Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 International Journal of Current Microbiology and Applied Sciences ISSN: 2319-7706 Volume 7 Number 03 (2018) Journal homepage: http://www.ijcmas.com Review Article https://doi.org/10.20546/ijcmas.2018.703.286 Soilless Cucumber Cultivation under Protective Structures in Relation to Irrigation Coupled Fertigation Management, Economic Viability and Potential Benefits-A Review Mahesh Chand Singh1*, Dilip Singh Kachwaya2 and Kapil Kalsi2 1Department of Soil and Water Engineering, Punjab Agricultural University, Ludhiana (PB)-141004, India 2Dr. Y S Parmar University of Horticulture and Forestry, Nauni, Solan (HP)-173230, India *Corresponding author A B S T R A C T The protected c ultivation of cucumber in soilless systems (substrate culture, hydroponic, Key words aeroponic etc.) under optimal operating microclimatic conditions has become an alternative for sustainable vegetable production as a result of shifting climatic scenarios Cucum ber, and soil related problems. The soilless cultivation in both open and closed systems with Fertiga tion, Potential efficient utilization of water and nutrients has significantly increased the greenhouse benefi ts, Protected cucumber productivity compared to conventional cultivation under both protective cEuclotinvoa mtioicn ,v Siaobiilllietsys , structures and open field conditions. A closed hydroponic system has helped in saving A rtic le Info water and nutr ients thereby increasing water and nutrient use efficiency of cucumber cultivation with no environmental pollution. Further, the year round cultivation of Accep ted: cucumber has become possible under protective structures making it as an economically 20 Fe bruary 2018 viable technolo gy for the growers. Here, an effort has been made to highlight the progress Avail able Online: made in soilless or hydroponic cucumber cultivation in past in relation to irrigation 10 M arch 2018 coupled fertigation management, economic viability and the potential benefits. The study also describes the methods for computing the components of water and nutrient use balance in both open and closed systems. Introduction 22.0°C and below 27.0°C (Singh et al., 2017a). It grows productively under Cucumber (Cucumis sativus L.) is one of the conditions of high humidity, high light, most popular vegetable crops cultivated fertilizers and moisture in plastic greenhouses broadly throughout the world (Soleimani et (El-Aidy et al., 2007). It can be planted by al., 2009) under plastic greenhouses for higher direct seeding or transplanting. The row to economic value in off-season cultivation row spacing is often kept from 120.0 to 150.0 (Chandra et al., 2000). It is thermophilic in cm and plant to plant spacing varies from 30.0 nature and frost susceptible crop (Bacci et al., to 45.0 cm. The suitable plant spacing and 2006), growing best at a temperature above pruning results in higher fruit yield of 24 51 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 cucumber (More et al., 1990). Cucumber is Effect of growing media on crop production cultivated for its tender fruits, which are consumed either raw as salad, cooked as The major limitations of protective cultivation vegetable or pickled in its immature stage are soil borne diseases (Baysal-Gurel et al., (Sumathi et al., 2008). For fresh consumption, 2012). The soilless cultivation is therefore, a cucumber fruits are picked at their full growth possible alternative for sustainable vegetable stage before attainment of physiological production, which reduces the soil related maturity (Kanellis et al., 1986) However, the problems experienced in the conventional crop quality and production of cucumber is now in cultivation (Olympious, 1995; Hussain et al., threat under changing climatic scenarios, 2014). Moreover, soilless growing media are growing media (occurrence of soil-borne easier to handle and may provide a better diseases on global basis) and the microclimate growing environment compared to soil within plant community. (Mastouri et al., 2005). Soilless cultivation also offers other benefits such as capability to Effect of climate change on crop production control water availability, pH and nutrient concentration in the root zone (Epstein and The climate change which is one of the main Bloom, 2005). Cocopeat is one of the soilless determinants of agricultural production has media which is suitable for cucumber started affecting the pattern of crop growth production. The major limitations of from last few couple of decades in various protective cultivation are soil borne diseases agro-climatic zones globally (Wang et al., (Baysal-Gurel et al., 2012). The soilless 2014). The production and quality is affected cultivation is therefore, a possible alternative directly or indirectly due to increased for sustainable vegetable production, which temperature and exposure to elevated levels of reduces the soil related problems experienced carbon dioxide. According to Minaxi et al., in the conventional crop cultivation (2011), the expected rise in average surface (Olympious, 1995; Hussain et al., 2014). temperature may be in the range of 1.1°C- Moreover, soilless growing media are easier to 6.4°C by the last decade of 21st century on handle and may provide a better growing global basis. Any change in climatic factors environment compared to soil (Mastouri et al., such as temperature is bound to have a 2005). Soilless cultivation also offers other significant impact on crop growth and benefits such as capability to control water production (Singh, 2016). In India, climate availability, pH and nutrient concentration in change has increased the intensity of natural the root zone (Epstein and Bloom, 2005). calamities such as floods, droughts, heat Cocopeat is one of the soilless media which is waves and cyclones (Goswami et al., 2006). suitable for cucumber production. The present The problems would further aggravate in the study was thus undertaken to study the years to come due to heavy dependence on nutrient and water use efficiencies of agriculture and limited arable land if no greenhouse seedless cucumber production in corrective measures are taken today. Apart soilless media under partially controlled from this, the application of agrochemicals greenhouse conditions. such as fungicide has also limited the production of high value fruits and vegetable The soil borne diseases both in protective crops such as cucumber (Bagi et al., 2014). structures and open field conditions have Numerous factors which affect the cucumber limited the cucumber cultivation (Baysal- growth and productivity to a significant degree Gurel et al., 2012). Under such circumstances, are described in Singh et al., (2017a). the soilless cultivation can be practiced for 2452 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 sustainable vegetable production, with a radiation of short wavelengths to create an significant reduction in soil related problems encouraging microclimate for plant growth experienced in the conventional cultivation and higher productivity (Tiwari 2006). In the (Olympious, 1995; Hussain et al., 2014). present scenario of continual demand of Furthermore, soilless growing systems vegetables and dwindling land holdings (substrate culture, hydroponic, aeroponic or drastically, protected cultivation of vegetable aquaponic etc.) may provide a better growing crops is also becoming popular in the hilly environment and are easier to handle regions of the country, which offers a great compared to soil (Mastouri et al., 2005). It scope for use of low cost naturally-ventilated also offers other benefits such as capability to polyhouses or greenhouses because of mild control pH and nutrient concentration in the climate (Mishra et al., 2010). Thus, protected root zone and water availability (Epstein and cultivation is now very much needed under Bloom, 2005). Indian conditions to improve the productivity and quality of the vegetable crops (Kohli et Microclimate within plant community al., 2007). The productivity of cucumber grown inside According to Spehia (2015), the productivity protective structure is extremely dependent on of cucumber inside a polyhouse is more than light, humidity, temperature, CO , irrigation four times of that obtained under open field 2 water, fertigation, method of cultivation and cultivation. Kumar et al., (2011) reported a 2.5 cultivars. The crop productivity also chiefly times increase in vegetable production from depends on the response of plants to 1991-92 to 2010-11 in India. FAO (2011) has environmental conditions. According to reported a cucumber production of 1.60 lakh Marcelis et al., (2005), the yield of vegetable tonnes with an average productivity of 6412.0 crops reduces by 0.8-1.0 percent for 1.0 kg ha-1 from an area of 25104.0 ha in India. percent reduction of solar radiation reaching The worldwide area under protected the plant canopy. From biomass production cultivation has increased significantly during and energy saving point of view, an air past couple of decades (Singh et al., 2016). temperature of 19.0°C for daytime and 15.0°C India, from where the cucumber originated is for night time and a canopy leaf area index not among the leading global (LAI) of 2.0-3.0 provides the most energy producers. However, the cucumber efficient conditions for greenhouse cucumber productivity and fruit quality in soil under cultivation under the winter climate conditions protective structures has been significantly (Luo et al., 2005). It is thus important to reduced due to soil-borne dieases (Hussain et cultivate cucumbers under protective al., 2014). Thus, soilless cultivation can be structures under optimal operating adopted for sustainable vegetable production. microclimatic conditions for improved yield and quality of fruit. Soilless cultivation Protected cultivation The soilless cultivation is a technique of growing crops without soil as rooting media A protective structure when supported with (Savvas et al., 2013). It has been favorable climatic conditions offers off-season acknowledged internationally due to its ability cultivation of vegetable crops which is more to maintain efficient and intensive plant profitable in the Northern Plains of India (Nair production (Barrett et al., 2016). It has and Barche 2014). A greenhouse traps solar become progressively more important 2453 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 worldwide during last fifty years capacity, EC and pH value can be determined (Schmilewski 2009). In recent past, numerous individually by different methods suggested in researchers cultivated cucumbers in soilless literature one of which is reported in media under different greenhouse conditions Verdonck and Gabriels (1992). EC and pH (Huber et al., 2005; Gul et al., 2006; Gul et values are the most sensitive parameters which al., 2007; Janapriya et al., 2010; Zhang et al., directly affect the water and nutrient uptake of 2012; Mazahreh et al., 2015). Subjected to the plants which in turn can limit the plant optimal operating microclimatic conditions for growth and development. greenhouse cucumber cultivation in soilless media (Singh et al., 2017b), desired yield Among several growing media, coco-peat is potential of cucumber can be achieved. considered as a wonderful growing media with suitable EC, pH and other chemical properties. Optimal microclimate for cucumber Coco-peat has high water holding capacity and cultivation its high air filled porosity results in very high seed germination rate and produces more The assemblage of climatological parameters stronger and fibrous seedlings (Fornes et al., forming around a living plant inside a 2003). According to Ghehsareh et al., (2011), protective strucutre is termed as greenhouse the water holding capacity (WHC) of coco- microclimate (Singh et al., 2016). Thus, it peat is 90.5 percent, while other substrates becomes important to maintain the such as perlite and date-palm peat are having microclimatic parameters to a desired range WHC of 96.7 percent and 78.3 percent for better crop growth and productivity respectively. through commonly used techniques viz. natural ventilation, shading and evaporative Nutrients requirement of cucumber cooling under summer climatic conditions (Singh et al., 2018) and heating in winter The cucumber plants grown in soilless media climatic conditions. Most recently, Singh et require a continuous supply of nutrient al., (2017b) reported the desirable range for solution containing both macro and air temperature in plant community, leaf micronutrients in appropriate proportion based temperature and root-zone temperature as on crop growth stage. The elements viz. 22.0-27.0°C, 20.5-25.1°C, 16.9-22.9°C carbon (C), hydrogen (H) and oxygen (O) are respectively. According to Singh et al., also essentially required by the plants and (2017b), relative humidity, solar radiation and obtained from water and air. The nutrients vapour pressure deficit should lie in the range (macro) are categorized as primary and of 60.0-85.0 percent, 100.0-169 Wm-2 and secondary macronutrients. The primary 0.53-1.10 kPa respectively for optimal plant macronutrients included nitrogen (N), growth and development. phosphorus (P) and potassium (K) and the secondary macronutrients included calcium Properties of soilless growing media (Ca), magnesium (Mg) and sulphur (S). The micronutrients included boron (B), iron (Fe), The knowledge of physiochemical properties manganese (Mn), copper (Cu), zinc (Zn) and of growing media is essential for limiting molybdenum (Mo). The micronutrients are them to desired range in order to offer better required by plants in small quantities. The plant growing conditions. These properties water soluble fertilizers are now easily viz. particle density, bullk density, air filled available in market and can be used as the porosity, total porosity, moisture content, field sources of macro and micronutrients. 2454 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 Preparation of nutrient solution the yield penalty (Sonneveld and Voogt 2009). The EC and pH values of nutrient solution and The nutrient solution can be prepared on daily growing media can be monitored on daily basis if possible or stock solution (hogland) basis using water proof testers. can be prepared according to literature (Hoagland and Arnon 1950; Sonneveld and Method of irrigation coupled fertigation Straver 1994). The prepared nutrient solution should be monitored on regular basis for EC, Drip irrigation is the most suitable method for pH value and deficiency of micronutrients as irrigating vegetable or fruit crops in both soil suggested by Shah et al., (2009). and soilless media. It is considered as a tool for increasing water and nutrient use Disposal of leachate efficiency (NUE) of the crop grown when managed carefully. It maximizes yield with a Suitable arrangement should be made to safely significant reduction in input water and dispose off the leachate coming out of the environmental pollution by improving the growing media in open system. In closed photosynthetic capacity of the plants. It system, the leachate is recirculated and reused minimizes the leaching of nutrients or causing no environmental pollution. chemicals from root-zone system of the plants (Gardenas et al., 2005). Numerous studies pH and EC of nutrient solution reported the positive effect of drip irrigation in improving yield of the crop grown (Ertek et The parameter pH is directly related to al., 2006; Vazquez et al., 2006; Castellanos et nutrient uptake by plants and should be al., 2013). adjusted to desired range for optimal plant growth. Several chemicals can be used to Crop water requirement adjust the pH of nutrient solutions. The crop water requirement (CWR) can be The most popular are phosphoric acid to lower estimated from crop transpiration under a pH and potassium hydroxide to raise pH. closed soilless or hydroponic system. The crop Other chemicals viz. mono potassium transpiration can be computed using the phosphate, nitric acid and sulfuric acid can formula derived from the Penman-Monteith also be used to lower pH. However, nitric acid equation (Medrano et al., 2005) with certain and sulfuric acid are much more dangerous modifications in model coefficients (equation than phosphoric acid. The pH value of the 1). Transpiration helps to improve irrigation nutrient solution should be kept in the range of control in soilless cultivation of crops under 6.0-6.40. greenhouse conditions as it is directly related to plant production (Watts and Goltz 1985). On the other hand, the increased EC value of Numerous studies related to transpiration from nutrient solution may consequently increase cucumber crop (evapotranspiration) have been the acidity (Adams and Ho 1989) thereby reported in literature (Medrano et al., 2001; affecting the crop growth and development. Weihong et al., 2004; Medrano et al., 2005; Cucumber is moderately salt-sensitive among Zhang et al., 2010). the vegetable crops having threshold EC value of 3.0 dS m-1 (Singh et al., 2017a). The A simplified model derived from Penman- preferable EC of nutrient solution is 2.7 dS m- Montheith equation with a few modifications 1 and should never exceed 3.0 dS m-1 to avoid in model coefficients for specific cop and 2455 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 climatic conditions can accurately predict the filters for each supply before its delivery to the hourly transpiration rate for soilless cucumber plant root system to prevent the clogging of crop (Medrano et al., 2001, 2005). Weihong et emitters. The nutrient solution is supplied al., (2004) also computed the canopy directly to the plants root system through transpiration of cucumber crop using Penman- emitters operating at a desired discharge rate. Monteith model. Medrano et al., (2005) evaluated the transpiration time course of Computation of water and nutrient balance cucumber plants cultivated in soilless culture during two cycles at low (up to 9.0 M J m-2 d- In soilless cultivation, the quantity of nutrient 1) and high (up to 20.0 M J m-2 d-1) radiation solution applied to the crop on daily basis can levels. Zhang et al., (2010) studied the be measured by means of measuring cylinders relationship between crop evapotranspiration installed at various locations. (ET ) of cucumber and climatic factors inside c a solar greenhouse. Volume of water application A(1ekLAI)I BLAIVPD V V T rad The total volume ( nsapp or wapp) of nutrient (1) solution applied per plant per day in a given time (t) can be calculated for a known Where, T = Crop transpiration (kg m-2 s- discharge rate of an emitter using the I following relationship. 1), rad= Solar radiation (W m-2), VPD =Vapour pressure deficit (kPa), k = V nqt Extinction coefficient,LAI = Leaf area index wapp (2) (m2 m-2), = Heat of vaporization of water (J n Where, = Total number of emitters kg-1), A= Equation parameter (dimensionless) operating, q= Emitter discharge (litre hr-1) andB= Equation parameter (W m-2 kPa-1) t and = Time to irrigate or fertigate (hr) The parametersAand Bare different for V V different seasons. If wuppand wdppbe the volume of water uptake and water drained per plant, the water Fertigation system balance can be expressed as V V V V ws wapp wdpp wupp (3) A drip fertigation system generally includes main pipeline, laterals and emitter pipe for V supplying nutrient solution to the plants. The Where, wapp= Volume of water applied per system also includes tanks for preparation of V plant (litre plant-1) and ws= Water stored in nutrient solution, electric motors or solar growing media on volumetric basis. system, pressure gauges, timers, filters and emitters. The operating pressure of emitters However for a closed hydroponic system can be regulated using pressure gauge to where only plant roots are in contact with deliver the nutrient solution safely and at V desired rate to the root zone of the plants. The running nutrient solution, wscan be taken as timers allow the application of nutrient zero and the equation 4 becomes as solution for a pre-determined time. The V V V 0 nutrient solution should be passed through wapp wdpp wupp (4) 2456 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 Nutrient balance dw nutrient in fruits (ppm), L= Dry weight of dw For measurement of leachate coming out of leaves (mg plant-1), S= Dry weight of stems the growing media specially designed trays (mg plant-1),dwF= Dry weight of fruits (mg can be installed at different locations in the plant-1). experimental trial. The collected leachate or ND waste nutrient solution can be measured using The nutrient drained can be computed as pp a measuring cylinder on daily basis and can be NA NU = pp pp (7) recycled for further use in a closed system. Having known the total volume of nutrient ND solution applied on daily basis and the Where, pp= Total nutrient drained per concentration of each nutrient in the applied plant (mg plant-1) nutrient solution, the quantity of applied nutrient to the crop can be computed as The total nutrient applied, plant uptake and NA c V nutrient drained can be computed on per plant pp ans nsapp (5) and per hectare basis (kg ha-1). The nutrient balance in an open soilless growing media can TNA Where, pp = Total nutrient applied per be expressed as c plant (mg plant-1), ans= Concentration of a N NA NU ND nutrient in the applied nutrient solution (mg sgm pp pp pp (8) V litre-1) and nsapp= Total volume of nutrient N solution applied per plant (litre plant-1) Where, sgm= Nutrient storage in growing media (mg plant-1) The nutrient concentration of plant can be computed analytically using inductively However, the nutrient balance for a closed coupled plasma atomic emission hydroponic system can be expressed as spectroscopy (ICP-AES) except for nitrogen (Grewal et al., 2011). The oven dried samples NA NU ND 0 pp pp pp (9) of leaves, stems and fruits are used for determination of nutrient elements. The Pollination concentration of nitrogen present in plant in amonical form can be computed using In cucumber, the male flowers emerge first standard ethods such as Kjeldahl digestion and female flowers shortly later. A female method. The total plant uptake of the nutrient flower contains a small immature fruit at the for each element can be computed as base while the male flower does not contain NU c dw c dw c dw any. Pollen is transferred from male to female pp L L S S F F (6) flower by bees or other insects. After proper pollination the female flower develops into a NU fruit. However, the parthenocarpic cucumber Where, pp = Nutrient uptake per plant (mg hybrids produce fruits without pollination. plant-1), cL=Concentration of a nutrient in Greenhouse cucumbers are naturally c parthenocarpic and the fruits produced are leaves (ppm), S = Concentration of a nutrient seedless. c in stems (ppm), F= Concentration of a 2457 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 Disease management The recommended fertilizers for fertigating cucumbers in soil and soilless systems can be The main entry gate of the greenhouse under taken from Anon (2018a). The fertigation experimentation should be provided with an schedule or nutrients (macro and micro air curtain to create a barrier between outside nutrients) required for preparing nutrient and inside air and blow away the insects solution to fertigate soilless (hydroponic and entering the greenhouse during entry of a substrate based) cucumbers can be taken from personal. The insect net (if any) of the Anon (2018b) or Anon (2018c) or greenhouse should be washed with a spray of Papadopoulos (2001). water under adequate pressure in order to remove insects attached to it before Water and nutrient use efficiency transplanting. The yellow sticky cards can be installed in the whole greenhouse at various The water use efficiency decreases with locations for controlling the whiteflies increase in irrigation water applied from incidence to the crop and a spray of Polo fruiting to the end of growth stage and (Diafenthiuron 50.0 % WP) can also be done increases with increase in irrigation water before date of transplanting. The growing from cucumber fruit setting to start of fruit media should be disinfected before using it for ripening (Mao et al., 2003). According to growing cucumbers next growing season. Wang et al., (2009) the IWUE decreased with Fungicides such as Ridomil Gold of Syngenta decrease in water input which also agrees with which is a mixture of systemic and contact Ayas and Demirtas (2009). A continous effort fungicide (Metalaxyl-M4.0%+Mancozeb has been made in the past to study the effect of 64.0% WP) can be used for disinfecting the irrigation (Sanchez-Guerrero et al., 2009; growing media. Grewal et al., 2011) on cucumber growth, yield and quality. Irrigation coupled fertigation The greenhouse seedless cucumber has a high Water and nutrients are two vital inputs for nutrient requirement and is very productive plant growth particularly in soilless media and with adequate supply of nutrients (Sonneveld their uptake by plants are two independent and Voogt 1978). Drip fertigation allows processes. Cucumber is considered to be precise and uniform distribution of water and sensitive to drought stress and should never be nutrients exactly to the root system of plant at short of water (Loomis and Crandall 1977) right time. According to van Os (1999), a and nutrients. Fertigation in soilless soilless system has higher water and nutrient cultivation provides efficient use of water and use efficiencies [IWUE or CWUE = yield (kg nutrients (Jensen 1997) through precise and plant-1)/irrigation water applied (m3 plant-1) uniform application of nutrients directly to the and NUE = yield (kg plant-1)/nutrient applied active root system of the plant (Rouphael et (g plant-1)]. A continous effort has been made al., 2008). Numerous researchers studied the in the past to study the effect of fertigation effect of irrigation amount (Mao et al., 2003; (Gul et al., 2007) on cucumber growth, yield Amer et al., 2009; Salcedo et al., 2017) and and quality. fertilizer coupled irrigation i.e. fertigation (Papadopoulos 2001; Lee et al., 2005; Ahmet While comparing the effect of open and closed et al., 2006; Zhang et al., 2011) on cucumber systems on cucumbers cultivated in vertical yield and water use efficiency (WUE). and horizontal bags fertigated with applied and drained nutrient solution, Tuzel et al., 2458 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 (1999) confirmed a 22.0 and 35.0 % reduction opportunities in recycling drainage water to in water and nutrient consumption under increase water and nutrient-use efficiency and closed system. According to Papadopoulos reduce the environmental impact of the (2001), the optimal nutrient application to a drainage water discharge. Salcedo and Reca crop throughout the entire cropping season in (2017) reported a good correlation between relation to changing needs of the plants for the crop water uptake (WU) and the leaf area nutrients is highly successful due to a greatly index (LAI) while evaluating the irrigation improved understanding of plant nutrition and water consumption of a soilless cucumber the availability of sophisticated nutrient crop under greenhouse conditions under a delivery systems. Huber et al., (2005) humid tropical climate in order to improve the cultivated cucumbers in a specially designed irrigation water and fertigation management. urethane based recyclable plant growth substrate (UBS) and rockwool (RW) under a Effect of growing media on nursery raising recirculating hydroponic system and and fruit quality of cucumber confirmed that irrigation was extremely important for crop production with a potential Selection of a suitable growing media is for UBS to evolve as a competitive plant extremely important as it influences the growth substrate for greenhouse soilless crop quality of seedlings produced. The growing production with some design modifications. media influences the performance of the ready Lee et al., (2005) reported increased plant seedlings before transplanting in the field and weight, number of leaves and total yield the performance of seedlings in the main field including marketable yield with increased after transplanting is evaluated based on their application of nutrient solution from 0.5 to 2.0 performance in the nursery raising in relation litre plant-1day-1 obtaining highest fruit yields to growing media and microclimatic at 1.5 and 2.0 litre plant-1 day-1 of fertigation conditions offered. According to Luoto level. While evaluating the effect of nutrient (1984), the colour of cucumber is affected by sources, organic manure and inorganic growing media at the end of harvest season. conventional nutrient solution on cucumber The quality of cucumber fruits can be tested cultivation in different locally available by determining texture, colour and taste by substrates, Gul et al., (2007) indicated a yield either chemical analysis or sensory evaluation reduction of 10.8% and 31.3% under organic (Luoto 1984). nutrient solution and solid organic manure compared to inorganic nutrient solution. Cucumber contains about 95.2% water, 1.7% sugar and 2.8 mg (100g)-1 vitamin C. Melo et While testing an EC-based irrigation strategy al., (2006) have reported the polyphenol in two greenhouse soilless cucumber crops concentration of 9.05±0.83 mg (100g)-1 and grown under Mediterranean conditions, ascorbic acid concentration of 1.49±0.85 mg Sanchez-Guerrero et al., (2009) concluded (100g)-1 in cucumber. Firmness is one of the that CO enrichment combined to an EC-based major quality parameters of pickling 2 irrigation scheduling improved the overall cucumbers (Suojala-Ahlfors 2005). However, water use efficiency of soilless greenhouse the transpiration from the cucumber fruits cropping systems with a drastic reduction of detached from the plants considerably reduces the leaching fraction. Grewal et al., (2011) the fruit water content and therefore the fruit confirmed a 33.0% reduction in potable water firmness. The chlorophyll present in the used for irrigation in hydroponic cucumber epidermis is responsible for dark green colour production while investigating the on the surface of cucumber fruit. The 2459 Int.J.Curr.Microbiol.App.Sci (2018) 7(3): 2451-2468 chlorophyll breakdown results in immediate a significantly higher fresh weight of degreening of cucumber fruit and the exposure cucumber plant grown in peat compared to of cucumber plants to heat stress during fruit that in perlite. Alifar et al., (2010) obtained development stage causes bitterness of fruits. the highest and lowest yield of cucumber fruit The nutritional and calorific value of from cocopeat and perlite-cocopeat by cucumber is very low, but it is a primary investigating the effect of five different source of vitamins, minerals and fibre for growing media viz. cocopeat, perlite: cocopeat human body. It is also a wonderful source of (50:50), perlite: cocopeat: peatmoss (50:20:30 carbohydrates and phosphorus (Yawalkar and 50:30:20) and perlite: peat moss on 1985). volume basis. Janapriya et al., (2010) reported highest yields of 113.9 t ha-1 and 96.1 t ha-1 in Soilless cucumber yield greenhouse and open field cultivation of cucumber under the treatment peat: The cucumber cultivation in soilless system vermicompost: sand with a highest benefit- has significantly increased during last couple cost ratio of 3.4 with 100.0% drip fertigation of decades (Gul et al., 1999; Lorenzo et al., inside the greenhouse. 1999; Tuzel et al., 1999; Huber et al., 2005; Al-Mulla et al., 2008; Janapriya et al., 2010; Ghehsareh et al., (2012) reported higher yield, Zhang et al., 2012; Mazahreh et al., 2015) biomass, plant height, root weight, leaf area than soil based cultivation due to occurrence index (LAI) and total soluble solids (TSS) of of soil borne diseases. Hickman and Klonsky cucumber fruit on using date-palm as growing (1993) reported a yield of 33.0 kg m-2 for media compared to the conventional soil cucumber cultivated in bag culture inside a system. While evaluating the weathered spent greenhouse. Gul et al., (1999) reported no mushroom substrate (SMS) as a growing significant difference in plant growth, yield medium for nursery raising of cucumber, and fruit quality both in open or closed Zhang et al., (2012) indicated a better systems while cultivating cucumbers in peroformance of plant height, leaf area, fresh vertical and horizontal bags containing perlite weight, dry weight and seedling quality under as growing media in open and closed systems. SMS: vermiculite of 2:1 and SMS: perlite of 4:1 growing media compared to peat and However, the plant growth and yield were perlite (1:1). While studying the effect for considerably higher in bags placed different growing media viz. perlite (1:0), horizontally. Lorenzo et al., (1999) obtained a cocopeat grow bag, perlite: cocopeat (1:1) and lower yield of cucumber under mulching with perlite: cocopeat (1:2) on volume basis on white film than conventional soilless culture cucumber yield, Mazahreh et al., (2015) due to lower temperature of air and growing reported highest (87.6 t ha-1) and lowest (46.0 media in the mulched cucumber during the t ha-1) yield of cucumber under perlite: vegetative development phase, inducing a cocopeat (1:1) and cocopeat grow bag lower leaf area and LAI thereby reducing the respectively. interception of radiation by the cucumbers under mulching. Economic feasibility of cucumber cultivation Al-Mulla et al., (2008) reported higher number of cucumber fruits per plant cultivated The economic viability of cucumber in wood straw than date palm straw as cultivation under protective structures mainly growing media. Peyvast et al., (2008) reported depends on the basic cost of construction of 2460
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