King- —tutEntomologiaUniversite,Bologna 18:293-304.
solver, U.S. National Museum of Natural . 1982. ByrrhusgeminatusLeConte:Speciead
History, Washington, D.C.; A. F. Newton, ampia geonemia olartica, XIII Contribute all co-
Jr. (formerly) and D. Furth, Museum of noscenza della famiglia Byrrhidae (Coleoptera).
Comparative Zoology, Cambridge; H. Uh- Bolletinade Museo Divisio Storia Naturella, Ve-
rona 9: 437-447.
lig, Humboldt-Universitat, Berlin; J. M. Forster,J. R. 1771. NovaeSpeciesInsectorum.Cen-
Campell,AgricultureCanada,Ottawa; R. L. turia I. T. Davies & B. White, London.
Davidson, Carnegie Museum of Natural Hamilton, J. 1894a. Catalogue ofthe Coleoptera of
History, Pittsburgh; F. G. Andrews, Cali- Alaska, with the synonymy and distribution.
forniaDepartmentofFoodandAgriculture, Transactionsofthe American Entomological So-
ciety 21: 1-38.
Sacramento; and William H. Clark, College
1894b. CatalogueoftheColeopteracommon
ofIdaho, Caldwell, for the loan ofmaterial to.NorthAmerica,northernAsia,andEuropewith
and/or permission to examine material un- distributionandbibliography.Transactionsofthe
der their care. Thanks are also extended to American Entomological Society 21: 345^16.
the entomologists ofthe CNC and Carleton Hatch,M.H. 1961. TheBeetlesofthePacificNorth-
University, Ottawa, fortheirhospitalityand west, part III: Pselaphidae and Diversicomia I.
University ofWashington Press, Seattle.
visitation grants from the CanaColl Foun-
Horion, A. 1955. Faunistik der Mitteleuropaischen
dation. Thanks are extended to A. F. New- Kafer: Stemoxia, Fossipedes, Macrodactyla, Bra-
ton, Jr., S. E. Miller, fortheircomments on chymera. Entomologische Arbeiten 4: 222-249.
early drafts, J. M. Kingsolver for consid- Horn, G. H. 1870. Contribution to the coleopter-
erable assistance with specimens, informa- ology of the United States. Transactions of the
tion, and reviewing ofmanuscripts, and to American Entomological Society 3: 69-142.
D. S. Chandler for comments on the draft Johnson, C. 1966. The British species ofthe genus
ByrrhusL., includingB. arietinusSteffahny(Col.,
key. Byrrhidae) new to the British list. Entomological
Monthly Magazine 101: 111-115.
Literature Cited Johnson,P.J. 1985. AnewspeciesofExomellafrom
Idaho, with notes on the biology of Exomella
Bonadona,P. 1975. LesByrrhus(sensulato)deFrance pleuralis (Casey) (Coleoptera: Byrrhidae). Cole-
(Col., Byrrhidae). L'Entomologiste31(6): 193-209. opterists Bulletin 39(2): 151-157.
Brown, W. J. 1932. New species ofColeoptera, II. 1986. AnewspeciesandakeytotheNearctic
Canadian Entomologist 64: 8-9. .
Casey, T. L. 1908. A new genus ofByrrhidae. Ca- species of Curimopsis Ganglbauer (Coleoptera:
Byrrhidae). Coleopterists Bulletin 40(1): 37-43.
nadian Entomologist 40: 281-282.
1912. DescriptivecatalogueoftheAmerican . 1990. Notesonthenaturalizationoftwospe-
BMyer..mroh1ii9dr1as4e..onMMeitmshceoeiClrolslaenooenpotuteshrenaoCt5o:els3e7oa8pn.tdernaew3:s1pe-c6i9e.s. cASiomecesireoitfcyaE.9u8rJ(oo4)pu:rena4an3l4Bo-yf^r4tr0hhe.idNaeew(CYoolreokptEenrtao)moilnoNgoirctahl
Dalla Torre, K. W. von 1911. Fam. Byrrhidae, pp. . 1991 (Inpress). TaxonomicreviewsofLioon
5-36. In Schenkling's Coleopterorum Catalogus, Casey and Listemus Casey, with descriptions of
pars 33. W. Junk, Beriin. twonewspecies(Coleoptera:Byrrhidae).Proceed-
ElMoursy,A.A. 1970. ThetaxonomyoftheNearctic ingsofthe Entomological SocietyofWashington.
speciesofthegenusByrrhusLinnaeus(Coleoptera: . (submitted). Synonymical notes for some
Byrrhidae).QuaestionesEntomologica6: 327-338. speciesofthe metallica-groupofSirnplocariaSte-
Erichson, W. F. 1843. [revisionary notes and mis- phens(Coleoptera:Byrrhidae).EntomologicaFen-
cellaneous species descriptions]. /«StefThany, G., nica.
ed., Tentamen Monographiae Generis Byrrhi. Johnson, P. J. and L. K. Russell 1978. Noteson the
Zeitschrift fiirdie Entomologie 4: 1-42. rediscovery, habitat, and classification of Exo-
Fall, H. C. 1907. The Coleoptera ofNew Mexico: mella pleuralis (Casey) (Coleoptera: Byrrhidae).
Descriptions ofnew species. Transactions ofthe Coleopterists Bulletin 32(2): 159-160.
American Entomological Society 33: 145-272. Kirby, W. 1837. Northernzoology, partIV, Insecta,
Fiori,G. 1951. AlcuniAppuntisuifivA'/jM^^L.,s.str.. pp.iii-xxxix, 1-325. InRichardson,J.,ed..Fauna