SOCIOBIOLOGY AN D CON FLICT SOCIOBIOLOGY AND CONFLICT Evolutionary perspectives on competition, cooperation, violence and warfare Edited by J. VAN DER DENNEN Polemological Institute, University of Groningen, The Netherlands and V. FALGER Department of International Relations, University of Utrecht, The Netherlands S CHAPMAN AND HALL London. New York. Tokyo. Melbourne. Madras UK Chapman and Hall, 2-6 Boundary Row, London SE1 8HN USA Chapman and Hall, 29 West 35th Street, New York NY10001 JAPAN Chapman and Hall Japan, Thomson Publishing Japan, Hirakawacho Nemoto Building, 7F, 1-7-11 Hirakawa-cho, Chiyoda-ku, Tokyo 102 AUSTRALIA Chapman and Hall Australia, Thomas Nelson Australia, 480 La Trobe Street, PO Box 4725, Melbourne 3000 INDIA Chapman and Hall India, R. Seshadri, 32 Second Main Road, CIT East, Madras 600 035 First edition 1990 © 1990 Chapman and Hall Softcover reprint of the hardcover I st edition 1990 Typeset in 10/12 pt Plantin Light by Leaper and Gard Ltd, Bristol Printed in Great Britain by St Edmondsbury Press, Bury St Edmunds, Suffolk ISBN-13: 978-94-010-7316-5 e-ISBN-I3: 978-94-009-1830-6 DOl: 10.1007/978-94-009-1830-6 All rights reserved. No part of this publication may be reproduced, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording or otherwise, or stored in any retrieval system of any nature, without the written permission of the copyright holder and the publisher, application for which shall be made to the publisher. British Library Cataloguing in Publication Data A CIP catalogue record for this book is available from the British Library. Library of Congress Cataloging-in-Publication Data Available Contents Contributors vii Acknowledgements IX 1. Introduction 1 PART ONE Conflict and Biology 2. Intergroup competition and conflict in animals and man J.A.R.A.M. van Hoof! 23 3. Selfish cooperation in social roles U. Motro and D. Cohen 55 4. The biological instability of social equilibria P.P. van der Molen 63 PART TWO Sociobiology and Enmity 5. The cerebral bridge from family to foe L. Tiger 99 6. The evolutionary foundations of revolution J. Lopreato and P. Green 107 7. Loyalty and aggression in human groups Y. Peres and M. Hopp 123 8. Territoriality and threat perceptions in urban humans M. Hopp and o.A.E. Rasa 131 PART THREE 'Primitive' Warfare 9. Origin and evolution of 'primitive' warfare J.M. G. van der Dennen 149 10. The Inuit and the evolution oflimited group conflict C. Irwin 189 11. Human nature and the function of war in social evolution P. Meyer 227 12. War and peace in primitive human societies U. Melotti 241 13. Primitive war and the Ethnological Inventory Project ].M. G. van der Dennen 247 PART FOUR The Conflict about Sociobiology 14. The sociobiology of conflict and the conflict about sociobiology U. Segerstnlle 273 Bibliography 285 Author index 325 Subject index 335 Contributors Dan Cohen Professor of Ecology and Evolutionary Biology at the Department of Botany, the Hebrew University, Jerusalem, Israel Johan M.G. van der Researcher at the Polemological Institute of the Dennen State University of Groningen, The Netherlands Vincent S.E. Falger Lecturer in International Relations at the Univer sity of Utrecht, The Netherlands Penny Anthon Green Teaches at the Department of Sociology at the University of Texas at Austin Michael Hopp Member of the Hebrew University School of Nutrition and has a private applied social research institute Colin Irwin Held a Killam Post-Doctoral Fellowship at Dalhou sie University at Halifax, Nova Scotia, Canada, while preparing his contribution to this volume Joseph Lopreato Professor of Sociology at the University of Texas at Austin and a former Chairperson of his Department Umberto Melotti Professor of Political Sociology and Cultural Anthropology at the universities of Rome and Pavia, Italy Peter Meyer Lecturer of Sociology at the Economics and Social Sciences Department at the University of Augsburg, FRG Popko P. van der Molen Conducts research at the University of Groningen, The Netherlands Uzi Motro Senior Lecturer at the Department of Genetics and the Department of Statistics of the Hebrew University of Jerusalem, Israel Yochanan Peres Associate Professor of Sociology at Tel Aviv University, Israel Olwen Rasa Associate Professor of Animal Behaviour at the University of Pretoria, South Africa Ullica SegerstnUe Assistant Professor of Sociology at Illinois Institute of Technology, Chicago Lionel Tiger Professor of Anthropology at Rutgers University in New Brunswick, New Jersey Acknowledgements The Sociobiology ofC onflict was the topic of the ninth meeting of the European Sociobiological Society, held on January 10 and 11, 1987. It was Michael Hopp's initiative to organize this conference in Jerusalem, Israel, a symbolic place in many respects. Thanks to the scientific and personal qualities of Professor Amotz Zahavi, from Tel-Aviv University, many non-Israeli partici pants were able to experience the naturalistic, geographical and political history of the country in an impressive guided tour which influenced clearly the presentations and discussion in the conference. Without the hospitality and financial support of the Van Leer Jerusalem Institute the meeting would not have been possible. The Institute's director, Professor Yehuda Elkana, and Mrs Rivka Ra'am, member of the Executive Board of the Van Leer Jerusalem Institute, in close cooperation with local organizer Michael Hopp, contributed very much to the success of the meeting itself. The Board of the European Sociobiological Society expresses its gratitude for this vital support. In the conference itself Vincent Falger, Lea Gavish, Johan Goudsblom, Anne Rasa, Avi Shmida, Jan Wind and Amotz Zahavi presented papers next to those elaborated and collected in this volume. ESS Board members Jan Wind, Hans van der Dennen and Vincent Falger organized those aspects of the conference not immediately connected with the meeting in Jerusalem. Finally, the patience and trust of Tim Hardwick, former senior editor with Chapman and Hall, and his successor Bob Carling, were indispensable for this book to be published. It is fortunate that in human society cooperation is not less essential than conflict. This volume combines both, not surprisingly. CHAPTER ONE Introduction 1.1 THE STUDY OF CONFLICT Polemos Pantoon Pater Heraclitus Conflict on all levels of organic existence is pervasive, persistent, ubiquitous. Conflict is the universal experience of all life forms. Organisms are bound in multiple conflict-configurations and -coalitions, which have their own dynamic and their own logic. This does not mean, however, that the more paroxysmal forms of conflict behaviour, naked violence and destruction, are also universal. Conflict and cooperation are always intertwined. Conflicts do, however, have a propensity to gravitate towards violence. There is, as Pettman (1975) pointed out, no accepted or agreed list of the social units by which conflicts might be classified. To talk of conflict in intra personal, inter-personal, familial, group, class, ethnic, religious, intra-state or inter-state terms is to assume, perhaps erroneously, that 'each kind of social unit, having its own range of size, structure, and institutions, will also have its own modes of interaction and thus its own patterns of conflict with other social units' (Fink, 1968) like and unlike itself. Such an assumption merits scrutiny on its own, since, despite the plausibility of some sort of analytical link between the parties to a conflict and the nature of the confrontation that ensues, the link should be demonstrated and not allowed to stand by assertion alone. This volume is devoted to one type of analysis of conflict, the socio biological one. In The Sociobiology of Ethnocentrism, a book closely related to many of the ideas and some authors of this volume, sociobiology was defined as 'the branch of biology that concerns itself with the explanation of social behaviour in all species, including our own. It is thus, essentially, evolutionary biology, and relies on Darwinian, Mendelian and Hamiltonian ideas - concepts such as natural selection, genetics, and, especially the individual's inclusive fitness - for its underlying explanatory schema' (Reynolds, Falger and Vine, 1987). Inclusive fitness theory, first proposed by Hamilton in 1964 and repeatedly referred to in this book, shows that genes will spread iftheir carriers act to increase not only their own fitness or reproductive success but also that of 1 Introduction other individuals carrying copies of the same genes. A person's inclusive fitness is his or her personal fitness plus the increased fitness of relatives that he or she has in some way caused by his or her actions. It may sound very deterministic or even 'geneticistic' to try to explain social phenomena on the basis of some supposed underlying biological process, but that would be too restricted an interpretation of the effort to draw attention to biology as one place to look. What is encouraging about a recourse to biology is that there is a body of theory and a wealth of empirical data relating to other species. This provides an excellent background against which to compare and contrast human processes and situations, the main focus of attention here. None of the authors in the present volume is proposing to reduce the study of conflict to biology, to account for it simply as any simple instinct. But all are alert to the existence of similar processes in animals and are trying to use the theory of inclusive fitness to explain the evolution of these processes. However, a sociobiology of human behaviour would not be taken seriously if it explains in terms of fitness something that can be better explained in strictly cultural terms or in terms of the market. That is why sociobiology of any human behaviour can and must seek links with social psychology, social anthropology, sociology, economics, political science and even history. Beyond that - and this is very important - it would be quite pretentious to assert that sociobiology could solve all questions that have arisen since the phenomena of conflict were studied in a systematic way. To know what these questions are and why it makes sense to add a biological-evolutionary oriented approach to the study of conflict and competition, it is worthwhile to draw some broad historical lines. Therefore, this introduction aims literally at introducing the reader to the traditional scientific discourse on conflict, which usually means human conflict. Then a condensed overview on the (socio)biol ogy of conflict and competition tries to make the non-specialist familiar with the most relevant theoretical and conceptual problems in this field. Those readers, however, who want to cut short and demand an immediate answer on the question why a sociobiological perspective on conflict and competition is a valuable contribution, should skip the next three main parts of this intro duction and turn straight to its fifth part. 1.2 THE STUDY OF CONFLICT IN PERSPECTIVE Conflict may be defined as: incompatibility of interests, goals, values, needs, expectations, and/or social cosmologies (or ideologies). Ideological conflicts especially have a tendency to become malicious (cf. Berger and Luckman, 1966). Webster's Dictionary defines conflict as 'Clash, competition or mutual interference of opposing or incompatible forces or qualities (as ideas, interest, wills)'. Coser (1956) defined social conflict as 'a struggle over values and claims 2