Simognathinae (Acarina: Halacaridae) from the Great Barrier Reef and Coral Sea, descriptions of thirteen new species

30 Pages·2000·14.3 MB·English

SIMOGNATHINAE (ACARINA: HALACARIDAE)FROMTHE GREATBARRIER
REEF AND CORAL SEA. DESCRIPTIONOF THIRTEENNEW SPECIES
JURGENC.OTTO
Otto,J.C. 20()() 06 30: Siinognatlimae(Acarina: Halacaridae)fromHieGreatBarrierReef
and Coral Sea, descriptionoftliirteen new species. Memoirs ofthe OneenslandMuseum
45(2): 505-534. Brisbane. ISSN 0079-8835.
The simognathine fauna ofthe Great Barrier Reefand reefs ofthe Coral Sea has been
mvestigatedandonenewspeciesoLicammaritisandtwelvenewspeciesofSimogriathus
were found: Acaromaniisdiazpulidoi Siniognathusabnomalus.,S. actius._S. aspidionis.
^
S, clypealits,S. conieatus,S. exoticus,S.platyaspis,S.pygmaeus,S. specicdis.,S. trachys^
S. versicolorandS. xandanis. A ke>' to AustralianspeciesofSimognatliinaeisprovided,
Acaromantis is redefmed, and the deilnition of Simognathus is coniinenled on.
Simognathiis, Acawmantis, SimognaThinae. Givar Batrier Reef, Coral Sea, Australia,
Halacaridae, mahnemites, Acati.
JiirgenC.Otto, (emaiL'j\otto{d:airfis,govMu),Australiaf7lmtiti^^
Townsville4810,Australia; 10September1999.
&
Mites of the family Halacaridae are benlhic Otto. 2000a,b.c: Otto Bartsch, 2000: Bartsch,
inliabilantsofmarineandfreshwaterhabitatsand 2000).onespeciesoT^carornantisand 12species
are assumedto have diverged from semiaquatic ofSiniognathusWQTQ found. These representthe
prostigmatid ancestors more than 200 million first records of Simognatliinae from the eastern
yearsago (Bartsch, 1996). Ofthe 14 subfamilies halfofAustralia.
of Halacaridae wliich arc currently recognised
METHODS
(Abe, 1998: Otto. 1999c). the Simognatliinae is
among the most conspicuous. Species of tliis Sand, coral aibble and pieces of dead coral
entirelymarinesubfamilyarespindle-shapedand
^^ere collected either inteilidally or subtidally
heavilyannotnedbylargedorsalplateswliicharc using SCUBAequipment. Mites were extracted
covered witlibroad shallow pits (fo\'eae). Their by wasliingthe substratesinabowlofwaterand
gnatliosoma is relatively short and the palps decanting the supernatant through a IOOijliu
wliichareinscrtedclosetoeachotherdorsallyon sieve. All material was collected by the author
the gnathosomal base ha\'e no more than three except where stated otlier\vise. Mites were
segments. The most remarkable feature is their cleared in lactic acid and mounted in PVA
frontlegswliichpossessa largeventralormedial (Boudreaux & Dossc, 1963). Drawings were
spine onthetibia anda hook-like unpairedclaw made withthe aid ofa camera lucida.
attheendofthetarsus.Theunpairedclawandthe
tarsusitselfaremovableandopposethespineon Inthe accountsofeach speciesonly one sex is
thetibia,anarrangementwliicliappearssuitedto described in detml, while for the opposite sex
catchingandholdingprey, although,feedinghas only characters that differ are described.
notbeenobsened. Measurementsare in micrometres.
Siniognathine mites are common in intcrtidal Tenninolog}^ follows Bartsch (1993a. 1994).
liabitatsandshallowwatersaroundtheglobebut To make a clear distinction between paranibu-
areyet to be recorded fromthe deep sea. The 34 lacral setae (pas) and \entral setae, only those
ventral, ventromedial or ventrolateral setae
describedspeciesaresubdi\idedintotwogenera.
Acarotnaritis Troucssart & Neumann, 1893 and whichare inserteddistalto the most distalofthe
dorsal setae fossar\ setae) are here referredto
SimognathiisTxoxKS^mt 1889. So far25 species as pas. In some cases the number of param-
tohfeSsehvforgonmathAuusstrlailaivea b(eBeanrtsdcehs,cri1b9e9d3,b,sev1e99n4)o.f bulacaral setae and \cntral setae may therefore
differ from those Bartsch (1994) described for
Acaromantisappearslessdi\'erse. Only ninespecies similar species. InAcaromantis and at least one
havebeendescribedandnone from Australia.
speciesofSimognathusthe tibiaandtarsusofleg
Duringthepresentstudy,whichfonnspartofa 1istwistedby 90°comparedwithotherlialacarid
broadersurveyoftlielialacaridfaunaoftheGreat genera. The tarsal claw^ is therefore pointing
Barrier Reef and Coral Sea (Otto. 1999a,b,c; inwards instead ofdownwards inuncompressed
506 MlilVlOIRS OF THE QUEENSLAND MUSEUM
FIG. 1.Acaromaniisdiazpulido!sp. nov '; A, idiosoma.dorsal view(AD,anteriordorsal plale;fo.fovcae; 2p,
gland pore_; OC. ocular plale; PD. posterior dorsal plate); B, idiosoma, ventral view (ads. adanal seta; AE,
anteriorepimeralplate:ev.epimeral vesicle:GA,geniloanalplate;GO.geniia! opening;os,ouiKingsetae;PE,
posterior epimeral plate; pgs, perigenital setae; sgs, subgenital setae); gnathosoma, dorsal view (gb,
gnathosomal base;p,palp;te,tectum);D,jnathosoma,ventralview(rO, rostrutn). Scalebars:A,B=100am;
C,D=50am,
NEW SPECIES OF SIMOGNATHINAE 507
specimens. To be consistentwithdescriptions of DESCRIPTION. Male. Idiosoma 354 long. AD
othergenerathe termsmedial, lateral, dorsal and longer than wide; foveate throughout (Fig. 1A);
ventral fortibiaandtarsus I areappliedas ifthese two pairsofsetae and pairofbarelyvisiblegland
segmentswereorientedintheusual waywiththe pores as illustrated; scattered canaliculi around
tarsal claw pointing downwards. Accounts of pores. OC reduced to more or less oval
chaetotaxy are from trochanter to tibia only. subcuticular platelet carrying few canaliculi.
Abbreviations: Descriptive- AD, anteriordorsal Anterior to OC with seta on a small subcuticular
plate; AE, anterior epimeral plate; GA, platelet. PD clearly longer than wide; anterior
GA
genitoanal plate; GO, genital opening; OC, margin rounded; with two pairs of setae.
ocular plate; PD, posterior dorsal plate; PE, foveate except for an anterior transverse smooth
posterior epimeral plate; P-2, P-3, second and area (Fig. IB); tGwOo pairs of branched outlying
third palp segments, respectively, counted from setae anterior to and one pair ofunbranched
base of palp; 1-IV, leg to leg IV; pas, setae level with anterior margin of GO; 14
parambulacral seta{e); pgs, Iperigenital seta(e); branched pgs surrounding GO; three pairs of
sgs, subgenital seta(e). Additional abbreviations short sgs. Adanal setae ventrally on anal cone.
in the illustrations are explained in the captions. Tectum with an upper split part and a lower
GBR. Great Barrier ReefMarine Park; I, Island; pointedpart (Fig. IC). Ventral gnathosomal base
Rf, Reef Otherabbreviations: ANIC, Australian with foveae posterolaterally and posteriorly,
National InsectCollection,Canberra(Australia), medially smooth (Fig. ID). Palp two-segmented
MTQ, Queensland Museum branch of the (Fig. 2A); P-2 with heavy ventral seta, one
Museum of Tropical Queensland, Townsville slender dorsal seta and two apical setae.
(Australia); ZMH, Zoologisches Museum
Outline of leas as shown in Fig. 2B-E.
Hamburg (Germany). All specimens with the
accession nuniber prefix QMS are deposited in Chaetotaxy: I 1-2-1-4-5 (Fig. 2B), II 1-2-2-4-5
the MTQ. (Fig. 2C), III 1-1-2-3-5 (Fig. 2D), IV 1-1-2-3-5
(Fig. 2E). Genu I with a strongly spinoseventral
SYSTEMATICS seta. One small branched seta dorsaily on tibiae
IIIand IV,twosuchsetaeongenuaIII andIV,one
AcaromantisTrouessart & Neumann on telofemur III and two on telofemur IV (oil
immersion). Tibial spine and tarsal claw ofleg I
Acaromantis Trouessart & Neumann 1893: 207; Andre, prior to compression ofslide medially directed.
1946: 138;Viets, 1956:696;Morselli. 1970: 109;Bartsch, Tibiae II-IV with pair of ventral coarsely
1974: 280; 1976: 664; 1977: 92(530); 1980: 401; 1983:
194;Bartsch& Schmidt, 1978: 22(644). bipectinate setae. Paired fossary setae of tarsi
II-IV slightly serrated. Tarsi ll-IV without
TYPE SPECIES. Acaromantis squilla Trouessart & median claw. Tarsus III withventral setaand pair
Neumann, 1893, by nionotypy andoriginaldesignation. ofpas. Tarsus IV with two ventral setae and pair
DIAGNOSIS. Body spindle-shaped. Dorsal ofpas. Pairedclaws oftarsi II-IVwith accessory
process andpecten.
platescoveredby foveae. Palpsseparatedby less
thantheirwidth,inserteddorsailyongnathosoma Female. Unknown.
and two-segmented. Tibia offront leg at least as
longasthreetimes itsheight, apically narrowing REMARKS.Acaromantis diazpulidoisp. nov. is
and equippedwith ahea\'y spine. Tarsus ofsame the first species of Acaromantis known from
leg shorter than 1/3 the length of tibia, with a Australia. With itstwopalpsegments, singleseta
heavy apical claw but no paired claws. Genu of ontelofemurI,presenceofaspinosesetaongenu
legIwithaspinoseseta.TelofemurIwithasingle I, short tarsus I, slender and in the proximal half
seta. conspicuously widened tibia I and lack ofpaired
Acaromantis diazpulidoi sp. nov. rcelparwesseontnattiavresuos{AIc,aAr.omdainatzipsu.liIdtodiiffiesrsaftr^o^mpicaalll
(Figs 1,2) itscongeners by the shapeofitstectum, which is
characteristically split in contrast to being entire
ETYMOLOGY.ForGuillermoDiaz-Pulidowhocollected andscale-like. Malescanalsobeidentifiedbythe
theholotype.
presenceofthreepairsofoutlyingsetae,ofwhich
MATERIAL. HOLOTYPE: QMS105316, 6, Coial Sea thetwo anteriomiostpairs arebranched. Such an
(QueenslandPlateau),ChilcoltL, 16^^56.5rS 150°0.4'E, 14 arrangement ol"setae is unknown for any other
Sep. 1998,G.A.Diaz-Pulidocoll.,coarsesandat10-15m. species ofAcaromantis.
508 MEMOIRS OF THE QUEENSLAND MUSEUM
B
mcl sp
F]G.2.Acaromanfisdiazpu/idoisp.nov., 6:A.palp,medial vicvvfP-l. P-2.palpsegment); B. leg1,medialview
(ba. basifenuir: ge, genu; mcl, median claw; pas, paranibulacral setae: sp. spine: ta, tarsus; le. telofemur; ti,
tmi,biat)n;eCd,iallevgiieI,wm)e;dEi,allevgiIeVw,(fmse,dfioaslsavriyeswe.laSec;aplaes,baprasr:aAmbu-laIcQr^aml;seBt-aeE;=pelS,Ojpuauimr.edclaw;vs,ventralseta);D,leg
SimognathusTrouessan T^TE SPECIES. Pachi^-naihm scidplm Btad>; 1875 (=
Pachygnathns miyniitis Ilodge, 1863; synonymy by
Simognathus lToucssan, 1889: 1180; Viots. 1927: 151; 1936: Fountain(1953)); byoriginal designation.
421; 1940: 94; 1956: 694. Andre. 1V46: 135; Fountain,
1953:357; Monniot, 1961- 5S5: 1%2 288; Barlsch. 1974: DIAGNOSIS. Body spindle-shaped. Dorsal
275: 1977: 87 (525t: 1*^^: 19; 19^:3: 193: 1985: 554; platescovetedbyfoveae.Palpssepamedbyless
1993b: 95: 1994: 135: Ncwcll. 1947: 23: 1971: 36; 1984: thantheirwidth,inserteddorsallyongnathosoma
264.
Ischyrognathus Trouessart, 1901; Newell, 1947: 23, 37; and three-segmented. Tibia offront legs usually
Vicls,1956: SynonynQ'byBartsch. 1974. shorterthanthreetimesitsheight(exception:S,
NEW SPECIES OF SIMOGNATHINAE 509
FIG. 3. Siwoi^ncuhm ahnormalus sp. nov., adult: A, idiosoma, dorsalview;B. 6 idiosoma,ventralView; S
genital opening; D, gnathosoma, dorsal view: E, gnathosoma, ventrnl view;F, posteriorhalfof9 idiosoma,
ventral view. Scale bars: A,B = IOOh-hi; C = lS\Lva; D,E-50|xm; F= lOOM-m.
oiwjrwate, 5 s(7//£7m\), equippedwith ahea\7 usually longer tb^ 1/3 the length of tibia
spineandtarsus olsaine legswithaheavyapical (exceptiont 5. dbnorrndtlus, 3. sCUta$m)i vdHk
clawandslenderseta-ltkepairedclaws. Tarsus I heavymediancJawandslender, sometimesseta-
510 MEMOIRS OF THE QUEENSLAND MUSEUM
FIG.4.Simognathusahnomalussp.nov.,adult:A,legI,dorsalview(racl,medianclaw;pel,pairedclaws;pas,
parambulacral setae); B, leg11,ventralview(fs,fossaryseta); C, legIII,ventralview;D,legIV,lateralview.
Scale bars: A-D -50^xm.
like paired claws. AU setae of genu of leg I Simognathus abnormalus sp.nov.
glabrous. Telo^mur I usually With two s^tae (Figs 3, 4)
(exception:S.pygmaeussp.nov).
HTt'MOl-OGY- F^eferring to the front legs" short tai^iis
andslendertibia, andthepeculiarorientation oftliese leg
segments.
1 ,
New SPECIES OF SIMOGNATHINAE 511
MATBRjAt. HOLoi vrc: (;msuwi7, 5. cjbR, species possess fused venin*! plates,oblOn^OC
Ig'^ISUS147^06.49^£^ Lpodsttmc itf, J2 Aiu. and arelatively slendertibia and short tarsus on
^fi^1:feat2nuPARArYl^:{3MSl{S3I^ 9,dm legl,acombinationofcharacters ihalisunknown
for othei species in the ^cnus. Siffmy^f/nlhiti
jPI1Cr84tafa9l*r°st*De6WZ.sae8at95n3«d*mRa:1tE4Ol17i2MzmS*a;Ib2e0Z3t5hM23R2rHf0.E,A2,352M?/.D^9e9*Oc,MSS5w]I,9W09G.57B,3K21c,oI4a,rAispgJer',.s^aOG1n.B9dH9R'8&.S. sabobcdnuyow«rumasanldbiyiashwaicvdaienngnubwnei-dttdeiirvsedtaoitrnesgaaulnisns)lhaeittdeisc>diiaarloll)ma'rcoSen.t
ROPoUMtscsSaeur1j0R.5f:c^...c2ha:i.lc61ol>t.QIi.).7M'1SS611^40555'63'i233r3,'SE.I85C00o°ar0.a.4l']SR9e.0aS1,(4QsSuaeenped.nast1l92a9mnR;.d theventraSliginnaotehioMHifolniiiailSh4ka(sica.»sp.nbv.
Ci.,\. Dia/-Pulido. course saiid at :0-I5m; QMSIO?>24
WIiOl'l;i".?s->,1..**c;i'.fs,IC!/oImS!'SSeIa4'>('Q3u8eTe,nsl1a5ndS^Ppl.atcaM). SCo}u^t.h FTYMOLOGY \ .urn. uao = {XtOfiTvitififtring tp !li&
Diaz-Pulido. coroi rubble (tiiifi in t")-10ni. spcviet;'occurrence on beaches.
nrStKII^ ION, Xfule. Idiosoma 412-424 long M.MIRIVL. MOtOTYPF: f^)\tSI0s32fv CBR.
(holoiyp: I424). AD or subequal length and W i<..!)'-)"S I47-G3.05*F. Cape Ferguson-.. S Nov. 1097,
j(swplFilijdaggtlh.chll;l3c}tA/v>».uilhtohiOncgalCeirnroctntaf-hnfdaibniiv^^euaiIalddeensa:eirtbeaab^nnatagnentrftaijieoirurfpbttedtouyn^l.a^iracPgl^ilaDnyT GsPiVnaBAt,ncRRQrdt,AMidTb;SWeY:oilP1lno0Fwg5sS3aa2:hn8QBid.egMah6Sdn1i,ntd,0iad5tbeIa3e6aa2m*sca"2hfrF,okrS,ihs,oii.s"F-elAd.5diMuinC-nip.'iccVenfnt4(i.,AdZdwcteMSppd1jHt0iuh5K^244^0U>le'c,).'nm.,'_i-.s;,
toildmlUtsD>tnciamltccscdsl;pacew)ci.itAmheLtnlsbureAoeaEdplaayinrdfsiGioseAfdsateolisCaoic.f\u((siIendicgl.tuo3dBiP)nEg.' dCQeoMpvteSh,1t10O45-2O30c3ct0t.-tr11.90953S3,4c.oa[5^sVasr,idGaBiURi,ghtLii/daerldev1^.,seMde4ninatatindt
(as shown for 5 in Fig,W^.AEandPE foveaJe Fi'nuilc. Idiosoma 244-272 long fholoiyp^'^Ti-
mtoughoui(Fig. 3B).GAfoveateexceptftrt'Area All dorsal setae short (I'lg. 5A'). AD unifoTtno*
along median a\i^ (Fig. 3B): two pairs of pale, either ot similar width in anterior and
outlvinL! selae anterioi To GO; 28-30 pgs ppsAftnoi hall^or ppstedorJiolf^Lij^Uy y^^^
MrfcnuidingGOin a circle (Tig- ?B,C)lGOwi:Ui ]1Qstenoiinafginrnunoearlaterollyvnmstuttered
threepairs ol shortSig^(F^ 3Q, pits. (K* n_-diu;ed In mini.iU' plnlcwith a setae PD
Gnafhosomalbas&\«ridei%em^ai^(P%.3D,C1: pear-slia|xd, vMtii ihicc pairs ofsetae ineludmg
tejTtun wide, naiTowing anieriorly (Fig. 3D); ad<in;d se(ae, Al, wilh Iovlmc only lalerally and
ventrally Ibvealcexceptforarelatively widearea along posierujr margin Fig. 5B); remainder of
along the median axis (Fig. 3F). r*alj> three- plate punctate; epimenil( vesicle hirge. OA wilh
segmented;P-2 withhea\7ventralsctaj i'-3 with foveaeinposterior2/3andalonganteriormargin;
1}irce.shortsdaeandonelongerstitoi remainderofplate punctate; ^4 pairs ofpgs.
OulUne of legs as in Fig. 4. (?hactotn\\ I Dorsal gnatlu»sonialbase with large uTcgularly
1.2-1-4-5 (Fiii. 4A|, II 1-2-3-4-5 -lliK 111 shaped loveae(Fig.5D). Tectum poiiUcd.Ventral
1-1-2-3-5 (Fig^.4C)JV I- N2-_V5 {Fig.41 lihia gnathosomal base Ibvcatc except medially (Fig.
1 long and slender, narrovviiig apieailj. All setae 5£), P*2 with A ventr^^i^tti and a poorly
ofgenu I slenderandglabrous. Intmcomprcsscd developedwhnalnrohibcmnccproximaltoseta.
specimens tibial spine and tarsal elaw oflcg I P-T lunger than halfthe length of
mediall) directed. Tarsus I! with the medial OuiliiH: oflegs as in Fig. 6. Tclofemqra^With
lbssar> seta apically widened and serrulcd, Tarsi fw
III and IV with single ventral seta and one consfpAirceuooeu,sonthlaeng Jonmorotehneurmclreogus&^('Fiimg.d U6MAx)i.t
mUiiienad[Mniidi^vIdcVlpaaapwsi.;caDplialsiytraSelfdspigarIia^trverdo?.fblfeaoansrdsian1rg1y-,1psVceciwaticetnohfasltmaoarns1gi C1.1hai-e.lolaxv: I 1-2-2-4-5((FFiigi\: o6CA)).,I1V1 i1--2l--2F-;4--35
MOSrtoftheshall,most(?to>S{>feuo«»QntarsusIV. p(Friogx.i6mDa)l. Hsewaelvlyinvge}n.traSflttss*piinedjoifitciebmia tIowi.tshpoiunte
Femalii. Idiosoma 417-420 long. GA with four branched (Fig. t^A^am$wed}.l]ai:SusI]withpair
pairs ofpgs and no sgs (Fig- 3P). OC in some ofpas btttwlltout vitrttr«rs8lta,lafsl TO and IV
N|)Leimens shorter and more mondcd than with single ventral seta hut -vMihoui pas. Paired
illuslraled for male. clawsoftarsi II-1V wilhapical claslcrofpeeten.
RF-MaRKS. Stnu^^ttuihti.s t//>//r//v»iij/«i is. iriii.sc SUie, Idiosonta 241-250 long, GA with 16-17
snnitertn'3. scuiafus Bttrtsch, W^b. Both pe^(Ing, jrOiSg^itotseen.
512 MEMOIRS OF THE QUEENSLAND MUSEUM
FIG,5.Sirnos^nathus act'iussp. now. adult: A. idiosoma. dorsalview(OC,ocularplate; ads,adanal seta); B, 9
idlosoma,ventral view;C, 6genitoanalplate;D,gnathosoma^dorsalview;E,gnathosoma,ventralview.Scale
b?urs:A-E=50|xm.
REMAKKS,SintognatiiKs acfiu.s sp. nov. differs Slightly in the striation of the membranous
£romitsctinge&ersbyhavingabranchedsetaon cuticle.InthespecimensfromCapeFergusonthe
tibiaIadjac^ttotheventt^spiAeandanatrow
bandoffoveaealongtheposteriormai^ofthe striation is conspicuously liner (Fig. 5A) than in
AE. all remaining specimens. This character is here
Althoughallexaminedspecimensareidentical notregardedassufficientevidenceforseparating
in the characters described above they diflfer Aesespecimensintodifferentspeciesalthoughit
NEW SPECIES OF SIMOGNATHINAE 513
FIG. 6. Simognathus actius sp. nov., adult: A, leg I, lateral view; B, leg 11, ventromedial view; C, leg III,
ventromedial view; D, legIV, ventromedialview. Scale bars: A-D = 50|jLm.
may indicate a certain amount of genetic RosserRf,ca. 15°37'S 145°33'E,8Oct. 1998,sandat2m;
isolation between them. QMS105342, c?,GBR,BoulderRf,ca. 15.24'S 145.27'E,
QMS
8 Oct. 1998, coarse intertidal sand; 1053431, 9,
Simognathus aspidiotus sp. nov. GBR, BoulderRf,ca. 15°24'S 145°27'E, 8Oct. 1998,A.
(Figs 7, 8) Thompson,coarsesandat2m;QMS105344, 9,CoralSea
(QueenslandPlateau),LihouRf,ca. 17°25'S 151°40'E,22
ETYMOLOGY. Greek, aspidiotes = shield bearer, July 1998,D.Fenner, sandat7m;QMS105345, 9, Coral
referringtothespecies'Hisedventral plates. Sea (Queensland Plateau), Flinders Rf, ca. 17°35'S
148°27'E, July 1998, D. Fenner, sand;
MATERIAL. HOLOTYPE: QMS105335, 9, GBR, QMS105346/S105347,2 6s, QMS105348, 9,Coral Sea
Myrmidon Rf, 18°16.69'S 147°23.2rE, 14 Apr. 1998, (Queensland Plateau), Herald Cays, 16°57.17rS
c9o,arAseNsIaCn,d&9,rubZbMleHat1A53m4./9P9A3R,AT9Y,PGEBSR:,QM19S°21005.31326'&,S s1a4n9'd^1at2.50-316'5Em,; 1Q6MSSep1.053149998,,9G,AC.oraDliaSz-ePaul(iQduoe,encsolaarnsde
149°02.85'E, Elizabeth Rf, 25 Dec. 1997, coarse sand Plateau), South Willis I., ca. 16°18'S 149°58'E, 15 Sep.
rubble at 3m; QMS1053371, 9, GBR, 18°42.03'S 1998, GA. Diaz-Pulido, coral rubble (fme) at 0-lOm;
r1u4b7b°l0e6.a5t4'E1,2-L1o5amd;stQoMneS1R0f,531328A/p1r0.51393998,,c2oar9sse,saGnBdR&, QI.,M1S6°15065.35510',S 915,0C°o0r.a4lE'S,ea14(QSueepe.n1s9l9a8n,dGPAla.teDaiu)a,z-CPhuillicdoot,t
18°38.25'S 147m42'E, John BrewerRf, 11 Apr. 1998, coarsesandat 10-15m.
coarse sand at 15m; QMS105340/S105341, 2 9s, GBR,
514 MEMOIRS OF THE QUEENSLAND MUSEUM
FIG. 7. Simognathus aspidiotus sp. nov., adult: A, idiosoma, dorsal view; B, 9 idiosoma, ventral view; C,
posterior part of 6 genitoanal plate; D, gnathosoma, dorsal view; E, gnathosoma, ventral view. Scale bars:
A,B,D,E= 100|xm; C =25|jLm.