A t l a s of H u m a n A n a t o m y Internal Organs 15th Edition v T'vV\i Edited try F Paulsen and J Waschko English Version with Latin Nomenclature 1 M MLR 1 HJU>* l«K)UA URBAN & FISCHER User's Guide to the Book Introductory pages: • Bulleted lists in figure captions as well as in tables help structu • The introductory pages provide all relevant anatomical informa ring complex facts and provide a better overview. tions concerning the subject of the chapter. Important details • Figures, tables, and text boxes are interconnected by cross- and connections are explained easily to understand. references. • The Dissection Link for each chapter comprises brief and con • Cross-references link the figures to the separate Table Booklet cise tips essential for the dissection of the respective body re with tables of muscles, joints, and nerves, thus providing a suf gion. ficient anatomical knowledge for the exam. • Exam Check Lists provide all keywords for possible exam • Clinical Remarks boxes provide clinical background knowledge questions. concerning the anatomical structures illustrated on the page. • The dissection link on the page indicates if a tip for dissecting Atlas pages: the illustrated anatomical region is available on www.e-sobotta. • The menu bar on top indicates the topics of each chapter, the com. bold print shows the subject of the respective pages. • Important anatomical structures in the figures are highlighted in Appendix: bold print. • List of abbreviations, general terms of direction and position • Small supplement sketches located next to complex views can be found at the end of the book. show visual angles and intersecting planes and, thus, facilitate orientation. • Detailed figure captions explain the relationships of anatomical structures. Perfect Orientation - the New Navigation System Thoracic Viscera The menu bar with the The subject of this page terms printed in bold indicates the subject of the current page. Sketches facilitate orientation in complex figures by showing visual angles and intersecting planes. Important anatomical structures are printed in bold. Figure captions explain anatomical connections The Clinical Remarks concerning the boxes describe medical illustrated structures. contexts to the anatomical structures illustrated on the page. Mostly, these clinical aspects are also of high For pages with this relevance for the exam. dissection link detailed dissection tips can be found on www.e-sobotta. com. The following contents can be found in the other two volumes: m 1 General Anatomy e t s Orientation on the Body -► Surface Anatomy -► Development -► Muskuloskeletal y S System -»Vessels and Nerves -► Imaging Techniques -► Integumentary System al t e el k s o 2 Trunk ol c Surface Anatomy -► Development -► Skeleton -► Imaging Muscles -♦ s u M Vessels and Nerves -► Topography, Back -> Female Breast Topography, Abdomen and Abdominal Wall d n a y m 3 Upper Extremity o at Surface Anatomy -► Development -► Skeleton -► Imaging -► Muscles n A Topography -> Sections al r e n e G 4 Lower Extremity 1 Surface Anatomy -► Skeleton -> Imaging -► Muscles -► Topography ol. Sections V 8 Head Overview -* Skeleton and Joints -*■ Muscles -► Topography -► Vessels and Nerves -► Nose Mouth and Oral Cavity Salivary Glands 9 Eye Development -► Skeleton -► Eyelids -► Lacrimal Apparatus -► Muscles of the Eye -► Topography -► Eyeball -► Visual Pathway y 10 Ear m o Overview -► Outer Ear -► Middle Ear -► Auditory Tube -► Inner Ear t a n Hearing and Equilibrium a o r u e N d 11 Neck n a k, Muscles -► Pharynx -► Larynx -► Thyroid Gland -► Topography c e N d, a e H 12 Brain and Spinal Cord 3 General -► Meninges and Blood Supply -» Brain -► Sections . Cranial Nerves Spinal Cord ol V Paulsen, Waschke Sobotta Atlas of Human Anatomy Latin Nomenclature Internal Organs Translated by T. Klonisch and S. Hombach-Klonisch Sobotta Atlas of Human Anatomy Internal Organs 15th edition Edited by F. Paulsen and J. Waschke Translated by T. Klonisch and S. Hombach-Klonisch, Winnipeg, Canada 363 Coloured Plates with 441 Figures URBAN & FISCHER München Editors Prof. Dr. Friedrich Paulsen Prof. Dr. Jens Waschke Dissecting Courses for Students More Clinical Relevance in Teaching In his teaching, Friedrich Paulsen puts great emphasis on the fact From March 2011 on, Professor Jens Waschke is Chairman of that students can actually dissect on cadavers of body donors. "The Department I at the Institute of Anatomy and Cell Biology at the hands-on experience in dissection is extremely important not only Ludwig-Maximilians-Universitat (LMU) Munich. " For me, teaching at the for the three-dimensional understanding of anatomy and as the basis department of vegetative anatomy, which is responsible for the for virtually every medical profession, but for many students also dissection courses of both Munich's large universities LMU andTU, clearly addresses the issue of death and dying for the first time. The emphasizes the importance of teaching anatomy with clear clinical members of the dissection team not only study anatomy but also relevance", says Jens Waschke. learn to deal with this special issue. At no other time medical "The clinical aspects in the Atlas introduce students to anatomy in the students will have such a close contact to their classmates and first semesters. At the same time, it indicates the importance of this teachers again." subject for future clinical practice, as understanding human anatomy "The dissection links in the atlas lead to online images that are means more than just memorization of structures." relevant for the dissection. You can print them and take them along. The offered dissection tips are not instructions, but make sure that Professor Jens Waschke (born in 1974) habilitated in 2007 after graduation from Medical School and completing a doctoral thesis at you are oriented exceptionally well and not 'cutting in the dark'." the University of Wuerzburg. From 2003 to 2004 he joined Professor Fitz-Roy Curry at the University of California in Davis for a nine months Professor Friedrich Paulsen (born 1965 in Kiel) passed the 'Abitur' in research visit. Starting in June 2008, he became the Chairman at the Brunswick and trained successfully as a nurse. After studying human Institute of Anatomy and Cell Biology III at the University of medicine in Kiel, he became scientific associate at the Institute of Wuerzburg, in 2005, together with his colleagues, Professor Waschke Anatomy, Department of Oral and Maxillofacial Surgery and the was awarded the Albert Koelliker Teaching Award of the Faculty of Department of Otolaryngology, Flead and Neck Surgery of the Medicine in Wuerzburg. In 2006, he was awarded the Wolfgang Christian-Albrechts-Universität Kiel. In 2002, together with his Bargmann Prize of the Anatomical Society. colleagues, he was awarded the Teaching Award for outstanding His main research area concerns cellular mechanisms that control the teaching in the field of anatomy at the Medical Faculty of the adhesion between cells and the cellular junctions establishing the University of Kiel. On several occasions he gained work experience outer and inner barriers of the human body. The attention is focused abroad in the academic section of the Department of Ophthalmology, on the regulations of the endothelial barrier in inflammation and the University of Bristol, UK, where he did research for several months. mechanisms, which lead to the formation of fatal dermal blisters in pemphigus, an autoimmune disease. The goal is to gain a better From 2004 to 2010 as a University Professor, he was head of the understanding of cell adhesion as a basis for the development of new Macroscopic Anatomy and Prosector Section at the Department of therapeutic strategies. Anatomy and Cell Biology of the Martin-Luther-Universität Halle- Wittenberg. Starting in April 2010, Professor Paulsen became the Chairman at the Institute of Anatomy II of the Friedrich-Alexander- Universität Erlangen. Since 2006, Professor Paulsen is a board member of the Anatomical Society and 2009 he was elected the general secretary of the International Federation of Associations of Anatomy (IFAA). His main research area concerns the innate immune system. Topics of special interest are antimicrobial peptides, trefoil factor peptides, surfactant proteins, mucins, corneal wound healing, as well as stem cells of the lacrimal gland and diseases such as eye infections, dry eye, or osteoarthritis. All business correspondence should be made with: Elsevier GmbH, Urban & Fischer Verlag, Hackerbrucke 6, 80335 Munich, Germany, mail to: [email protected] Addresses of the editors: This atlas was founded by Johannes Sobotta t, former Professor of Professor Dr. med. Friedrich Paulsen Anatomy and Director of the Anatomical Institute of the University in Institut für Anatomie II (Vorstand) Bonn, Germany. Universität Erlangen-Nürnberg German editions: Universitätsstraße 19 91054 Erlangen 1st edition: 1904-1907 J. F. Lehmanns Verlag, Munich Germany 2nd—11th edition: 1913-1944 J. F. Lehmanns Verlag, Munich 12th edition: 1948 and following editions Urban & Schwarzenberg, Munich Professor Dr. med. Jens Waschke Institut für Anatomie 13th edition: 1953, editor H. Becher 14th edition: 1956, editor H. Becher Ludwig-Maximilians-Universität 15th edition: 1957, editor H. Becher Pettenkoferstraße 11 16th edition: 1967, editor H. Becher 80333 München Germany 17th edition: 1972, editors H. Fernerand J. Staubesand 18th edition: 1982, editors H. Fernerand J. Staubesand Addresses of the translators: 19th edition: 1988, editor J. Staubesand Professor Dr. med. Sabine Flombach-Klonisch 20th edition: 1993, editors R. Putz and R. Pabst Urban & Schwarzenberg, Munich Professor Dr. med. Thomas Klonisch Faculty of Medicine 21st edition: 2000, editors R. Putz and R. Pabst Department of Human Anatomy and Cell Science Urban & Fischer, Munich 22nd edition: 2006, editors R. Putz and R. Pabst University of Manitoba Urban & Fischer, Munich 745 Bannatyne Avenue 23rd edition: 2010, editors F. Paulsen and J. Waschke Winnipeg Manitoba R3E 0J9 Elsevier, Munich Canada Foreign editions: Bibliographic information published by the Arabic edition Deutsche Nationalbibliothek Modern Technical Center, Damaskus The Deutsche Nationalbibliothek lists this publication in the Deutsche Chinese edition (complex characters) Nationalbibliografie; detailed bibliographic data are available in the Ho-Chi Book Publishing Co, Taiwan Internet at http://www.d-nb.de. Chinese edition (simplified Chinese edition) Elsevier, Health Sciences Asia, Singapore All rights reserved Croatian edition 15th Edition 2011 © Elsevier GmbH, Munich Naklada Slap, Jastrebarsko Czech edition Urban & Fischer Verlag is an imprint of Elsevier GmbH. Grada Publishing, Prague Dutch edition 11 12 13 14 15 5 4 3 2 1 Bohn Stafleu van Loghum, Houten English edition (with nomenclature in English) For copyright concerning the pictorial material see picture credits. Elsevier Inc., Philadelphia English edition (with nomenclature in Latin) Elsevier GmbH, Urban & Fischer All rights, including translation, are reserved. No part of this French edition publication may be reproduced, stored in a retrieval system, or transmitted in any other form or by any means, electronic, Tec & Doc Lavoisier, Paris Greek edition (with nomenclature in Greek) mechanical, photocopying, recording, or otherwise without the prior written permission of the publisher. Maria G. Parissianos, Athen Greek edition (with nomenclature in Latin) Maria G. Parissianos, Athen Acquisition editor: Alexandra Frntic, Munich Hungarian edition Development editor: Dr. Andrea Beilmann, Munich Medicina Publishing, Budapest Editing: Ulrike Kriegel, buchundmehr, Munich Indonesian edition Production manager: Sibylle Hartl, Munich; Renate Hausdorf, Penerbit Buku Kedokteran EGC, Jakarta buchundmehr, Gräfelfing Italian edition Composed by: Mitterweger & Partner, Plankstadt Elsevier Masson STL, Milan Printed and bound by: Firmengruppe appl, Wemding Japanese edition Illustrators: Dr. Katja Dalkowski, Buckenhof; Sonja Klebe, Aying- Igaku Shoin Ltd., Tokyo Großhelfendorf; Jörg Mair, Munich; Stephan Winkler, Munich Korean edition Cover illustration: Nicola Neubauer, Puchheim Cover design: SpieszDesign, Neu-Ulm Elsevier Korea LLC Polish edition Printed on 115g Quadro Silk Elsevier Urban & Partner, Wroclaw ISBN 978-0-7234-3732-1 Portuguese edition (with nomenclature in English) Editora Guanabara Koogan, Rio de Janeiro Portuguese edition (with nomenclature in Latin) Editora Guanabara Koogan, Rio de Janeiro Russian edition Reed Elsevier LLC, Moscow Spanish edition Editorial Medica Panamericana, Buenos Aires/Madrid Turkish edition Beta Basim Yayim Dagitim, Istanbul Ukrainian edition Elsevier Urban & Partner, Wroclaw Current information by www.elsevier.de and www.elsevier.com fable of contents V is c e ra o f th e T h o ra x Heart....................................................................................... 4 Lungs...................................................................................... 28 Oesophagus .......................................................................... 42 Thymus.................................................................................. 50 Topography............................................................................ 52 Sections ................................................................................. 62 V is c e ra o f th e A b d o m e n Development.......................................................................... 72 Stomach ................................................................................. 74 Intestines ............................................................................... 86 Liver and Gallbladder............................................................. 102 Pancreas................................................................................. 120 Spleen .................................................................................... 128 Topography ............................................................................ 130 Sections ................................................................................. 148 P e lv is a n d R e tro p e rito n e a l S p a c e Kidney and Adrenal Gland..................................................... 160 Efferent Urinary System ........................................................ 174 Genitalia ................................................................................. 182 Rectum and Anal Canal.......................................................... 220 Topography ............................................................................ 228 Sections ................................................................................. 236
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