Study of Asian Strongyliini (Coleoptera, Tenebrionidae) : XI. Ten New Small-sized Species of the Genus Strongylium from Southeast Asia

18 Pages·2001·21.5 MB·English

by  Masumoto, K

Ely1t・a,Totkyo,29(2):401-418,November15,200]
Studyof AsianStrongyliini (Coleoptera,Tenebrionidae)
XI. TenNewSmall-sizedSpeciesoftheGenus
SlrongyliumfromSoutheastAsia
Kimio MAsUMoTo
InstituteofHumanLivingSciences,0tsumaWomen'sUniversity,
Tokyo,102-8357Japan
Abstract ThisistheeleventhpartofthestudyoftheAsianStrongyliinianddeals
with10newspeciesofthegenusSt,-o1lgylit1,n fromSoutheastAsia,whicharedescribed
underthefollowingnames:St,-ongy/llm,nolg/fan11sp nov.,S nagafisp nov.,S nofsui
sp nov.,S.walshaesp nov.,S.1nu1ot1sp nov.,S.stt'ambensesp nov.,S moerianumsp
nov.,S tsuyukie11umspnov.,Snagale111m1sp nov.,andSdo1okensesp nov.
Thispaper istheeleventhpartofmystudy oftheAsianStrongyliinianddeals
withtennewspeciesofthegenusStrongy11umfromvariousareasofSoutheastAsia.
All thespeciestobedescribedissmall inbodysize(lessthan10mm),sothatonlya
fewmembershavebeenhistoricallyrecognized.Pfcdescribedlessthanfivespecies.
GEBIENpreparedtodescribemorethantenspecies,buthedidnot finishitbecauseof
theWorldWarIIandtheconfusionfollowedlt.Iamgoingtodescribetennewspecies,
includingGEBIENs' invalidones.
Thespecimensexaminedaresubmittedtomefor taxonomicstudy fromthecol-
lectionsoftheMuseumNationald'HistoireNature11e,Paris,and theMuseumand In-
stituteofZoology,PolishAcademyofSciences,Warsaw.0thermaterials,besidesthe
specimensinmycollection,arealsosubmittedtomepersonally fromDr.Wolfgang
ScHAwALLER,Staatliches Museumftir Naturkunde,Stuttgart, Ing.Stanislav BECvAR,
InstituteofEntomology, Czech Academy ofSciences, Dr. MasatakaSAT0,Nagoya
Womens' University,Messrs.ShigeoTsuYuKl,ZushiCity,andMinoruTAO,Yokohama
City.
IwishtoexpressmycordialthankstoDr.ClaudeGIRARDandMileJeanneCHAR-
BoNNEL,MuseumNationald'HistoireNature11e,Paris,Mr.MartinJ.D.BRENDELL,the
Natural HistoryMuseum, London, Ing.Stanislav BEcvAR, andDr. KiyoshiANDo,
OsakaPrefecture, for their invaluablesupport inthe courseof thepresentstudy. I
thankDr. Michel BRANcUccl andDr. Daniel BURcKHARDT, theNatural History Mu-
seum, Basel, andDr. Dariusz IwAN, theMuseum and Instituteof Zoology,Polish
AcademyofSciences,Warsaw, for permittingmetoexaminetheGEBIENcollection
preservedintheMuseums.Appreciationisduetotheabovepersonswhoofferedme
402 KimioMAsUMOT0
materials,andalsotoMr.Seiji MORITAandMr.KaoruWADA,Tokyo, for takingpho-
tographsanddrawingfiguresbothinserted in thispaper.Finally,my deepest thanks
shouldbeexpressedtoDr.Shun-IchiUENo,National ScienceMuseum(Nat.Hist),
Tokyo, forhisconstantguidanceinmytaxonomicstudies.
Depositoriesoftheho1otypestobedesignatedaregivenineachdescription.The
abbreviations used herein are as follows: NSMT- Nationa1 Science Museum (Nat.
Hist),Tokyo;MNHNP-MuseumNationald'HistoireNature11e,Paris; NHMB-Nat-
ura1HistoryMuseum,Basel;SMNS-StaatlichesMuseumfiirNaturkunde,Stuttgart.
Sfron l加mmodiglianll'sp nov
(Figs.1,11-12)
Brown, head and pronotum blackish brown with feeble dark greenish lustre,
scutellum and elytrapartly with feeblegolden lustre, head andpronotum weakly,
sericeouslyshining,scutellum,elytraandventralsurfaceslightlyalutaceouslyshining,
legsmoderatelyshining;almostglabrous.Elongated fusiform, longitudinallyconvex,
thoughweakly attenedinmedic-basalhalvesofelytra.
Head transverselyelliptical,very feeblycoveredwith isodiametricmicrosculp-
ture, closely, finely punctate;clypeussomewhat transversely hexagonal, attened in
basal part, gently inclined forwards, truncateat apex, fronto-clypeaI border gently
curved and indistinctly sulcate; genaeoblique, raisedoutwards, with rounded outer
margins; fronsrather wide,gently inclinedanteria diatone 12 times the width of
transversediameterofaneye,withanimpunctateareaatthemiddleofposteriorpart.
Eyesmedium-size subreniform, convex latera triangularly,shallowly inlaid into
head.Antennaereachinghumeriofelytra, ratioof thelengthofeachsegment from
base to apex:0.32,0.2,0.59,0.39,0.32,0.27,0.23,0.26,0.23,0.25,0.29.
Pronotumwidelyhexagonal,1.36timesaswideaslong; apexsublinear, finely
rimmedoneachside,vaguelymarginedinmiddle;basegentlysinuousoneachside,
borderedandweaklyridgedinmiddle;sidessteeplydeclinedtolateralmargins,which
areproducedandentirelyfinelyridge eachwithatoothslightlybehindthemiddle;
frontanglesrounded,hindanglesobtuselyangulate;discgentlyconvex,weakly im-
pressedclosetobaseoneachside,weaklydepressedinmedic-basalparts,veryfeebly
coveredwithisodiametricmicrosculpture,rathercloselypunctate.Scutellumtriangular
with rounded sides, feebly elevated, very weakly covered with isodiametric mi-
crosculpture,sparselyscatteredwithmicroscopicpunctures.
Elytraelongatedsubovate,1.7timesaslongaswide,3.65timesthelengthand
1.30timesthewidthofpronotum;dorsumlongitudinallyconvex, though attened in
medic-basal part, weakly depressed in area along scutellar striole; disc punctate
groove 5thgroovedeepenedandtouchingbase,thepuncturessmall, ratherclosely
set, becoming smaller posteriad; intervals gently convex, attened posteriad, very
weaklycoveredwithisodiametricmicrosculpture,sparselyscatteredwithshallowmi-
croscopicpunctures,very feebly, rather transverselywrinkled;humerigentlyswollen;
StudyofAsianStrongyliini,XI 403
apicesroundlyproduced.
Maleanalsterniteweakly,sube11iptica1lydepressed inapicalpart. Legsrather
slender, no special modification; ratiosof the lengthsofpro-,meso- and metatar-
someres:0.27,0.24,0.22,0.23,1.2;1.2,0.56,0.52,0.29,1.27;2.39,0.62,0.32,1.22.
Malegenitaliasubfusiform, 1.65mm in length and 0.3mm inwidth, feebly
curvedinlateralviewwithbasalpiecesubovateanddistinctlywiderthanlaterallobes;
fused lateral lobeselongated triangular, 0.7mmin lengthwithrounded, somewhat
spatulateapices.
Bodylength:7.7mm.
Holotype: , “SUMATRA/SIBOGA/X. 90 e 111. 91/E. MODIGLIANI//
MUSEUMPARIS/1952/COLL.R.0BERTUR”(MNHNP).Paratypes:1ex.,samedata
as fortheholotype; l ex.,“SUMATRA,MEDAN/Env.DeDo1ok-Baros/2esemestre
1905”.
Notes. This new speciessomewhat resemblesStrongylzum diverslco1or PIC,
1940, fromTonkin,butcanbedistinguished fromthelatterbythelargerandwider
body,withdifferentcoloration(whollydarkgreenishinSdiverslco1or).
Strongyliumnagauspnov.
(Figs2,13-14)
Blackishbrown,posteriorpartofheadandpronotumwithdarkgreenishlustre,
scutellumchocolatebrown,elytrapurplishwith feeblegolden lustre; dorsalsurface
weakly,sericeouslyshining;ventralsurfaceandlegsmoderately,slightlyalutaceously
shjnjng; almost glabrous.Elongated fusiform, longitudinally convex, thoughgently
flattenedinbasalpartsofelytra.
Head subdecagona1, feebly covered with isodiametric microsculpture, closely,
finelypunctate;clypeusrathertransverse,flattenedinbasalpart,bentventradinapical
part, fronto_clypea1bordergentlycurved;genaeratherstronglyraisedoutwardswith
roundedoutermargins; fronsratherwide,gently inclinedanteria diatone 0.9 times
thewjdthoftransversediameterofaneye,withaweakimpressionat themiddleof
posterjorpart,andalsowithanmi punctateareaatthemiddleclosetofronto-Clypea1
border.Eyesmedium-size convexlaterad,obliquelyinlaidbygenaeatthemiddleof
anterjormargjns.Antennaereachinghumeriofelytra,withfiveapicalsegmentsna一「
rowedateachbase,ratioofthelengthofeachsegment frombasetoapex:0.39,0・2,
0.72,0.62,0.43,0.36,0.26,0.27,0.24,0.23,0.32.
pronotumwjdelyhexagonal,1.3timesaswideas long;apexsublinear, finely
rjmmedoneachsjde;baseslightlysinuousoneachside,borderedandweaklyridged
wjdelyjnmjddle;sidesrathersteeplydeclinedtolateralmargins,whicha「eenti「ely
finelyrimmed,eachwithcrenulationsinmedialpart;frontan9lesroundedthou9hfee-
bly produced, hjndangles obtusely angulate; disc gently convex, weakly, Va9uely
obljquely impressedinbase-lateralparts,andalsomoreclearly imP「eSSedCloseto
baseoneachsjde,coveredwithisodiametricmicrosculpture,rathercloselypunctate,
404 KimioMAsUMoTo
1・ 3
一
/.・
.・・・・ 1・・一/
f4、 5 6
Figs. 1-6. HabitusofSI,・o,tgyliumspp fromEastAsia. - l,Smodig/!an!1sp nov.,holotype, ;2,s
'1agatlSp nov.,holotype, ;3,Snolsuispnov.,holotype,3;4,S.walshaespnov.,holotype,6;5,s
';1u1otispnov.,holotype,3;6,S.sl,-a,nbe,1sespnov.,holotype,
.
thepuncturessparselyintermixedwithsmallerpunctures.Scutellumsubcordate,gent_
1yelevatedposteriad,weaklyridgedlongitudinallyatthemiddle, feeblycoveredwjth
isodiametricmicrosculpture,sparselyscatteredwithmicroscopicpunctures.
Elytraelongatedsubovate,2.1timesaslongaswide,3.83timesthelengthand
StudyofAsianStrongyliini,XI
405
l
、
、
/
7 8
、 10
Figs 7-10. HabitusOfSt''ongylitt'nspp. - 7,S. /ttoe''ta川″11sp nov.,hOlotype, ; 8,S tsllytlkie//urn
spnov.,holotype, ;9,Snaga1e/1u,nspnov.,holotype,(5;10,Sdo/oke,1sesp nov.,holotype,3.
1.22timesthewidthofpronotum;dorsumlongitudinallyconvex,weaklydepressedin
interiorpartsofbasal t/5; discpunctategrooved, 5thgroovedeepenedclose tobase,
thepuncturessmall,rathercloselyset,becomingfinerposteriad; intervalsgentlycon-
vexinmajoranteriorparts, becominglowerposteriad,weaklycoveredwithisodiamet-
ricmicrosculpture,sparselyscatteredwithmicroscopicpunctures, feebly,rathertrans-
406 KimioMAsUMoT0
verselywrinkled;humerigentlyswollen;apicesroundlyproduced.
Maleanal sternite rather weakly,subelliptically depressed inapical part. Legs
ratherslender;malemetatibiaewithinterior facesweaklygougedinbasalparts;ratios
of thelengthsofpro-,meso- andmetatarsomeres: 0.34, 0.31, 0.32, 0.28, 1.2; 1.73,
0.89,0.73,0.41,1.29;2.21,0.91,0.49,129.
.
Malegenitaliashort fusiform,1.3mmin length,0.28mm inwidth,withbasal
partcurvedinlateralview; fusedlateral lobesratherelongatedtriangular,0.75mmin
length,withroundedapices.
Bodylength:8.6mm.
Holotype: ,nearKeningau,Sabah,N.Borneo,Malaysia,14-III-l988,M. ITO
leg. (NSMT). Paratypes: 1 ex., Kimanis Road, near Keningau, Sabah, Borneo,
5-V-1994; 1 ex., Crocker Range, 1,400m alt., NW of Keningau, Sabah, Borneo,
2~14-V-1984,S.NAGAl leg;1ex.,Keningau,Sabah,N.Borneo,11-V-1989,M.ITO
leg;2exs.,Keningau,N.Borneo,29-V-1989,M. ITOleg;1ex.,“SABAH:Crocker/
RangeNP,Gn.Emas/6.-18.VI.1996, l600m/leg.J.KoDADA”(SMNS).
Notes. This new species resemblesthepreceding species,Slrongyliummod1-
glian11sp nov., inhaving characteristicantennaeandsomeothermajor features,so
thatthesetwospeciesformaspecies-group.Itcanbedistinguishedfromthepreceding
speciesbythedorsalsurfacerathercoveredwithisodiametricmicrosculpture, lateral
marginsofpronotumcrenulateinmiddle,andtheelytrapunctateandwrinkled.
Stron Z加mnofsM'sp
nov.
(Figs3,15-16)
Darkreddishbrown,majorposteriorpartofheadandpronotumexcept forbasal
partwithdarkbluishlustre,basalpartofpronotum,scutellumandelytrawithcoppery,
partlyslightlygoldenlustre,mouthpartsandclawslighter incolour;head,pronotum
andscutellumsericeouslyshining,elytraratherstronglyshining,ventralsurfacesome-
whatalutaceouslyshining;eachsurfacealmostglabrous.Ratherelongate, longitudi-
nallyconvex,thoughweaklydepressedatbasal t/4ofelytra.
Headnearly rounde gentlyconvexinmiddle,weaklyimpressedmedially,cov-
eredwith isodiametricmicrosculpture,closely, finelypunctate;clypeussemicircular,
gently flattenedinbasalpart,bentventradinmiddle, truncateatapex,withashort
transverseimpressionbefore fronto-clypeaIborder;genaesomewhatobliquely rhom_
bic,raisedoutwards,withobtuseoutermargins; fronswidelyT-shaped,rathersteeply
inclined anteria diatone0.67timesthewidthoftransversediameterofaneye.Eyes
medium-size subreniform,convexlatera obliquelyinlaidintohead.Antennaemiss_
ingfourapicalsegmentsinthetypespecimen: ratioofthelengthofeachsegment
frombaseto7thsegment:0.38,0.2,0.6,0.49,0.41,0.39,0.31
Pronotumsomewhattransverselyhexagonal,1.33timesaswideaslong;apexa_l
moststraight,finelyborderedandrimmedoneachside;baseslightlysinuousoneach
side,clearlyborderedand rimmed, therimbecomingbolderwidely inmiddle;sides
StudyofAsianStrongyliini,XI 407
steeplydeclinedtolateralmargins,whichareseparatedfromventralpartsbyfinerims,
roundlyproducedinmiddleandfeeblysinuousbeforebaseindorsalview;frontangles
nearlyrounde hindanglessubrectangular;discgentlyconvex,coveredwithisodia-
metricmicrosculpture,moderatelycloselypunctate, thepunctureslargerandsparser
thanthoseonhead.Scutellumtriangular,gentlyconvex,coveredwithisodiametricmi_
crosculpture,sparselyscatteredwithveryfinepunctures.
Elytraelongatedsubfusiform,2.31timesaslongaswide,4.5timesthelengthand
1.5timesthewidthofpronotum;dorsumlongitudinallyconvex, thoughweakly flat-
tenedinareasaroundbasal t/5andalongscutellarstricto;discpunctategrooved,5th
groovetouchingbase,thepunctureslongitudinalanddeep,becomingsmallerandshal-
lowerposteriad;intervalsgentlyconvex,flattenedposteriad,coveredwithisodiametric
microsculpture,verysparsely scatteredwithmicroscopicpunctures,whicharevery
sparselyintermixedwithlargerpunctures;humeriweaklyswollen;apicesslightlypro-
duced.
Male anal sternite without any modification, covered with isodiametric mi-
crosculptureandmicroscopicallypunctate. Legsratherslender;malemetatibiaewith
interior faceweakly gougedat basal t/4; ratiosof the lengths of pro-, meso- and
metatarsomeres: 0.35, 0.33, 0.31, 0.27, 1.2; 1.28, 0.79, 0.67, 0.39, 1.27; 1.91, 0.89,
0.41 1.22.
Malegenitaliafusiform,0.85mminlengthand0.2mminwidth,weaklycurved
inlateralview; fusedlateral lobessomewhattriangular,0.35mminlength, feeblyde-
pressedlongitudinallyinposteriorpart,scatteredwithmicroscopicandsomewhatlon-
gitudinalpuncturesinapicalhalves.
Bodylength:6.3mm.
Holotype: (5, Sabah, Borneo, 14-IV~19-V-1984, S. NAGAl leg. (NSMT).
Paratypes: 1ex.,samedataas for theholotype; 3exs.,16milesNWofKeningau,
1,400malt., Borneo, Malaysia, 12~19-Vm-1983, Y. NoTsu leg; 1ex., Crocker
Range,1,400malt.,NWofKeningau,Sabah,Borneo, l2-V-1984,S.NAGAl leg;1
ex.,Keningau,Sabah,N.Borneo,10~20-X-1988,M.ITOleg.
Motes. Thisnewspeciessomewhat resemblesStrongyfzummodzgfzamfsp nov.,
butcanbedistinguishedfromthelatterbythesmallerandslendererbody,withelytra
moredeeplypunctate-striate.
Strongyliumwalshaesp nov
(Figs4,17-18)
Yellowishbrown,posteriorpart ofheadandelytraexcept formedic-basalpart
brownishblackwithdarkgreenishorbluelustre,apicalmarginandapico-media1part
of pronotum brownish black, scutellum and medic-basal part of elytra chocolate
brown; head and pronotum weakly, rathersericeously shining,scutellumvitreously
shining,elytra feeblysericeously,somewhatmetallicallyshining,ventralsurfaceand
legsrather alutaceouslyshining;eachsurfacealmost glabrous.Elongated fusiform,
408 Kimio MAsUMoTo
longitudinallyconvex,thoughweaklyflattenedinmedialpartsofelytra.
Headtransverselyelliptical,coveredwithisodiametricmicrosculpture,closelyru-
gose-punctate;clypeussemicircular, feeblydepressedinbasalpart,gradually inclined
forwards,gentlybentandtruncateinfront,fronto-clypea1bordergentlycurvedandnot
clearly impressed;genaeobliquelysubrhombic, ratherstronglyraisedoutwards; frons
rather finelyT-shaped,steeply inclinedanteriad,diatone1/8timesthewidthoftrans_
versediameterofaneye,withalongitudinalimpressionatthemiddleofposteriorpart,
weaklysulcatealongthebordersofeyes.Eyesnoticeablylarge,subreniform,strongly
convex latera approximatewitheachother.Antennaeratherfiliform,reachingbasal
1/3ofelytra,ratioofthelengthofeachsegment frombasetoapex:0.64,0.2,0.81,
0.79,0.61,0.63,0.62,0.6,0.58,0.59,0.63.
Pronotum widely subhexagona1, 1.33 timesaswideas long; apex sublinear,
rimme therimthickenedinmiddle;baseslightlysinuousoneachside,clearlybor_
deredandrimmed;sidessteeplydeclinedtolateralmargins,whicharenot defjned
fromventralparts,producedinmiddle,andgentlysinuousbeforethebaseindorsal
view; frontanglesobtuse,hindanglessubrectangular;discgentlyconvex,weaklyde_
Pressedinmedic-longitudinalpartanddeepenedinapicalandbasalparts,weaklyim_
pressedclosetobaseoneachside,veryfeeblycoveredwithisodiametricmicrosculp_
ture, rather closely punctate, thepuncturesoften fusedwithoneanother.Scutellum
Slightlyelongatedtriangular,feeblyconvex,almostsmooth,sparselyscatteredwjthmj_
croscopicpunctures.
Elytraelongatedovate,2.18timesas longaswide,4timesthelengthand
12
timesthewidthofpronotum;dorsumlongitudinallyconvex,though
attenedjnmedja1
Pa「t;discwithrowsofpunctures,5throwclosetobase,thepunctureslargeandtrans_
Ve「Se,Closelyset, oftensomewhathexagonal,becomingsmallerposterjad; jntervals
na「「owandweaklyconvex,ratherridgedinlateralparts,weaklycoveredwjthjsodja_
met「ICmiCrosculpture,sparselyscatteredwithmicroscopicpunctures,wjth2ndjnter_
valflattened;humeriweaklyswollen;apicesroundlyproduced.
Maleanalstemitesemicircularlydepressedandpubescentinapjca1part Legs
medium-Sizedforamemberofthisgenus,malemetatibiaeflattenedininterior facesof
basalhalves,weaklytwisted,withdorso-interna1edgesroundlyproduced;ratjosofthe
fen9thSofp「o-,meso-andmetatarsomeres:0.32,0.23,0.25,0.22,1.2;1.g6,0.g5,0.77,
0.44,1.32;2.03,0.69,0.48,1.28.
Male9enitaliasubfusiform,1.5mm in lengthand0.3mm inwjdth,aml ost
St「al9htinlateralview; fusedlaterallobeselongatedtriangular,0.7mmjnlenglhwith
slightlyprolongedratheracuteapices.
Bodylength:6.32mm.
Holotype: “SUMATRA/Reg de Benkoelen/Moeara Tenam /Mme M E
,
WALSH/1935”(MNHNP).
Notes. ThisnewspeciesresemblesStrongyliumbenakatenseMAsUMoTo,1gg7,
f「om SouthSumatrainhavingmodifiedmalemetatibiae,butcanbedjstjngujshed
f「omthelatterbythedifferentlycoloredbody,withlargereyes,thelongerelytra,and
StudyofAsianStrongyliini,XI 409
theantennaeandlegswithdifferentratiosofsegmentlengths
Strongyliummulotisp nov.
(Figs 5,19-20)
Yellowishbrown, antennae, apical halvesof tibiaeandmetepisternadarker in
colour, head and pronotum blackish brown withweak dark bluish lustre, elytra in
majorpartswithgreenishbluelustre,thoseinantero-media1parts feeblywithgolden
greenishlustre,andthoseinpostero-1atera1partsfaintlywithpurplelustre;headand
pronotumweakly,sericeouslyshining,scutellumvitreouslyshining,elytrametallically
andsomewhatsericeouslyshining, ventral surface rather alutaceouslyshining, legs
moderately shining;eachsurfacealmost glabrous. Elongated fusiform, longitudinally
convex,thoughweakly attenedinantero-media1partsofelytra.
Headtransverselysube11iptica1,weaklycoveredwithisodiametricmicrosculpture,
closely, finelypunctate;clypeussemicircular,gentlyinclinedapica truncatein front,
fronto-clypealborder curvedandfinely impressed;genaeremarkably oblique, raised
posteriadandoutwards,withobtuseoutermargins; fronsratherT-shape steeply in-
clined anteria diatone1/9 timesthewidthoftransversediameterof aneye, witha
medic-longitudinalimpunctateimpressionat theleveloftheposteriorpartsofeyes.
Eyes large,subreniform, convex latera obliquely inlaid intohead. Antennaerather
filiform,reachingbasal t/4ofelytra,ratioofthelengthofeachsegment frombaseto
apex:0.78,0.2,0.79,0.72,0.56,0.55,0.53,0.56,0.51,0.53,0.59.
Pronotumshortbarrel-shape 1.21timesaswideaslong;apexveryweaklypro-
duce borderedand rimme therimmicroscopicallypunctateand feeblythickenedin
middle;baseslightlysinuousoneachside,clearlyborderedandrimmed,therimno-
ticeablysmoothinmiddle;sidessteeplydeclinedtolateralmargins,whicharenotde-
fined fromventralparts,roundedinmiddleandgentlysinuousbeforebaseindorsal
view; frontanglesobtuse,hindanglessubrectangular;discgentlyconvex,weaklyde-
pressedinmedic-basalpart,veryweaklyimpressedclosetobaseoneachside,weakly
coveredwithisodiametricmicrosculpture,closelypunctate, thepuncturessometimes
fusedwithoneanother. Scutellumtriangular with rounded sides, almost smooth,
sparselyscatteredwithmicroscopicpunctures,sparselysomewhatlongitudinallyacic-
ulate.
Elytraelongatedsubfusiform,2.04timesaslongaswide,3.8timesthelengthand
1.4 timesthewidthofpronotum;dorsumlongitudinallyconvex, though attened in
medic_longitudinalpart;discwithrowsofpunctures,thepunctureslarge,transverse
andclosely set,sometimes rather hexagonal, becoming smaller posteriad; intervals
narrowandconvex,ridgedinlateralparts,with2ndinterval attened,weaklycovered
wjth isodiametric microsculpture, sparsely scattered with microscopic punCtu「eS;
humerinotsodistinct;apicesroundlyproduced.
Maleanalsterniteweaklydepressedandpubescent inapicalpart. Legsmedium-
sized foramemberofthisgenus,maleprotibiaecurvedventrad;malemetatibiae at-
410 KimioMAsUMoT0
tonedininterior facesofbasalhalves,weaklytwistecLwiththedorsaledgesroundly
produced; ratiosofthelengthsofpro-,meso-andmetatarsomeres: 0.27,0.22,0.24,
0.26,12;2.05,0.96,0.82,0.6,1.42;2.18,0.86,0.59,1.37.
.
Male genitalia subfusiform, 1.7mm in length and 0.3mm in width, weakly
curvedinlateralview; fusedlateral lobeselongatedtriangular,0.75mminlength,de-
pressedmedic-longitudinally,withslightlyprolonged,ratheracuteapices.
Bodylength:5.5-6.5mm.
Holotype: “Borneoocc./Goenong Ampar/Mu1ot 1897//Museum Paris/ex
,
Coll./R.0BERTHUR”(MNHNP).Paratypes:1ex.,“BorneoOcc./Pontianak/1901”;3
exs.,“EstBorneo/Batanbessi/MeM.E.WALsH/1937';1ex.,Keningau,Sabah,7-
V-1981,M.TAOleg;1ex.,nr.Kpg.Poring,Sabah,BorneoIs.,Malaysia,4-I-1985,
M. NlsHIKAwA leg; 2exs., BORNEO: SABAH/Bingkor N Keningau/400-500m,
19.20.XI i996/leg.W.SCHAWALLER”(SMNS).
Notes. ThisnewspeciescloselyresemblesStrongyliumbenakatenseMAsUMoTo
etMAKIHARA,1997, fromSouthSumatrainhavingmodifiedmalemetatibiae,butcan
bedistinguished fromthe latterby thedifferently coloredbody, witheyes largerand
closertoeachother,pronotummoreconvexlateradandelytrashorter.
Thisandtheprecedingnewspeciesarethemembersofthespecies-groupofS.
benakatense,whosemalesarecharacterizedbymodifiedlegs.
Strongyliumsirambe'tsesp nov.
(Figs6,21-22)
Darkyellowishbrown,elytra,basalpartsofantennaeandtibiaelighter incolour,
dorsalsurfacemoderately,vitreouslyshining,ventralsurfacesomewhatalutaceously
shining,sixapicalsegmentsofantennaenotshining; eachsurfacealmost glabrous.
Elongated fusiform, longitudinallyconvex, thoughweakly attened in antero-media1
partsofelytra.
Head transversely sube11iptica1, very weakly micro-shagreen closely, finely
punctate,thepunctureslightlyfusedwithoneanother inareabetweenposteriorparts
ofeyes;clypeussemicircular, rathersteeply inclinedapicad,truncatein front, fronto-
clypea1bordercurvedandratherstronglyimpressed;genaeoblique,raisedposteriad
andoutwards,withobtuseoutermargins; fronsfinelyT-shape steeply inclinedante_
riad,diatone1/12timesthewidthoftransversediameterofaneye,withanindistinct
impressionat themiddleofposteriorpartofeyes.Eyeslarge,approximatetoeach
other,convexlaterad,ratherbroadly inlaidintohead.Antennaesubfiliform,reachjng
basal t/3ofelytra,ratioofthelengthofeachsegment frombasetoapex:0.41,02,
0.59,0.45,0.41,0.6,0.59,0.62,0.61,0.60,0.79.
Pronotumsubquadrate,1.3timesaswideaslong;apexalmoststraight,entirely
bordered and rimme therimsparselyscatteredwithmicroscopicpunctures; base
slightlysinuousoneachside,clearlyborderedandrimme therimbolderthanthat of
apexandalmostimpunctate;sidessteeplydeclinedtolateralmargins,whicharesepa_