Studies of Tiger Beetles. CLV. New data from Thailand and Cambodia (Coleoptera: Cicindelidae)

2005·1.7 MB·

by  Cassola, F

BULLETfN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE ENTOMOLOGIE, 75: 13-21,2005
BULLETIN VAN HET KONfNKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN ENTOMO!'.OGIE, 75: 13-21,2005
Studies of Tiger Beetles. CLV . New data from Thailand and Cambodia
(Coleoptera: Cicindelidae)
by Fabio CASSOLA
Abstract List of species
Thirty-seven tiger beetle species are here examined which were col 1. Tricondyla annulicornis SCHI\'IIDT-GOEBEL, 1846
lected in Thailand and Cambodia during several recent missions of the THAILAND, Loei: Na Haeo, 21-29.IV.00, Malaise trap,
Royal Belgian Institute of Natural Sciences (RBINS). Seven species o
P. Grootaert, I (RBINS).
are newly recorded from Cambodia, and moreover a new species,
Cylindera (lfasina) khmer, is also described from Cambodia. Neocol Described as a distinct species (SCHMIDT-GOEBEL,
lyris (Neocollyris) vitalisi (W. HORN, 1924) is tentatively raised to full 1846) but for long time considered to be a subspecies
speci fie status. of T. cyanea Dejean, 1825 (WIESNER, 1992), this species
has been reinstated in its full specific status by NAVIAUX
(2002). It is also kt10wn from Cambodia (SAWADA &
Resume
WIESNER, 1999).
Les 37 especes de Cicindelides sont etudiees ici qui ont ete recueillies
lors de plusieures missions recemment menees en Thailande et Cam 2. Protocollyris probsti NAVIAUX, 1994
badge par I'Institut Royal des Sciences Naturelles de Belgique CAMBODIA, Siem Reap: Phnom Kulen, 24.V.03, light trap,
(IRSNB). Sept especes sont nouvellement indiquees du Cambodge, J. Constant & K. Smets, I ~ (RBINS).
et une nouvelle espece, Cylindera (lfasina) khmer, est aussi decrite Described from Thailand and Laos (NAVIAUX, 1995),
du Cambodge. Neocollyris (Neocollyris) vitalisi (W. HORN, 1924) est
tentativement elevee au rang d'espece propre. new to Cambodia.
Key words: Coleoptera, Cicindelidae, Thailand, Cambodia.
3. Neocol/yris (Neocollyris) b. bonellii (GUERIN-MENE-
VILLE, 1834)
THAILAND, Loei: Na Haeo (bio. station), 5-l2.V.OI, secondary
Introduction
forest, Constant & Grootaert, 1 (5 (RBINS); Na Haeo, 22-
o
29.V.03, Malaise trap 4, P. Grootaert, I I ~ (RBINS). Ra
Due probably to heavy long-lasting wars and the subse nong: Koh Samed Island, 15-17. V. 0 I, Constant & Grootae11,
quent difficult political conditions, Cambodia is still a I ~ (FCC).
poorly explored countly, as far as its entomological fauna CAMBODIA, Siem Reap: Angkor Thom, 23.V.03, sweeping,
is concerned. Apatt from Walther Horn's preliminary J. Constant, K. Smets & P.Grootaert, 1 0 (RBINS); Prek Toal
lists of species, which were related to the whole of Indo (Tonie Sap Lake), 27-28.V.03, sweeping, J. Constant, K. Smets
oo
china (HoRN, 1913, 1914), only 44 tiger beetle species & P. Grootaert, 2 (RBINS); 8 km N of Sre Noi (road to
have been formally recorded from this countty so far Anlong Vaeng), 29-30.V.03, dry dipteroc. forest, day, J. Con
o
(Table 1). Future research will certainly add some more stant & K. Smets, I (FCC), I ~ (RBINS).
A common widespread species (NAVIAUX, 1995), how
species to this list. Thailand, in contrast, is a much better
ever not yet formally recorded from Cambodia.
known count1y, thanks especially to the dedicated re
search efforts by NAVIAUX (1989, 199la, b, 1995,
1996a, b, 2002, 2004) and NAVIAUX & PINRATANA
4. Neocollyris (Neocollyris) moesta (SCHMIDT-GOEBEL,
(2004). My own reckoning for Thailand presently indi
1846)
cates a total of 133 tiger beetle species (with 25 or 18.7%
THAILAND, Loei: Na Haeo, 22-29.V.03, Malaise trap 4,
endemics), what definitely ranges Thailand among the o
P. Grootaert, I (RBINS).
countries of the world with the richest cicindelid biodi CAMBODIA, Siem Reap: 8 km N of Sre Noi (road to Anlong
versity (PEARSON & CASSOLA, 1992; CASSOLA & PEAR Vaeng), 29-30.V.03, dry dipterocarp forest, day, J. Constant &
SON, 2000). More precisely, Thailand has turned out to be K. Smets, 2 ~ (RBINS); Angkor Thom, 2.VI-31.VIL03, net
o
the I 0111 country of the world as far as the total number of catching, IG 30.192, Daniel R. Jump, I (RBINS).
tiger beetle species is concerned, and even the 6111 as to the Neither WIESNER (1992) nor NAVIAUX ( 1995) men
surface or kt112/species ratio (3,848). tioned this easily recognizable species from Cambodia.
14 Fabio CASSOLA
Table I Checklist of the tiger beetles (Coleoptera: Cicindelidae) of Cambodia.
Species References
I. Tricondyla (Tricondyla) annulicornis Schmidt- Goebel, 1846 WIESNER, 1992; SAWADA & WIESNER, 1999
2. Tricondyla (Tricondyla) stricticeps Chaudoir, 1864 SAWADA & WIESNER, 1999; NAVIAUX, 2002
3. Protocollyris probsti Naviaux. 1994 CASSOLA, this paper
4. Neocollyris (Neocollyris) b. bonellii (Guerin-M., 1834) CASSOLA, this paper
5. Neocollyris (Neocollyris) moesta (Schmidt-Goebel, 1846) HORN, 1913; CASSOLA, this paper
6. Neocollyris (Neocollyris) batesi (W. Horn, 1892) WIESNER, 1992
7. Neocollyris (Neocollyris) jitscitarsis (Schmidt-Goebel, 1846) CASSOLA, this paper
8. Neocollyris (Neocollyris) rujipalpis (Chaudoir, 1864) HORN, 1914; WIESNER, 1992; SAWADA & WIESNER, 1999
9. Neocollyris (Orthocollyris) crassicornis (Dejean, 1825) FLEUTIAUX, 1902; NAVIAUX, 1995 ("presque tout le territoire")
I 0. Neocollyris (Leptocollyris) !. linearis (Schmidt-Goebel, 1846) WIESNER, 1992; SAWADA & WIESNER, 1999
II. Neocollyris (Leptocollyris) cylindripennis (Chaudoir, 1864) WIESNER, 1992 (sub cochinchinensis)
12. Neocollyris (Leptocollyris) 1. linearis (Schmidt-Goebel, 1846) WIESNER, 1992; CASSOLA, this paper
13. Neocollyris (Paclzycollyris) fascia/a (Chaudoir, 1864) WIESNER, 1992; NAVIAUX, 1995
14. Prothyma (Paraprothyma) exornata Schmidt-Goebel, 1846 I-lORN, 1913; RIVALIER, 1964; WIESNER, 1992;
SAWADA & WIESNER, 1999
15. Prothyma (Genoprothyma) heteromalla (Macleay, 1825) RIVALIER, E., 1964 ("dans toute l'lndochine"); CASSOLA, this paper
16. Prothyma (Genoprothyma) bouvieri W. Horn, 1896 WIESNER, 1992
17. I-feptodonta eugenia Chaudoir, 1865 WIESNER, 1992; SAWADA & WiESNER, 1999
18. Calochroa .f jlavomaculata (Hope, 1831) HORN, 1913 (sub se.xpunc/ala); WIESNER, 1992;
SAWADA & WIESNER, 1999
19. Calochroa 1nouhotii (Chaudoir, 1865) HORN, 1913; RIVALIER, 1961; WIESNER, 1992;
SAWADA & WIESNER, 1999
20. Calochroa elegantula (Dokhtouroff, 1882) WIESNER, 1992 (sub mouhoti elegan!llla);
SAWADA & WIESNER, 1999 (sub mouhoti elegantula)
21. Calochroa interruptofasciata (Schmidt-Goebel, 1846) I-lORN, 1913; RIVALIER, 1961; WIESNER, 1992;
SAWADA & WIESNER, 1999
22. Calochroa harmandi (Fieutiaux, 1893) I-lORN, 1913; RIVALIER, 1961; WIESNER, 1992
23. Calochroa brwnani (Dokhtouroff, 1882) WIESNER, 1992 ; SAWADA & WIESNER, 1999; CASSOLA, this paper
24. Calome1·a plumigera (W. H.) ssp. scoliographa (Rivalier, 1953) RIVALIER, 1953; WIESNER, 1992;
SAWADA & WIESNER, 1999 (sub p. macrograplina)
25. Cosmodela aurulenta (Fabr.) ssp. juxta/a (Acciavatti & Pearson, HORN, 1913; RIVALIER, 1961 (sub a.jlavomaculata);
1889) WiESNER, 1992; SAWADA & WIESNER, 1999
26. Cosmodela virgula (Fleutiaux, 1893) HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999
27. Lophyra (Lophyra) fit!iginosa (Dejean, 1826) HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999
28. Lophyra (Lophyra) cancel/ala (Dejean, 1825) RIVALIER, 1961; WIESNER, 1992; SAWADA & WIESNER, 1999
29. Lophyra (Spilodia) striolata (llliger, 1800) HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999
30. Lophyra (Spilodia) lineiji·ons (Chaudoir, 1865) RIVALIER, 1961; WIESNER, 1992; SAWADA & WIESNER, 1999
31. Naviauxella ramai Naviaux, 1991 CASSOLA, this paper
32. Cylindera (/fasina) foveola/a (Schaum, 1863) CASSOLA, this paper
33. Cylindera (lfasina) viduata (Fabricius, 180 I) WIESNER, 1992; SAWADA & WIESNER, 1999; CASSOLA, this paper
34. Cylindera (lfasina) khmer n. sp. * CASSOLA, this paper
35. Cylindera (/fasina) decempunctala (Dejean, 1825) HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999
36. Cylindera (/fasina) discreta (Schaum, 1863) (I) HORN, 1913; WIESNER, 1992
37. Cylindera (Eugrapha) minuta (Olivier, 1790) RIVALIER, 1961 ("majeure partie du territoire");
CASSOLA, this paper
38. Cylindera (Eugrapha) venosa (Kollar, 1836) RIVALIER, 1961; WIESNER, 1992
39. Myriochila (Myriochila) sinica (Fleutiaux, 1889) WIESNER, 1992; SAWADA & WIESNER, 1999
40. Myriochila (Myriochila) specularis (Chaudoir, 1865) HoRN, 1913 (sub speculifera); RIVALIER, 1961;
WIESNER, 1992 (sub speculifera)
41. Abroscelis tenuipes (Dejean, 1826) WIESNER, 1992
42. Callytron nivicinctum (Chevrolat, 1845) WIESNER, 1992
43. Callytron andersoni (Gestro, 1889) WIESNER, 1992; SAWADA & WIESNER, 1999
44. Enantiola hewittii (W. Horn, 1908) WIESNER, 1992
Excluded species:
Tricondyla pulchripes White, 1844 HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999;
NA VIAUX, 2002
Tricondyla gestroi Fleutiaux, 1893 HORN, 1913; WIESNER, 1992; SAWADA & WIESNER, 1999;
NA VIAUX, 2002
Neocollyris (Neocollyris) stiengensis (W. Horn, 1914) WIESNER, 1992; NA VIAUX, 1995
Neocollyris (Neocollyris) rugosa (Chaudoir, 1864) WIESNER, 1992; NAVIAUX, 1995
Neocollyris (Neocollyris) impressifi"Oizs (Chaudoir, 1864) WIESNER, 1992; NAVIAUX, 1995
Calomera a. angulata (Fabricius, 1798) I-lORN, 1913; RIVALIER, 1961, sub Lophyridia sumatrensis
("presque tout le territoire")
"' Endemics: I
(II This record has probably to be referred to a different species of the discre/a-group,juergellwies/leri Naviaux. 1991, known from Thailand. The true discrela occurs in
Sulawesi only (CASSOLA, 1991) and, with different subspecies. in other Indonesian islands and in the Philippines.
Tiger beetles from Thailand and Cambodia 15
However, it had been recorded from such a country by I 0 (RBINS); Na Haeo (bio. station), 5-12.Y.OI , secondary
HoRN (1913). The new data cited above clearly confirm forest, Constant & Grootaert, I (RBINS); Dan Sai: Bantun,
HORN's record. ll.V.OI, Constant & Grootaert, I ~ (RBINS); Na Haeo field
res. sta., 15-19.V.03, day catch, Constant & K. Smets,
5. Neocollyris (Neocollyris) batesi (W. HoRN, 1892) I 0 (RBINS), I ~ (FCC); Na Haeo, Chang Tok, waterfall,
18. Y.03, day catch, K. Smets, I 6 (RBINS). Kanchanaburi:
THAILAND, Ranong: Koh Samed Island, 15-17.V.O I, Constant & Sai Yok N.P., 4-5.Vl.03, J. Constant & K. Smets, I 6 (RBINS),
Grootaert, I ~ (RBINS). I 6 (FCC).
Already recorded from Thailand by WIESNER ( 1992)
Also a small widespread species, particularly common
and NAVIAUX (1995). Known also from Cambodia
in northern Thailand (NAVIAUX, 1995). The particular
(WIESNER, 1992).
shape of the pronotum makes it easily recognizable, but
there is apparently some variability
6. Neocollyris (Neocollyris) intermedia NAVIAUX, 1994
THAILAND, Loei: Na Haeo, Station 20023, 25.V.OO, P. Groo
taert, I ~ (RBINS). 12. Neocollyris (Leptocollyris) subti/is (CHAUD.) ssp.
Described at first as a subspecies of N. (N.) orichalcina brachycephala (W. HoRN, 1893)
(W. HORN, 1896) (NAVIAUX, 1995), intennedia has been THAILAND, Loei: Na Haeo, Chang Tok, 17.V.03 (23035),
subsequently raised to full specific status (NAVIAUX, P. Grootaert, I ~ (RBINS).
2004). Described from "Carin Cheba" (Myanmar), this
subspecies replaces s. subtilis in northern Thailand
7. Neocollyris (Neocollyris) siamensis Naviaux, 1991 (NAVIAUX, 1995).
THAILAND, Kanchanaburi: Sai Yok N.P., 4-5.Yl.03, J. Constant
& K. Smets, I ~ (RBINS), I ~ (FCC).
13. Neocollyris (Leptocollyris) variitarsis (CHAUDOIR,
Described from the Khao Yai Nat. Park in the Nakhon
1860)
Ratchasima province (NAVIAUX, 1991 b), this species is
THAILAND, Loei: Na Haeo (bio. station), 5-12.V.OI, secondmy '
said to be fairly common in the whole of Thailand, except
forest, Constant & Grootaert, 3 ~~ (RBINS); Na Haeo,
for the north-western provinces (NAVIAUX, 1995).
29.Y-5.Yl.03, Malaise trap 4, P. Grootaert, I ~ (FCC); Na Haeo
field res. sta., 15-19.V.03, day catch, Constant & K. Smets, I 6
8. Neocollyris (Neocol/yris) jitscitarsis (SCHMIDT- (RBINS).
GOEBEL, 1846) A widespread species, occtm·ing from Eastem Nepal to
THAILAND, Loei: Na Haeo (bio. station), 5-12.Y.O I, secondary Vietnam and southern China (NAVIAUX, 1995).
forest, Constant & Grootaert, I 0 I ~ (RBINS); Na Haeo (field
res. st.), 15-19.Y.03, day catch, Constant & K. Smets, I 6
(RBINS). 14. Neocollyris (Pachycol/yris) vitalisi (W. HORN,
CAMBODIA, Siem Reap: 8 km N of Sre Noi (road to Anlong 1924)
Vaeng), 29-30.V.03, dry dipteroc. forest, day, J. Constant & THAILAND, Loei: Na Haeo (bio. station), 5-12.V.Ol, seconda1y
K. Smets, 1 6 I ~ (RBINS). forest, Constant & Grootaert, I ~ (RBINS).
A widespread easily recognizable species, already known Described as a subspecies of N. (P.) .feae (W. HORN,
from Thailand (NAVIAUX, 1995) but new to Cambodia. 1893), but quite distinct because of the smaller size, the
darker colour and its geographical distribution, vitalisi is
9. Neocollyris (Neocollyris) rufipalpis (CHAUDOIR, herein tentatively raised to full specific status.
1864)
THAILAND, Loei: Na Haeo, river banks, 5-12.V.OI, Constant &
Grootaert, I 6 (RBINS). 15. Prothyma (Paraprothyma) schmidtgoebeli W. HORN,
Already known from Thailand (NAVIAUX, 1995), this 1895
species has also been recorded from Cambodia and Viet THAILAND, Loei: Na Haeo, Station 20023, 25.Y.OO, P. Groo
taert, I 6 (RBfNS), I 6 (FCC); Na Haeo, 29.IY-6.Y.OI,
nam (SAWADA & WIESNER, 1999).
Constant & Grootaert, I 6 I <jl (RBINS); Na Haeo (bio station),
5-12.V.Ol, Malaise trap, P. Grootaert, I ~ (RBINS); Na Haeo
I 0. Neocollyris (Leptocollyris) I. linearis (SCHMIDT- (bio station), 5-12.V.Ol, light trap, Constant & Grootaert, 6 66
GOEBEL, 1846) 5 n (IRSNB), I 6 I ~(FCC); Na Haeo (bio station), 5-l2.V.OI ,
THAILAND, Loei: Na Haeo (bio. station), 5-12.Y.Ol, secondary secondary forest, Constant & Grootaert, I 6 2 ~~ (RBINS), I c)
forest, Constant & Grootaert, I ~ (RBINS); Na Haeo (bio. (FCC); Na Haeo (bio station), 5-12.Y.OI , Constant & Groo
station), 7-14.V.OO, Malaise trap, P. Grootaert, I ~ (RBINS). taert, station I (small trees, dense ground vegetation, open
CAMBODIA, Siem Reap: Angkor Thom, station 24048, canopy), I ~ (RBINS); Na Haeo (bio station), 5-12.V.Ol,
30.Vli.04, P. Grootae11, I 6 (RBINS). Constant & Grootaert; pitfall station 3 (bamboo, very short
A small widespread species, occurring from Myanmar ground vegetation, thin litter; recently burnt), 2 c)c) (IRSNB), '
to southern China (NAVIAUX, 1995). I 6 (FCC); Na Haeo (field res. stat.), 15-19.V.03, light trap,
J. Constant, K. Smets & P. Grootaert, I ~ (RBINS); Na Haeo
I I. Neocollyris (Leptocollyris) variicomis (CHAUDOIR, field res. sta., 15-19.V.03, clay catch, J. Constant & K. Smets,
1864) I 6 (RBINS), I c) (FCC).
THAILAND, Loei: Na Haeo, 31.V-6.Yl.99, P. Grootaert, A well-known species, distributed from Myanmar to
1 <jl (RBINS); Na Haeo, station 20023, 25.V.OO, P. Grootaert, Vietnam (WIESNER, 1992).
16 Fabio CASSOLA
16. Prothyma (Genoprothyma) heteromal/a (MAC LEAv , 20. Heptodonta eugenia CHAUDOIR, 1865
1825) THAILAND, Loei: Na Haeo, 31.V-6.Vl.99, P. Grootaert, I ~
CAMBODIA, Siem Reap: Angkor Tom, station 24048, 30.Vli.04, (RB!NS); Na Haeo, 24-30.1V.OO, Malaise trap, P. Grootaert,
P. Grootaert, I ~ (REINS). 2 cScS (RBINS); Na Haeo, 14-20.V.OO, Malaise trap, P. Groo
Never formally recorded from Cambodia previously taert, I c5 I ~ (RB!NS); Na Haeo (Bio Station), 22-29.VII.OO,
(RJYALIER, 1964; WIESNER, 1992). Malaise trap, P. Grootaert, I c5 (RBINS); Na Haeo, 29.IV-
6. V .0 I, Malaise trap, P. Grootaert, 3 c5 c5 2 ~~ (RBINS); Na
Haeo, 29.IV-6.V.Ol, Malaise trap, Constant & Grootaert,
17. Prothyma (Genoprothyma) bouvieri W. HoRN, 1896 4 cScS I ~ (RBINS), I cS I ~ (FCC); Na Haeo (bio station),
CAMBODIA, Siem Reap: Siem Reap, 2-3.11.03, neon light 5-12. V .0 I, secondaty forest, Constant & Grootaert, 2 c5 c5 2 ~~
trap, Stephane De Greef, I ~ (REINS); Angkor Thom, (RBINS); Na Haeo (bio station), 5-12.V.OI , Constant & Groo
2.VI-3l.Vll.03, net catching, !G 30.192, Daniel R. Jump, I c5
taert; station I (small trees, dense grolll1d vegetation, open
(FCC). canopy), I c5 (RBINS); Na Haeo, 5-12.V.Ol, secondary forest,
Formerly mentioned from Cambodia just by WIESNER Malaise trap, Constant & Grootaert, I ~ (RBINS); Na Haeo,
( 1992), this species has definitely to be considered as 22-29.V.03, Malaise trap 3, P. Grootaert, I c5 (RBINS); Na
occuning in such a country. Haeo, 29.V-5.VI.03, Malaise trap 4, P. Grootaert, 2 cScS
(RBINS).
18. Prothyma (Genoprothyma) fallaciosa RIVALIER, CAMBODIA, Siem Reap: Angkor Thom, 23.V.03, day catch,
1964 J. Constant, P. Grootaert & K. Smets, I c5 (RBINS); Phnom
THAILAND, Loei: Na Haeo, 24-30.1V.OO, Malaise trap, P. Groo Kuen, day catch, 24-25.V.03, J. Constant & K. Smets, I ~
taert, I c5 I ~ (REINS), I ~ (FCC); Na Haeo (bio station), (RB!NS), I ~ (FCC); Angkor Thom, 2.VI-3l.VTI.03, net catch
ing, IG 30.192, Daniel R. Jump, I c5 (RBINS), I c5 (FCC).
5-12.V.OI, pitfall station 3 (bamboo, very short ground vegeta
tion, thin litter; recently burnt), Constant & Grootaert, 2 cScS A common widespread species, usually collected on
(RBINS); Na Haeo (bio station), 5-12.V.OI , pittfall station 4 small vertical slopes or roadcuts (NAVIAUX & PINRATANA,
(secondmy forest, tall trees, dense ground vegetation, densely 2004). Its occurrence in Cambodia had already made
covered with litter), Constant & Grootaert, I c5 (RBINS); Na known by SAWADA & WIESNER, 1999).
Haeo (bio station), 5-12.V.Ol, pittfall station 6 (abandoned
field: grasses, rough vegetation and open areas), Constant &
21. Therates pseudoprobsti PROBST & WIESNER, 1994
Grootaert, I c5 (RBINS); Na Haeo (bio station), 5-12.V.O I,
THAILAND, Loei: Na Haeo Nat. Park, 7.V.OI, Constant &
pittfall station 7 (bamboo forest, very thin litter, almost no
Grootaert, I ~ (RBINS).
ground vegetation), Constant & Grootae11, I ~ (REINS), I ~
Apparently a Thai endemic (PROBST & WIESNER, 1994;
(FCC); Na Haeo, 5-12.V.Ol, secondaty forest, Malaise trap,
P. Grootaert, 2 cScS (RBINS), I c5 (FCC); Na Haeo, 5-12.V.OI, NAVIAUX & PrNRATANA, 2004).
secondary forest, Malaise trap, Constant & Grootaert, I ~
(RB!NS); Na Haeo (bio station), 5-12.V.OI , light trap, Constant 22. Caloc/zroa f. jlavomaculata (HOPE, 1831)
& Grootaert, I cS I ~ (IRSNB), I cS (FCC); Na Haeo FIRS, THAILAND, Loei: Na Haeo (bio station), 5-12.V.01, light trap,
16.V.03, station 23030, P. Grootaert, I c5 (REINS); Na I-laeo, Constant & Grootaert, I ~ (RBINS); Na Haeo (bio station), 5-
18-31.V.04, Malaise trap 4, P. Grootaert, I c5 (FCC). 12.V.O I, light trap, Constant & Grootaert, 3 cScS 8 ~~ (REINS),
Described from Laos and the "haut-Mekong" I ~ (FCC); Na Haeo (bio station), 5-12.V.Ol, light trap, pitfall
(RIVALIER, 1964), and often confused with schmidt station I (small trees, dense ground vegetation, open canopy),
goebeli in many collections, this species has already Constant & Grootaert, 1 c5 (RBINS); Na Haeo (bio station), 5-
12.V.O I, light trap, pitfall station 2 (bamboo, some old trees,
proved to be quite widespread in Thailand (NAYIAUX,
1991). very short ground vegetation), Constant & Grootaert, 1 ~
(RB!NS); Na Haeo (bio station), 5-12.V.OI, light trap, pitfall
station 6 (abandoned field: grasses, rough vegetation and open
19. Protltyma (Genoprothyma) rapillyi NAVIAUX, 1989 areas), Constant & Grootaert, I ~ (RBINS); Na Haeo (field res.
THAILAND, Loei: Na Haeo, station 20023, 25.V.OO, P. Grootaert, stat.), 15-19.V.03, light trap, J. Constant, K. Smets & P. Groo
2 cScS I ~ (RB!NS), I c5 (FCC); Na Haeo, station 20025, taert, 2 cScS 2 ~~ (REINS), I c5 (FCC); Na Haeo (edge pond),
26.V.OO, P. Grootaert, I ~ (RB!NS); Na Haeo (bio station), 17.V.03, light trap, J. Constant & K. Smets, I Cj? (REINS).
5-12.V.O I, light trap, Constant & Grootaert, 2 cScS (REINS), CAMBODIA, Siem Reap: Phnom Kulen, 24.V.03, light trap,
I ~ (FCC); Na Haeo (bio station), 5-12.V.O I, secondary forest, 1. Constant & K. Smets, I c5 3 Cj?Cj? (RBINS), I Cj? (FCC); Angkor
Constant & Grootaert, I c5 (RBINS); Na Haeo (bio station), Thom, 26.V.03, night collecting, IG 30.192, Daniel R. Jump,
5-12.V.O I, pitfall station 3 (bamboo, very short ground vegeta I c5 I Cj? (RB!NS), I Cj? (FCC); 8 km N ofSre Noi (road to Anlong
tion, thin bitter, recently burnt), Constant & Grootaert, 1 c5 Vaeng), 29.V.03, light trap, J. Constant & K. Smets, 2 Cj?Cj?
(FCC), I ~ (RBINS); Na Haeo (bio station), 5-12.V.O I, pitfall (RBINS), I c5 (FCC); 8 km N of Sre Noi (road to Anlong
station 7 (bamboo forest, almost no ground vegetation), Con Vaeng), 29-30.V.03, dty dipterocarp forest, day, J. Constant
stant & Grootaert, 3 cScS (IRSNB); Na Haeo, 29.V-5.Vl.03, & K. Smets, I c5 (RB!NS); Angkor, Preah-Kahn, Temple,
Malaise trap 4, P. Grootaert, I ~ (RBfNS); Na Haeo FIRS, 31.V.03, light trap, J. Constant & K. Smets, I c5 (RBINS);
station 23030, 16.V.03, P. Grootaert, I c5 (FCC). Angkor Thom, 2.VI-3l.Vll.03, net catching, IG 30.192, Danic;,l
Described from eastern Thailand (NAVIAUX, 1989), this R. Jump, I Cj? (RBrNS).
species apparently is a Thai endemic. Its proper separa Quoted by former authors under sexpunctata Fabr.
tion from schmidtgoebeli sometimes appears to be rather (ACCIAVA TTl & PEARSON, 1989), this is a common,
delicate. However, the two species even belong to differ well-known species, widely distributed from India to
ent subgenera. the Philippines, and often collected at lights by night.
Tiger beetles from Thailand and C<;~~nbodia 17
23. Calocltroa moultotii (CHAUDOIR, 1865) A common species, widely distributed from Myanmar
THAILAND, Loei: Na Haeo (bio station), 5-12.V.OI, secondary to Vietnam and southern China {NAVIAUX & PINRATANA,
o o
forest, Constant & Grootaert, I 2 ~~ (RBINS), I (FCC). 2004).
A polymorphous species, from which elegantula at
least can be considered as specifically distinct (SAWADA
26. Calocltroa bramani (DOKHTOUROFF, 1882)
& WIESNER, 2002). Co-occurrence of both species in the
CAMBODIA. Siem Reap: Angkor Thom, 23.V.03, sweeping,
Naeo station clearly confirms such a separate status. J. Constant, K. Smets & P.Grootaert, I 0 (RBINS); Phnom
Kulen, 24.V.03, light trap, J. Constant & K. Smets, I
24. Calochroa elegantula (DOKHTOUROFF, 1882) (FCC); Phnomo Kulen, 24-25.V.03, day catch, J. Constant &
THAILAND, Loei: Na Haeo, station 20023, 25.V.OO, P. Grootaert, K. Smets, I (RBINS), I ~ (FCC); Angkor Thom, 2.Vl-
I o (RHTNS), I o (FCC); Na Haeo, station 20025, 26.V.OO, 31.VII.03, net catching, IG 30.192, Daniel R. Jump, I 0 2 ~~
P. Grootaert, I o (RBINS); Na Haeo, 21-29.1V.O I, Malaise (RBINS).
trap, Constant & Grootaert, I 0 (RBTNS); Na Haeo, 5- The above recorded data definitely confirm the occur
12. V .0 I, seconda1y forest, Malaise trap, Constant & Grootaert, rence in Cambodia of this beautiful species, already
o
I (FCC); Na Haeo (bio station), 5-12.V.OI , secondary forest, known from eastern Thailand to Vietnam (SA\-VADA &
Constant & Grootaert, I 0 (RBJNS); Na Haeo (bio station), 5- WIESNER, 2002; NAVIAUX & P!NRATANA, 2004).
12.V.OI , pitfall station 2 (bamboo, some old trees, very short
o
ground vegetation), Constant & Grootaert, I (RBTNS);
Na Haeo (bio station), 5-12.V.Ol, pitfall station 3 (bamboo, 27. Cosmodela auru/enta (FABR.) ssp. juxtata (ACCIA-
ve1y short ground vegetation, thin litter, recently burnt), Con VATTI & PEARSON, 1989)
stant & Grootaert, I 0 (RBTNS); Na Haeo (bio station), 5- THAILANoDo, Loei: Na 1-laeo, Station 20023, 25.V.OO, P. Groot
12. V.O I, pitfall station 4 (secondary forest, tall trees, dense aert, 2 4 ~~ (RBTNS), I 0 (FCC); Na 1-laeoo (bio station),
ground vegetation, densely covered with litter), Constant & 5-12.V.Ol, light trap, Constant & Grootaert, I I ~ (RBINS),
Grootaert, I o (RBINS); Na Haeo (bio station), 5-12.V.O I, I ~ (FCC); Na Haeo (riverbanks), 5-12.V.OI, Constant &
pitfall station 6 (abandoned field: grasses, rough vegetation Grootaert, I 0 (RBTNS); Na 1-laeo, Chang Tok Wateifalls,
and open areas), Constant & Grootaert, I o (RBTNS); Na Haeo IO.V.OI, light trap, Constant & Grootaert, I ~ (RBTNS); Na
field res. sta., 15-19. V .03, day catch, J. Constant & K. Smets, 1-laeo (field res. stat.), 15-19.V.03, light trap, J. Constant,
I o (RBlNS); Na Haeo, 22.V.03, Malaise trap 2, P. Grootaert, K. Smets & P. Grootaert, 4 00 I (RBlNS), I ~ (FCC); Na
I ~ (RBINS). l-loae o (edge pond), 17.V.03, light trap, J. Constant & K. Smets,
I (RBINS).
Co-occurrence of both elegantula and mouhotii at the
A common widespread species, occurring with nom
Naeo station clearly confirms their separate specific
inal aurulenta from the Sunda Islands to Malacca and
status, as proposed by SAWADA & WIESNER (2002).
with subspeciesjuxtata from north-eastern India to Viet
nam and southern China (ACCIAVAT Tl & PEARSON, 1989;
25. Calocltroa interruptofasciata (SCHMIDT-GOEBEL,
NAVIAUX & PINRATANA, 2004).
1846)
THAILAND, Loei: Na Haeo, station 20022, 24.V.OO, P. Grootaert,
I o (RBINS); Na Haeo, station 20023, 25.V.OO, P. Grootaert, 28. Lophyra (Lopltyra) fuliginosa (DEJEAN, 1826)
4 oo (RBINS); Na Haeo, station 20025, 26.V.OO, P. Grootaert, THAILAND, Ranong: Kon Samed Island, 15-17.V.Ol, Constant &
I o (RBINS); Na Haeo (bio station), 5-12.V.OI , light trap, Grootaert, 2 o" 5 <n (RBINS), I o (FCC).
Constant & Grootaert, I ~ (RBINS); Na Haeo (bio station), 5- A fluviatile species, widely distributed from the Sunda
12.V.O I, pitfall station I (small trees, dense ground vegetation, Islands to Malacca, Myanmar, Thailand, Indochina and
open canopy), Constant & Grootaert, I ""(RBINS); Na I-Iaeo, 5- China. It is not known from India, and former odd records
12. V.O I, pitfall station 3 (bamboo, some old trees, ve1y short from Sri Lanka proved to have to be discarded (NAV IAUX,
ground vegetation), Constant & Grootaert, I 0 (RBINS); Na 1984; ACCIAV A TTl & PEARSON, 1989).
Haeo (bio station), 5-12.V.OI , pitfall station 6 (abandoned
field; grasses, rough vegetation and open areas), Constant &
Grootaert, I ~ (RBINS); Na Haeo (bio station), 5-12.V.Ol, 29. Lopltyra (Spilodia) line~frons (CHAUDOIR, 1865)
sIe oco nId a1(yF CfoCr)e;s tN, Ca oHnasetaon t( b&io Gsrtoatoiotane)r,t ,5 2- 1020.V .2O I~, l(iRghBt INtrSap),, T&H GAIrLoAoNtDae, rLt,o eI io:o N ( Ra B1-llaNeoS,) ;2 4N-a3 01.-lIaVe.oO, O2, 5M.Va.lOaOis,e S ttraatpio, nC o2n0s0t2an3t,
Constant & Grootaert, I ~ (RBINS); Na Haeo, 5-12.V.Ol, sec P. Grootaert, I I (RBTNS); Na Haeo (bio. station), 5-
ondary forest, Malaise trap, Constant & Grootaert, I 12.V .0 I, pitfall station 6 (abandoned field; grasses, rough
(RBINS); Na Haeo Khring Tok, station 23043, 19.V.03, vegetation and open areas), Constant & Grootaert, I 0
P. Grootaert, I _ (RBINS); Na 1-laeo (field research station), (RBINS); Na Haeo (bio. station), 5-12.V.O 1, light trap, Con
15-18.V.03, white tray, J. Constant, K. Smets & P. Grootaert, stant & Grootaert, 2 _~ (RBINS), 2 ~ (FCC); Na l-laeo (bio.
I o (RBINS); Na 1-laeo field res. sta., 15-19.V.03, day catch, station), 8-15.1V .0 I, Malaise trap, P. Grootaert, I ~ (RBlNS);
J. Constant & K. Smets, I ~ (RBINS). Kanchanaburi: Sai Yok Na l-laeo (bio station), 5-12.V.Ol, light trap, Constant &
N.P., 4-5.VI.03, J. Constant & K. Smets, I " (RBINS). Grootaert, I (RB INS).
CAMBODIA. Siem Reap: Angkor Thom, 23.V.03, day catch, CAMBODIA, Siem Reap: Angkor Thom, 23.V.03, day catcf.,
J. Constant, P. Grootaert & K. Smets, I 0 (RBINS), I 0 J. Constant, P. Grootaert & K. Smets, 2 00 (RBTNS); Angkor
(FCC); Angkor Thom, 26.V.03, night collecting, IG 30.192, Thom, 2. Vl-31. VI1 .03, net catching, IG 30.192, Daniel R.
Daniel R. Jump, I o (RI31NS); Angkor Thom, 2.VI-31.VII.03, Jump, 2 oJ I (RBlNS), I o (FCC).
net catching, JG 30.192, Daniel R. Jump, I 0 (FCC), I A common species, widely distributed in continental
(RBINS). Asia from northern India to Vietnam (WIESNER, 1992).
18 Fabio CASSOLA
I I
30. Naviauxella ramai NAVIAUX, 1991 nal plates shiny blue-green; fully glabrous except for t\'Vo
THAILAND, Loei: Na Haeo, Chang Tok, 17.V.03 (23035), setae or setigerous punctures on fixed loci near eyes.
P. Grootaert, I (j (RBTNS). Striations fine on frons, concentric on vertex, stronger
CAMBODIA, Siem Reap: Phnom Kulen, 24.V.03, light trap,
on eyes, waved and oblique behind on neck. Eyes pale
J. Constant & K. Smets, I ~ (FCC). yellow, globose. Labrum with a longitudinal carina in the
A small species, described from Thailand and thus new
middle, distinctly wider than long, a bit longer and trian
to Cambodia. However, the above mentioned female
gular shaped in the female, slightly retracted in front in
specimen from Phnom Kulen, Cambodia, is a bit darker
the male, with a small tooth protruding in front in both
and moreover it has a dark, pitchy black, slightly metallic
sexes; fully metallic bronze, with eight submarginal setae
labrum.
some distance from f01ward edge, setigerous punctures
apparent, bright metallic blue-green. Mandibles yellow
31. Cylimlera (lfasina) foveo/ata (SCHAUM, 1863) ish-testaceous, more or less darkened on outer half. La
THAILAND, Loei: Na Haeo (bio station), 5-12.V.OI , light trap, bial and maxillary palpi testaceous, the last joint dark
Constant & Grootaert, I (j (RBfNS). metallic greenish with the extreme tip testaceous. Anten
CAMBODIA, Siem Reap: Angkor Thom, 26.V.03, night collect
nae slender, long, reaching over the half of the elytral
ing, IG 30.192, Daniel R. Jump, I 0 (RBfNS).
length in the male, slightly shorter in the female; anten
A fairly distinct, common species, widely distributed
nomeres 1-4 metallic bronze-green, segments 5-11 dark
from southern India, the Sunda Islands and the Philip
blackish-brown, dull, finely and evenly pubescent.
pines up the whole of continental SE Asia (WIESNER,
Thorax: pronotum subsquare, approximately as wide as
1992). However, it had never been recorded previously
long, parallel-sided, glabrous except for some scattered
from Cambodia, thus formally being new to such a
setae on sides, with fine descending wrinkles in the
country.
middle of disc; cupric-bronze with green reflections on
sides, transversal constrictions poorly marked. Episterna
32. Cylindera (Ifasina) viduata (FABRICIUS, 1801)
sparsely setose. Female mesepisternal coupling sulcus a
THAILAND, Loei: Na Haeo (bio station), 5-12.V.O I, light trap,
n small rounded pit in the middle.
Constant & Grootaert, I 0 6 ~~ (RBfNS), 2 (FCC); Na Haeo
Elytra proportionally wide, convex, with well-marked
(bio station), 5-12.V.O I, light trap, pitfall station 6 (abandoned
shoulders, colour dark purple with blue-green round
field: grasses, rough vegetation and open areas), Constant &
Grootaert, I (j (RBfNS); Ranong: Kon Samed Island, 15- punctures, the punctures being stronger and especially
n
17.V.O I, Constant & Grootaert, 3 (j(j 2 (RBINS), I (j evident in front third of elytra, shallower behind; a darker,
(FCC). Kanchanaburi: Sai Yok N.P., 4-5.VI.03, J. Constant & dull, unpunctured, nanow striped area runs longitudinally
K. Smets, I ~ (RBTNS). on disc, some distance from the suture. Apical border
CAMBODIA, Siem Reap: 8 km N of Sre Noi (road to Anlong distinctly rounded with a small sutural spine. Epipleura
Vaeng), 29.V.03, light trap, J. Constant & K. Smets, I (j (FCC). light coloured, almost testaceous, with some metallic hue.
This is a well known, widespread South-east Asian Elytral markings consisting of a conspicuous roundish dot
species, distributed from northern India and Nepal east on front third of disc, which obviously represents the end
ward into China, Malaysia, Indonesia and the Philippines. of a non-existing humeral lunule; a transversal, slightly
All the above mentioned specimens have the subapical oblique, middle band, made up by a transversal sublateral
dot more or less narrowly and briefly pointed toward the spot obliquely connected with a subtriangular, rather
suture.
large, discal spot, which lies in the dull, longitudinal,
striped area of elytra; and a large, subrectangular, com
33. Cylindera (Ifasina) khmer n. sp. (Fig. 1 a-d) ma-shaped, subapical spot, which is obliquely directed in
front on disc and shortly expanded along the subapical
TYPE MATERIAL Holotype, c), from Cambodia (Siem Reap margin toward the apex. Humeral dot lacking in both
province): Angkor, Preah-Kahn, Temple, 3l.V.03, light sexes. Moreover, in the allotype and paratype specimens
trap, J. Constant & K. Smets; allotype, ~' also from Cam only, a very narrow, poorly visible, lineole is detectable
bodia: Phnom Kulen, 24.V.03, light trap, J. Constant & which runs along the lateral border of elytra and almost
K. Smets; one paratype, c), from Angkor Thom, 26.V.03, connects the middle band to the subapical spot.
light trap, J. Constant, K. Smets & P. Grootae11; one Underside metallic dark green, with white sparse pubes
paratype, ~' from a different locality of Cambodia (Siem cence on sides of sternum and the abdominal stemites.
Reap province): 8 km N ofSre Noi, road to Anlong Vaeng, Trochanters dark rufous, femora metallic bronze green,
29.V.03, light trap, J. Constant & K. Smets. Holotype, tibiae and tarsi more or less tinged with metallic dark
allotype and paratype ~ in REINS, paratype 6 in FCC. violaceous and some greenish reflections; a few rows of
DIAGNOSIS: A small Cy/indera (lfasina) species similar spinifonn setae on tibiae and tarsi, the setae being more
to, and obviously a close relative of, viduata, however numerous on the apical half of the middle tibiae (toilet tool}.
syntopically occurring with it and differing because of the Aedeagus fusiform, straight, tapering, slightly curved
oblique, narrow, continuous middle band and the larger, downward apically, very similar to that of viduata; inner
comma-shaped subapical dot. sac typical of the genus Cylindera, with a long narrow
DESCRIPTION: Head bronze with greenish and cupreous flagellum whose upper convolutions are crossed by a V
reflections especially on eyes, clypeus and cheeks, anten- shaped sclerite ("equerre") (RIVALIER, 1950).
Tiger beetles from Thailand and Cambodia 19
8
~ r~~
.......
1-... ' //",
-;
/
D
,•. .·.
C); ··o
..
~
Q
"
A
Fig. I. - Cylindera ((j'asina) khmer n. sp., male holotype from Preah-Kahn, temple, Angkor, Cambodia. A. habitus; B. labrum;
C. aedeagus and inner sac; 0. female allotype from Phnom Kulen, Cambodia: labrum . Scale-lines: I mm and 0.5 mm.
20 Fabio CASSOLA
Length: 7.5-8 mm (without labrum). line singular emendation (spino/ai) by FLEUTIAUX ( 1902),
ETYMOLOGY: This new distinctive 1fasina species is so considered to be justified by ACCIAVATTI & PEARSON
named in memory and celebration of the Khmer civiliza ( 1989), would appear to be untenable according the the
tion and empire (IX-XV centuries), whose famous capital Zoological Code (ICZN, 1999).
and jungle temple of Angkor, where the holotype speci
men has been collected, stands as one of the great jewels 35. Cylimlera (lfasina) juergemviesneri NAVIAUX, 1991
of the human cultural heritage. Special thanks are to be THAILAND, Loei: Na Haeo, river before village, 8.V.O I, Con
expressed to the actual collectors of this new species. stant & Grootaert, 3 CSCS 3 ~ (RBINS), 3 CS" 2 ~~ (FCC); Dan
REMARKS: C. (1.) khmer n. sp. is obviously a close relative Sai, Bantu11, II.V.OI, Constant & Grootaert, I (S I ~ (RBINS).
Apparently a Thai endemic (NAVIAUX, 1991; NAVIAUX
of C. (!.) viduata (FABRICIUS, 180 I), and it could even be
& PINRATANA, 2004), this small species is clearly a close
considered a subspecific form of it, if it were not for the
remarkable stability shown by viduata over all its enor relative of C. (1.) discreta (SCHAUM, 1863). Past records
mous geographical range. In particular, however, viduata of the latter species from Cambodia (HORN, 19 13; WIES
has smaller, almost effaced elytral dots, it lacks any NER, 1992) may well refer, in reality, to juergenwiesneri
(Table 1).
subhumeral dot, it has a smaller and narrower subapical
spot, and moreover it was found to syntopically occur
36. Cylindera (Eugrapha) minuta (Olivier, 1790)
with khmer in one Cambodian locality at least (see
THAILAND, Loei: Na Haeo FIRS, 16.V.03, station 23032,
above); other SE Asian lfasina species, such as humillima n
P. Grootaert, 3 (S(S 2 (RBINS), I (S I ~ (FCC).
(GESTRO, 1893), macilenta (SCHAUM, 1862), dilatotarsa
CAMBODIA, Siem Reap: Tonie Sap Lake, Prek Toal, 27.V.03,
(W. HORN, 1924), kaleea (BATES, 1866), decempunctata light trap, J. Constant, K. Smets & P. Grootaert, I ~ (RBINS).
(DEJEAN, 1825) and modica (GESTRO, 1893) have a nar This common widespread species, known to range from
row and complete apical lunule (HORN, 1938); pronotalis India into Southeast Asia and Indonesia (ACCIAVATTI &
(W. HORN, 1922) has much smaller elytral dots and PEARSON, 1989; WIESNER, 1992), had not yet been formally
a different, shagreened, pronotal sculpture; humerula recorded from Cambodia.
(W. HORN, 1905) has a complete humeral lunule; the
viridilabris-group species [namely viridi/abris (CHAUDOIR, 3 7. Myrioclzila (Myriochila) sinica (Fieutiaux, 1889)
1852), somnuki NAV IAUX, 1991, and fallaciosa (W. HORN, CAMBODIA, Siem Reap: Phnom Kulen, 24-25.V.03, day catch,
1897)] have a roundish, instead of comma-shaped, sub J. Constant & K. Smets, I (RBfNS); 8 km N of Sre Noi (road
apical spot, as also have spinolae (GESTRO, 1889) and to Anlong Vaeng), 29.V.03, light trap, J. Constant & K. Smets,
subtilesignata (MANDL, 1970), which, in addition, have a I ~ (RBINS). Angkor, Preah-Kahn, Temple, 31.V.03, light trap,
differently coloured, much reddish pronotum; paucipilina J. Constant & K. Smets, 3 ~~ (RBINS), I (FCC);
(ACCIAVA TTl & PEARSON, 1989) and paeninsu/aris A Southeast Asian species, known to occur in muddy
areas from Myanmar to Vietnam and southern China
NAVIAUX, 1891, have a testaceous/pale labrum. The most
(NAVIAUX & PINRATANA, 2004), but puzzlingly (perhaps
similarly looking species, as far as the general shape and
erroneously) indicated from Java too (WIESNER, 1992). 1t
the elytral markings are concerned, is balabacensis
was firstly formally recorded from Cambodia by SAWADA
NAVIAUX & SAWADA, 1996, described from Balabac I. in
& WIESNER (1999).
the Philippines, which, however, lacks any middle band
(NAVIAUX & SAWADA, 1996).
34. Cylindera (lfasina) spillolae (GESTRO, 1889)
Aknowledgments
THAILAND, Loei: Na Haeo (bio station), 5-12. V.O I, secondary
forest, Constant & Grootaert, I CS (FCC); Na J-laeo (bio station),
5-12.V.O I, Malaise trap, P. Grootaert, I CS (RBINS); Na Haeo, Thanks to the courtesy of Mr. Alain Drumont (Royal Belgian
Institute of Natural Sciences, Bruxelles, Belgium), I have had the
Chang Tok Waterfalls, I 0. V.O I, Constant & Grootaert, I ~ opport1mity of examining the tiger beetle specimens collected in
(RBINS); Na Haeo Khring Tok, 19.V.03, 23041, P. Grootaert, Thailand and Cambodia during several field research missions that
I (S (RBINS). the above mentioned Institute (RBINS) carried out in the years 2000-04
A common species, wide! y distributed from eastern (by P. Grootaert, J. Constant, K. Smets and D.R. Jump). Such materials
provide some interesting additional data, including a species new to
Nepal and north-eastern India eastward to Vietnam
science, from Cambodia, which are duly recorded below. The speci
(WIESNER, 1992; SAWADA & WIESNER, 1999). The mascu- mens cited are either in the RB!NS or in author's (FCC) collections .
..
Tiger beetles from Thailand and Cambodia 21
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