Some Species of the Genus Oxycentrus : Coleoptera, Carabidae, Harpalini

10 Pages·1996·11.5 MB·English

by  Ito, N

Elyt1・a,Tokyo,24(1):31-40,May15,1996
SomeSpeciesoftheGenusOxycentrus
(Coleoptera,Carabidae, Harpalini)
Noboru ITO
1-7-l8Higashiuneno, KawanishiCity,HyogoPref.,666-01Japan
Abstract Three new speciesof thegenus 0.、、,,(、entrusCHAuDolR, 0.bae1/1・i
mes, , and0.g・lad加al・.・lIsfromThailandand0.a_fl/鑰n,71 fro Tonkin ar described an
aredescriptionof0.acutulusBATESfromMyanmarisgiven.
Introduction
In thispaper, I amgoing to redescribeOxycentrus(s.str) acutu/usBATES
fromMyanmarandtodescribethreenewspecies, Ox;ycentrus(s.str)baehr1and
0. (0、ycentropsfs)gracf加arsfsfromThailandand 0.(0、.yce,7fro- s)act,flpemzs
fromTonkin. 0)cycentrusbaehr1resembles0. acutulusBATES,but isdifferent from
the latter in having theantennaeslenderer, theapical angleslessprominent and
theelytramostlynot reddishbrown. 0x;ycent1usgracilitarsisischaracterizedby
the subsquare pronotum and the slender tarsi. The species of the subgenus
Oxycentr〔)pslshavehitherto been known only from the Malay Peninsula to the
SundaIslands,andisfirstlyrecorded fromtheIndo-chineseRegion.
IwishtoexpressmydeepgratitudetoDr.MartinBAEHRoftheZoo1ogische
Staatssammlung,Miinchen, Dr.FritzGUsENLFITNERoftheLandesmuseum, Linz,
and Dr. Ivan LoBLof theMuseumd'Historienature11e, Geneve, for their kind
offeringmanyvaluablematerials. 0、1ycentrusbaehri isnamedafter Dr. Martin
BAEHR. I alsoheartilythank Dr.RobertPoGGloftheMuseodiStoriaNaturale,
Geneva, forhiskind loanoftypespecimensandMr.TaichiSHIBATA forhiskind
guidanceinmystudy.
Oxycentrus(s.str)acutulusBATES
(Figs.1,5)
()xycent1・us acutulus H. W. BATES, 1892, Annli. Mus. civ. Stor nat. Geneva, (2), 12: 344. -
KIRscHENHoFER,1992, Linzerbiol.Beitr.. 24:51.
Bodysubcylindrica1,elongate,parallel-sided,moreor lesssimilarinformto
small examples of Oxycentrus (s. str ) melas (ScHMIDT-GoBEL), dark reddish
brown,shiny,withoutiridescentlustre;headblackexcept forpalpiandantennae
32 NoboruITO
light reddishbrown and labrum andmandibles dark brown, femora yellowish
brown.
Headnarrow,alittlelessthan two-thirds thepronota1width, rather convex,
very sparsely and minutely punctate; labrum transversely quadrate, straight at
apex;clypeusgentlyconvex,obscurelyrugoseatinnersideofeachlateralsetiferous
pore;clypea1sutureclearandmoderatelydeep; frontal impressionssubarcuately
divergentbehind, relativelydeep,shallowednearsupraorbitalgrooves;eyessmall,
not strongly prominent; temple long, ahalf theeyelength, drawinga gently
emarginatecurve; genuineventral marginofeyewidelyseparated frombuccal
fissure; mandibles moderately long, robust, clearly curved inwards; antennae
submoniliform,short,notprolongedbackwardsbeyondelytra1bases,3rdsegment
ratherwelldilateddistad,sparselypubescentinapicalhalf,aslongasthe4thand
ahalflonger than the2nd;3rdsegmentoflabialpalpuswell thickenedmedially
andone-eighthlongerthan the2nd;ligulainverted-trapezoidal, truncateatapex;
parag1ossaemoderatelycurvedinportionsfreefrom ligula,prolonged in fronta
littlebeyondit;mediantoothofmentummoreweaklyporrectthanthoseofother
speciesof thegenus, transversely triangular,ep1iobesweaklywidened forwards;
microsculptureinvisibleunder80xmagnification.
Pronotumalmostsquare,widestatapical two-fifths,alittlewiderthanlong
(1.14inratio),indistinctlycontractedin frontandbehind,uniformlywellconvex,
verysparselyandminutelypunctateondiscandmoderatelyandrather coarsely
soinbasalfoveae;sideswidelybordered,weaklyroundedinfront fromthewidest
point,shallowlysinuatealittlebehindmiddle, thencesubpara11e1tobase; apex
finelybordered,ratherdeeplyemarginate,straightatthebottomoftheemargina-
tion;baseone-seventhwiderthanapex,feeblyarcuate,brokenlybordered;apical
anglesmoreor lessporrect,narrowly rounded;basalanglesalmostrectangular,
edentateat tips;lateral furrowsnarrowthroughout,eachwithamarginalsetaat
apical fifth; basal foveae ill-defined, only flattened in inclined of disca1
area
convexity; frontandhindtransverseimpressionsobliterated;medianlinefineand
clear,reducednearapexandsomewhatdeepenednearbase;microsculpturemostly
invisible,detectableasisodiametricmeshesinthefurrowsandasvaguetransverse
lines in basal foveae.
Hindwingscomplete. Elytraelongateandwellconvexasinthespeciesof
Batoscelis DEJEAN, one and three-fourthsas longaswide, very sparsely and
microscopicallypunctate;sidesalmostparallel,slightlyarcuateinhumeri,wjth
more or less deep apical sinus; apices not produced behind, widely rounded,
narrowlyseparatedfromeachother,bluntatsuturalangles;basesalmosttruncate,
subrectangular anddistinctly toothedat humeralangles;striaewide, deepand
Clearlycrenulate,scutellarstrictoveryshort;intervalsweaklyraisedondisc,more
swollenlaterallyandapically,withoutsetiferousporeon3rdinterval;margjna1
seriesdivided into twogroups, the foregroupconsistingof6umbilicatepores
and thehindoneof7-8pores;microsculpturevisibleonlyineachlateral three
SomeSpeciesoftheGenus0:x;ycentrus
33
intervalswhicharetransverselymeshedinmostareasandisodiametrica11ysoin
part.
Ventralsurfacesparselyandrathercoarselypunctateonmeso-andmetepi-
stemaandlateralportionsofmetasternum,withveryshortandsparsepubescence
medially onmetasternumand2nd to 6th abdominalsegments; metepisternum
distinctlynarrowedbehind, twiceaslongaswide;6thabdominalsegment in
bisetoseoneachside,widelyandgentlyarcuateat tip.
str) baehrisp nov;3, 0. (0xycentropsis) graciiitarslssp nov; 4, 0. (0xycentropsis)
acuflpennl'ssp nov.
34 Noboru ITO
Legsrelativelyshort; femur tumid, hind femur with twosetaealonghind
margin; foretibiathick,distinctlydilatedapicad,weaklyobliqueinexternalhalf
ofapex,with threelongspinesalongapico-externa1margin, terminalspur long
andsimple;hindtarsusalittleshorterthanwidthofheadincludingeyes(0.86in
ratio), 1stsegment aslongasthe2nd and3rdcombined,2nd one-fifth longer
thanthe3rdandtwicethe4th,clawsegmentbisetosealongeachventralmargin.
Aedeagus(Fig 5)slender,weaklyarcuate;apical lobesubquadrate, rounded
and thickly bordered at distal margin; apicalorificenarrowly open, inner sac
armed with several conical sclerites.
Length:6.3mm.Width:2.2mm.
Specimen e1:camlned 1 (holotype), Tennasserim, Thagata, (Myanmar),
IV-1887,FEAleg.(preservedin theMuseodiStoriaNaturale,Geneva).
Thisspeciesisallied to 0)cycentrus(s.str)melas(ScHMIDT-GOBEL),but iS
distinguished fromthelatterbytheeyessmallerandlessprominent, thetemples
longerandthicker, theantennaemoretumid,theapicalanglesofpronotummore
stronglyproducedandmorenarrowly rounded, thehindanglesnot blunt, and
theelytra1dorsalporelacking.
Oxycentrus(s.str) baehrisp
nov
(Figs2,6,7)
Bodyelongate,butlessnarrowerthanin01,cycentrus(s.str)melas(SCHMIDT-
G〔I)BEL),subpara11e1atsides, pitchy black and often brownish, shiny, without
iridescent lustre; pronotumblack tobrownishblack, palpi, antennae, legs and
lateralmarginsofpronotumlight reddishbrown,labrum,mandibles,andlateral
marginsandsutural intervalsofelytrareddishbrown.
Headmoreor lessnarrow, two-thirdsthepronota1width,wideat interocular
spacewhich isalittlemorethanthree-fourthsthewidthofhead, ratherconvex,
very sparsely and minutely punctate; labrum transversely quadrate, angularly
roundedatapical corners;clypeussomewhat thick,smoothorbearingarugosity
like foveoleat internal sideofeach lateral seta;clypea1sutureclearly marked,
considerablydeep in individuals, even indepth; frontal impressionsdeep,weakly
arcuatelyrunningbehind,reachingsupraorbitalgrooves;eyesnotlargeandgently
convex; genuineventral marginofeyenarrowlyseparated from buccal fissure;
mandibles long and slender, abruptly curved before middle, coarsely
rugose
mediallyondorsalsurface,retinacular toothofrightmandibleminutelyproduced;
antennaesubmoniliform,slightlysurpassingpronota1base,3rdsegmentpubescent
inapical two-thirds,one-seventhlonger than the4thandoneand two-fifthsthe
2nd;3rdsegment of labialpalpusrelativelytumid inmiddle,equal in length to
the2nd; ligulaparallel-sided,weaklynotchedatapex;parag1ossaenarrow, fully
prolongedbeyondligula;epi1obesofmentumnarrowandnotwidened forwards;
microsculptureinvisibleunder80x magnification.
SomeSpeciesoftheGenus 0.、,1,(・ent1・u1. 35
Pronotumsubquadrate,widestatapicalthird,alittlewiderthanlong(about
1.2inratio),gentlyanduniformlyconvex,verysparselypunctulateondisc,rather
coarselyandmoderatelypunctateinlateralfu1Towsandbasalfoveae;sidesweakly
convergent towardsbothapexandbase,shallowlysinuatebe「orebase,withthick
borders; apexveryshallowlyemarginate,straight inmiddle,entirelybordered;
base a little wider than (1.1-1.2in ratio), almost truncate, vaguelyand
apex
brokenly bordered; apical angles not prominent, rather widely rounded; basal
anglesangulateand slightly larger than rectangle; lateral furrows deep, wholly
carvedinaline;basalfoveaesmall,onlyflat,shallowlyandlongitudinallygrooved
at innersides; front transverseimpressionshortandobscure,hindoneobsolete;
median line fine, shallow and reachingboth apex and base;surfaceobscurely
microsculpturedonlyinlateralfurrowsandbasalfoveaeandtransverselymeshed.
Hindwingsentirelydeveloped.Elytraelongate-oblong,alittlemorethanone
andseven-tenthsaslongaswide,almostflatondiscandabruptlydeclivousnear
sides, without punctures;sides weakly arcuateat humeri, a littlemoredeeply
sinuate before apices than in 0. melas; apices moreprominent than those of
0.melas,moreorlesswidelyrounded,notornarrowlyseparatedfromeachother,
sharpatsuturalangles;baseshardlyemarginate,humeralanglesangulate,alittle
obtuse and minutely toothed at tips; striae shallower than thoseof 0. nlelas,
clearlycrenulate,scutellarstrictoobliteratedorveryshort;intervalsflatorweakly
convex on disc, becoming more convex laterad and apicad, 3rd interval not
bearingsetiferouspore;marginal serieswidely interruptedinmiddle, composed
of8十(7-8)umbilicatepores;microsculpturevaguelyobservableaslinesondisc,
clearlydetectedastransversemeshesonthelateraldeclivousareas.
Ventral surface mostly smooth, coarsely punctate on lateral portions of
metasternum; metepisternumelongate, four-fifths longer than wide; abdomen
sparsely pubescent alongmiddleof2nd to6thsegments, 6thbisetoseon each
sidein bothsexes,subtruncatein andclearlyarcuatein
Hind femur bisetosealonghindmargin; foretibiastronglyexpandeddistad
andgentlyarcuateinwards,not incised inexternalhalfofapex,quadrispinous
apico-externa11y, terminalspur robust andlanceolate; tarsi relativelyshort, 1st
segmentsofforeandmidtarsinotbearingadhesivesquamaonventralside,hind
tarsus five-sixths in andabout three-fourths(0.78 in ratio) in the width of
head,1stsegmentaslongasthe2ndand3rdtogether,3rdthree-fourthsthe2nd
andone-fifthlonger thanthe4th,clawsegmentbl-or trisetosealongeachventral
margin.
Aedeagus(Fig 6)slender,almoststraightlyprolongeddistad;apexslightly
hookedabove;apicallobeaslongaswide,roundedandborderedatdistalmargin;
apicalorificewidelyopen, innersacbearingmanyconicalsclerites.Stylus(Fig.
7)slender andgentlycurved,withashortsetabasallyatexternalmargin;basal
segmentbisetoseapico-externally;valvifer bisetoseatapex.
Length: 6.2-7.0mm.Width:2.3- 2.7mm.
36 Noboru ITO
Holotype: Doi Inthanon,Chiang Mal Prov., Thailand, 8~9-V-1990,
,
MALIcKYleg. (preserved in theZoo1ogischeStaatssammlung, Miinchen). Para-
types:2 ,samedataastheholotype; thefollowingexamplesbearsamelocality
and collector as theholotype, I , 17~24-IV-1990, 1e, 25~29-IV-l990, I e,
1~3-V-1990,2 ,4~6-V-1990, 1e,19,10~11-V-l990,19, 12~14-V-l990.
ThisnewspeciesiscloselyalliedtoOx:ycentrus(s.str)acutuhtsBATES,but
differsinhavingtheantennaeslenderer, thepronotumlessprominentandmore
widelyroundedatapicalanglesandmorestronglycontractedbehindatsidesand
theelytralesselongateandmostlyblackinsteadofbeingreddishbrown.
Thenewspeciesisalsosimilarto 0)cycentrus(s.str)melas(ScHMIDT-GoBEL),
but is distinguished from the latter, in addition to the characters mentioned
above, by thebody wider and brownish in color instead of being black, the
pronotummore thickly bordered, and theelytrabearing reddish margins and
sutural intervalsinsteadofbeingblack.
Figs. 5-7. GenitaliaofOx:ycentrusspp. - 5, 0.(s.str)acutulusBATES;6&7, 0.(s.str)
baeh・tisp nov;6,male;7, female;a,dorsalview;1,lateral view.
SomeSpeciesoftheGenusOxvcent・,us 37
Orycentrus(0xycentro -s')graeill'tarsissp
nov.
(Figs3,8)
Body oblong, relatively convex, pitchy black,shiny,slightly iridescent on
elytra;maxillae,maxillaryandlabialpalpiyellowishbrown,antennaelightbrown,
legsbrown,mandiblesandlateralmarginsofpronotumdark reddishbrown.
Head evenlyconvex, narrow, about two-thirds the pronota1width, very
sparsely andminutely punctateonly on frons; labrum transverselytrapezoidal,
shallowly emarginate at apex; clypeus narrowly depressed along apex, gently
humpedbehindthedepression,withadeeprugosityneareachside;clypea1suture
fineandclear,evenindepththroughout; frontalimpressionsdeeplycarved,gently
arcuateoutwardsandreachingsupraorbitalgrooves;eyescomparativelylargeand
hemispherica11y convex; temples thinandshort,subrectangularlymeetingwith
neckconstriction;genuineventralmarginofeyeadjoiningbuccalfissure;antennae
slender andextendinga littlebeyondelytra1base, 3rdsegment slightly dilated
distad,slightlyshorter thanthe4th(0.95inratio)andthree-fifthslonger thanthe
2nd;mandiblesrobustandmoderatelysharp, leftmandibleweaklytriangular at
retinacular tooth, the tooth of right mandibleminutely protuberant; apical
segmentsofmaxillaryandlabialpalpitumid;ligulanarrow,triangularlyproduced
at apical corners, fused with parag1ossae just behind apex; parag1ossaewell
prolonged forwardsbeyondligularapex,narrowat theprolongedparts;mentum
narrow, with epi1obesstrongly widenedapicad; microsculpture invisibleunder
80x magnification.
Pronotumsubsquare,weaklycontractedbehind,widestatapicaltwo-fifths,
one-fourth wider than long, relatively convex, smooth on disc, coarsely and
moderately punctate in lateral furrows and basal foveae; sides rather thickly
bordered, gently curved in front; apex evenly and shallowly emarginate; base
three-tenthswider thanapex,weaklyarcuateatsides;bothbordersofapexand
basecomplete;basalanglesmoreor lessobtuseandangulate,withaveryminute
protuberanceateachtip;lateral furrowsnarrow,graduallywidenedbehind,linked
withbasal foveae,whichareill-defined, ratherwideandbearsmallgroovesnear
innersides;fronttransverseimpressionhardlyobservable,hindonealittleclearer
than the front one; median line fine, clear and reachingboth apex and base;
microsculpture detected only in the furrows and foveae,consistingofobscure
transversemeshes.
Hindwingsfullydeveloped.Elytraoblong-oval,two-thirdslonger thanwide
and three-tenths wider than pronota1width, uniformly well convex, without
punctures;sidesgentlyarcuateinhumeri,shallowlysinuatebeforeapices;apices
weaklyproduced,narrowlyroundedattips,closetoeachother,angularlyrounded
atsutural angles;basesshallowly emarginate, forminganobtuse, angulateand
edentateanglewitheach side;striaedeep, wideand clearly crenulate,scutellar
stricto comparatively long than usual; intervals rather convex even on disc,
NoboruITO
38
becomingmoreconvexapicadandlaterad,3rdintervalwith foursetiferouspores
along2ndstria;marginalserieswidelyinterruptedinmiddle,composedof8十9
umbilicatepores;microsculptureunobservable.
Ventralsurfacemostlysmooth,withseveralpunctureslaterallyinmetaster-
num,not pubescentexcept forprosternumand2nd to6thabdominalsegments
sparselypubescent;metepisternumwellnarrowedbehind, two-thirdslongerthan
wide;6thabdominalsegmentin bisetoseoneachsideandsubtruncateatapex.
Hind femur bisetoseonhindmargin; fore tibiaslender, deeply incised in
externalhalfofapex,bispinousapico-externa11y,with a clear sulcus on dorsal
surface;foretarsislendererthanusual,1stsegmentofmidtarsuswithoutadhesive
biseriatesquamae,hindtarsusone-tenthlongerthanthewidthofhead,1stsegment
alittleshorterthanthe2ndand3rdtogether(0.93inratio)andoneandthree-fifths
the 2nd, 3rd one-third longer than the4th, claw segment bisetose along each
ventral margin.
Aedeagus (Fig 8) gently arcuate, more or less thickened and obliquely
hookedaboveat apex;apical lobesquare, roundedat tip; innersacarmedwith
agroupoffivelargeconicalscleritesonrightside,oneofsmallconicalsclerites
andofnumerousslender peg-shapedscleritesonleftsideandoneof largepeg-
shapedscleritesnearbasalpart.
Length:7.7mm.Width:3.0mm.
8
Figs. 8-9. GenitaliaofOxycentrusspp. - 8, 0. (0.?ycentropsls)g,aciiitarsissp nov;9,
0. (0xycent1・ops1's)acutipennissp nov;a, dorsalview.
SomeSpeciesoftheGenusOx;ycentt・us 39
Unknown.
Holotype: 300-400m, KaengKrachan Nat.Pak.,Phetchaburi,Thailand,
,
17-XI-1985, BURcKHARDT & LoBL leg. (preserved in the Museum d'Histoire
nature11e,Geneve).
This new species is somewhat similar to 0)cycentrus (s. str) algutoroides
(BATES),but iseasilydistinguished fromthelatterbytheantennaeslenderer, the
pronotumwider, theapicesof foretibiaemoredeeplyincisedinexternalhalves
andthetarsislenderer,besidesthesubgenericcharacteristics.
Orycentrlls(0xycentro-s')aeMtl'penm'ssp
nov.
(Figs 4,9)
This new speciesresembles Oxycentrus (0xycentropsts)gracilitarsis, but is
different fromthelatter inhavingthebodysmaller,theeyeslessprominent, the
pronotum lesstransverseandsharper at basal angles,and thesuturalanglesof
elytra not rounded. Thenew species isalso allied to Ox;ycentrus(s.str)melas
(ScHMIDT-GoBEL), but is distinguished from thelatter, inaddition to thesub-
genericcharacteristics,bythesidesofpronotumnotsinuatebeforebase,theelytra
not toothedathumeralanglesandwithsharpsuturalangles,andthetibiaemuch
slenderer.
Pronotumsubqudrate,weaklycontractedforwardsandbackwards,withsides
rather thickly bordered,sparsely punctate in basal foveae. Hind wingsentire.
Elytrasubpara11e1-sided, ratherconvex,notpunctate;scutellarstrictoveryshort.
Sixthabdominalsegment in bisetose on eachside.
Aedeagus (Fig 9)slender, almost straightly prolonged distad; apex thin,
knob-shapedat tip;apicalorificewidelyopen,innersacarmedwith twogroups
ofscleriteswhichareconicaland ratherelongate.
Length:6.9mm. Width:2.6mm.
Unknown.
Holotype: Tonkin, with label inscribed 0.xycentrus deb出sScHAuB.,
a
,
(preserved in the Landesmuseum, Linz).
要 約
伊藤 昇: 0;xycentrus属の数種について. - 0xycentrus(s.str)acutulusBATESを再記載した.
また, タイからOx),centrus(s.str)baehriおよび0.(0xycent・topsls)gracitita,s:isをそれぞれ記載し
た.前者は, 0.acutulusにきわめてよく似てぃるが,触角がより細いこと, 前胸背板側縁がよ
り強く狭まること, 上通がより短いことなどで区別できる. また後者は, Ox;ycentrus(s.str)
argutoro,dos(BATES)に似てぃるが, 触角および付節がより細いこと,前111?節先端の外側が深く
切れ込むことなどで区別できる. さらに, ベトナム(Tonkin)より0xycentrus(0xycentropsis)acu-
tipenn,sを記載した. この極は, 0.grac流tarsisに似てぃるが, より小型なこと,前胸背板はよ
り横長で後角がより鋭いこと, 上通会合線先端角が鋭いことなどの点で異なる. 0x:ycentro-
40 NoboruITO
ps,s亜属の種は,従来マレイ半島からスンダ諸島にかけての地域だけから知られていたが,今
回インドシナ半島より初めて記録された.
References
BATES, H w.,1892. viaggiodiLeonardoFEAinBirmaniaeregioni vicine.XLIV. List of the
Carabidae.Annli. Mus.c、ll. Stet・.nat.Get1ova,(2),12:267-428.
ITO,N.,1993. StudyonAsianCarabidae.VI.Speciesofthegenus0.,cycentrusCHAUDOIR(1).T「ans.
Shikokuent.Soc.(20):51-59.
KIRscHENHoFER,E.,1992. StudieniiberdiemitOxycentruspara1/eluSCHAUDOIR1854Ve「Wandten
Arten(Col.Carabidae,Harpalini:Harpalina).Linzerbio1.Beit1'.,24:35-58.
NooNAN,G.R.,1985. ClassificationandnamesoftheSelenophoriGroup(Coleoptera:Carabidae:
Harpalini)andofninegeneraandsubgeneraplacedinincertaesediswithinHarpalina.Milwaukee
Pub. Mus.Contr. Biol. &Goof.,(64):1-92.
ScHAUBERGER,E.,1934. ZurKenntnisder indo-orientalischenHarpalinen(FiinfterBeitrag). Ent.
Anz..14:89-92.
Elytra,Tlokyo,24(1):40,May15,1996
NewRecordofStaphylinidBeetlesfrom
Minamidaito-jima Island, theRyukyus
Yasuaki WATANABE
LaboratoryofEntomology,TokyoUniversityofAgriculture,
Setagaya,Tokyo, l56Japan
Inthepreviousshortreport (WATANABE,1991,Elytra,Tokyo,19,p.18),SiXSpecies
ofstaphylinidbeetleswererecordedfromMinamidaito-jmi aIsland,theRyukyus.
ThroughthecourtesyofDr.K.KuRosA,Tokyo,somestaphylinidbeetlesobtained
onMinamidaito-jimaIslandweregiventome.Theycontainedfourspecies,threeofwhich
arenew to the faunaofthisisland.Theyarerecordedbelow,allhavingbeenobtainedby
Mr.J.0KUMAonJune13th,1977,atIkenosawaonMinamidaito-jimaIsland,theRyukyus.
I thankDr.K.KuRosAforhiskindnessingivingmethespecimens.
1. Phi1onthusdiscoideusGRAvENHoRsT,1?.
2. Ph11onthusrectangulusSHARP,2
.
3. Atanygnathusterminalis(ERIcHsoN),1(;.