Revision of the Genus Myosides Roelofs (Coleoptera:Curculionidae)

29 Pages·1993·4.1 MB·English

by  Morimoto, Katsura

九州大学学術情報リポジトリ
Kyushu University Institutional Repository
Revision of the Genus Myosides Roelofs
(Coleoptera:Curculionidae)
Morimoto, Katsura
Entomological Laboratory, Faculty of Agriculture, Kyushu University
Lee, Chang Eon
Entomological Laboratory, Faculty of Agriculture, Kyushu University
http://hdl.handle.net/2324/2564
出版情報：ESAKIA. 33, pp.59-86, 1993-03-31. 九州大学農学部昆虫学教室
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権利関係：
,AUIASE (33): 59-86. March 31, 1993 59
Revision of the Genus Myosides Roelofs
arustaK M ORIMOTO
l,aycriog toyaltrolomubocataLnFE ,feorutluuhcsiurygKA
,y t,iaskroeuvkiun FU218napaJ
and
noE gnahC L EE
Deparhnent fo ,ygol oeigBelloC fo laru t,aeNcnkeoiocpSgnuyK
la n,oyittiasNre v,iungUeaT 107-207aeroK
Abstract. ehT suohtangoleda sliveew fo eht suneg Myosides ni eht ebirtbus
anihcotP srucco ni eht raF tsaE gnitneserper 21 .seiceps eniN wen seicepsera
:debircsed
formosanus, yonagunianus, ishigakianus, okinawanus, amamianus,
dna ylwen si relleH
tokarensis, ejimai, kiiensis chejuensis. Trachyodes marshalli
derrefsnart ot siht.suneg
ehT suneg saw dehsilbatse yb sfoleoR )3781( htiw a elgnis seiceps
Myosides seriehispidus
tnes e.rnPa pm aoh Jrt)fo6b9 8p 1r d(aye hbdssSdauawryp dinnaieolhpyhc ae rb sTienfrhiottleoR
noitagitsevni slaever eht ecneserp fo evlewt seiceps ni siht suneg morf nretsae aisAgnidulcni
relleH . sihT suneg sawderrefsnart
Trachyodes marshalli
morf i neihetolhpyh coatrT ienhithcnyhro iytbO o&K )0391( dna ot eht inihcotP usnes rettieR
)3191( ( = inirecihpyC ees
morf eht
sehsub dna eht yssarg secalp yb gnitaeb dna .gnipeews yehT era yllausu derevoc yb a ytrid-atsurcni
noit dna nac ton eb deifitnedi tuohtiw gnitaert ni a tnegreted noitulos ni na cinosartlu renaelc niynam
.sesac
laireta Mdenimaxe ni siht repap era yltsom detcelloc yb .rD .S arumoN dna yltrap yb .srsseM.K
1) -goegoiB dna noitulovE ehT“ no margorP ecneicS evitarepo-oC napaJ-aeroK eht morf stluseR
yhpar fo eht stcesnI ni eht tsaE .”aisA .oN.41
2) noitubirtnoC morf eht lacigolomotnE ,yrotarobaL ytlucaF fo ,erutlucirgA uhsuyK,ytisrevinU
akoukuF .reS( ,4 .oN.)84
60 K MORIMOTO & C. E. LEE
,abaB .H ,amijoK .H ,arahikaM .Y ,ekatayiM .S ,awakayiM ,arihoN:A .K ,atagO .H ,ijasaS.K
adawaS dna .S .onCU oT‘ eseht stsigolomotne eht roines rohtua sehsiw ot sserpxe sih erecnis.sknaht
lnaaier resotaKaw melba ottce l hletoci cnwdant i snksiiusos iasryaavw yb .rM e agJnoey,MoJ
rotceriD ,lareneg dna .rD gnoJ kuK ,miK fo eht ts eh,rceortFauetsiet Rs.nrIM gnueS 1uK ,kraP
lanoi tnaoNew n efageoKnL uno aY.hfC o dr,nPnaoit attnSemire ptxsEe ruobF-gn urfhooCtceriD
,ytisrevinU ot mohw ew tnaw ot sserpxe ruo yltraeh.sknaht
suneG Myosides sfoleoR
Myosides ,sfoleoR ,3781 .nnA .toS ent. ,.gleB 461:IVX .sedieolhpyhca r:Ts(eiceps-epyT Myosides
seriehispidus ,sfoleoR .)cisabonom - ,prahS ,6891 .snarT .tne .toS ,.dnoL .39:6981 - ,onoK
1930, J. .taF .ordgiAa k.kp om,HI.vinU ;inihcnyhrroit0(122:)5(VIXX .)y ek - ,otomiroM
1962, J. .taF ,u.hvsiuny UK,.rgA ;anihcotP:inihcotP(25:21 .)y ek - ,otomiro M,1891
J. Fat.
,.rgA uhsuyK ,.vinU .281:52
mreD h t.,is e weetgrs anun dsciteyaellcdleluaulHaseraci ctnstdn silnomduacuumrttisgonrol
eht ta tsehgih ,xevnoc ylneve seyE .murtsor fo esab eht naht reworran elttil a seye neewteb daeherof
.elddim murtsoR retrohs naht ,daorb suounitnoc ot snorf ni eht emas enalp no ,musroddedis-lellarap
y ldrleoad rn, eaattpiaax gle nydyerelhewdttoip rprraoarfn a etcrnoahts soitd sedis; efmootsipe
emotsipe ,erab ylprahs detimiled ylroiretsop yb -V ro depahs-U anirac dna htiw a erabecaps
dniheb , trioiret slolpaw fo eht anirac htiw 8-6 sriap fo tnebmuced ro devruc gnol ;eataeisgyretp
;dekram-llew 1annetna seborts ,lasrod ne p.oylro ieeatnsnoept n,Atsubor epacs gnihcaerrorietna
nigram f,,omd ueettloncnei omntrguspe f1st-n 7e,mtgseus btourob aeciwt sa gnol sa ,daorbdn2
tnemges a elttil retrohs naht ,tsl dr3 ot ht6 stnemges tuoba sa gnol sa ,daorb ,suorefilinomht7
tnemges ,esrevsnart bulc .lavo selbidnaM htiw lacipa htoot ,tnulb lartnev gnittuc egdir llew,dekram
h t, hihnh cwtataaoiieo dw t3egm ntdounloaba 5 tr o,hesate se hrtetr oshesno l/2 - 2/3 h teghntel
fo eht regnol .seno mutnemerP htiw 3 ro 4 sriap fo ,eates eht tsom laretal riap netfo.tcnitsidni
tsonnig rraomire tet,nhueatbs nl ea ssune sibruoall utcuooh t,ig wnn oarlhetda oxrabrohtorP
-etsop ,noitanimaxe esolc yb sedis ta sebol raluco mrof ot decudorp yltniaf yrev dna thgiarts yltcaxe
t, senrot io mamgi slcu rrdaantfaauoomorn rto sbrr ap ey, ndl nraatrio hohogretrgrihbiaetlmtsna
tuo bmaumetso rfhpotgn enlaidem 2/3 ylla uwsour reusfrevsn alratcipa b,umsuton ohfrtopg n eehlt
etunim mulletucS .worruf eht morf detarapes lla ta ton ro ylworran eaxocorp ,edis lartnev ta tcnitsid
ro ,laig idtesleavecnoc yb selacs fi.tneserp
artylE ,lavo ylneve dednuor ta ,sedis tsedaorb ta ro tuoba 3/l morf ,xepa iremuh ,etelosboeairts
reworran naht ,slavretni ht9 airts .etelpmoc muidigyP yletelpmoc delaecnoc yb.artyle
.dedis-l e,sllwsaloem arce.rrotsaarsupnpoo suy cgls MieeltdianatxooiocrvcbaocrP -retsateM
mun ,tro hescnat sniedewteb -osem dna eaxocatem sa gnol sa dr3 .e tliarntrnectvsipeetreuMtus
elbisiv ni eritne htgnel ni delacs ro deduned .snoitidnoc retneV htiw tsrif etirtnev etaunis taroiretsop
t,sn o idms,gnlagrianahomelb eaxoc dn o cseas etirtnev ne ksaettir t g dnhddsnsnetnraoaav4a3l
.rehtegot
sgeL htiw aromef ,etavalc hcae htiw a llams prahs ,htoot eaibit tsomla ,thgiarts ,etanorcum israt
,tsubor tsrif tnemges sa gnol sa owt sgniwollof ,denibmoc dnoces tnemges ,esrevsnart drihttnemges
REVISION OF
MYOS~DES 61
,debolib hcum redaorb naht ,dnoces swalc ,elpmis.eerf
:elaM htneveS etigret etacnurt ta ,xepa ht8 etigr e,tetanigrame ht8 etinrets htiw ehtnaidem
ed mle,uazslr itut sgtrcsfunoaiaoo irppg nttes earalht rarctin bpas9ne,mprttdeeefuemstoovstrsuc
yllasrod ta eht thgir edis fo .sinep nemgeT yletelpmoc ,degnir seremarap ,laigitsevmuirbunam
neli tageisru naad,n ibm doehf yettte,lvfidrkr ow n aa ur ,isle,censiuwrgainbenhneudoteptnlpels
no ils , tat teyaru, adr ztrgrieh intne svcntesogsg uaovlninkmh roniatdelehenlpcwacaser -nevorp
sulucirt seni pflsol uhft irewotarep s,ayldepahs -dZedl onfet f,ogn oc lalsanret n,ixarohtos enmi
in
basal area.
:elameF htneveS etigret etacnurt ta ,xepa ht8 etigret ,etanigrame ht8 etinrets htiw gnolmulucips
.elartnev rotisopivO ,gnol htiw lasab suonarbmem trap hcum regnol naht eht lacipa dezitorelcs,trap
tuohtiw ,ilyts aera fo anigav ,gnol delioc eno dna flah semit ni ,esoper asr uxbirtalup olclamsot
laretal ,esab ta devruc ebol lasab htiw acehtamreps ,rotisopivo elohw naht regnol hcum netfo ,egral
ebol ,trohs latsid ebol devruc ralugna-thgir ot lasab.ebol
oyteK
species
l(14) emotsipE htiw sti dnih elgna dednuor dna gniyl erofeb ro no eht level neewtebroiretna
nigram fo ,aigyretp htiw redaorb erab ecaps dniheb.emotsipe
Z(7) eannetnA htiw dr3 dna ht4 stnemges fo elcinuf a elttil regnol naht.daorb
3(4) htiw artylE ,dn2 tsedaorb artylE .esab ta thgieh dna htdiw emas eht fo slavretni ht4 dna dr3
ta eht ,elddim neht yldaorb dednuor ylroiretsop ni lasrod .tcepsa.nawiaT
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .ps .von
Myosides formosanus
4(3) artylE htiw dr3 lavretni hcum redaorb naht gnirobhgien slavretni ta.esab
5(6) -roiretsop deworran ylthgiarts tsomla ,reworran elttil a artyle ,htgnel ni mm 4.3-1.3 ,rellamS
yl nroh lacipa l/3 ,sec itpnaioj ndon c uootr tcerebus deta lyildthg i,lrsewor rsaenl acs -lacipa
.yl .nawiaT . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. (Heller)
Myosides marshalli
6(j) ,regra L0.4 mm ni ,ht ganretly l,el ayvlo ddaeodrnbuor yllacipa ni lasrod ,tcepsa tcerebus
selacs worran ,ralugnairt tuoba eciwt eht htgnel fo lacipa .htdaerb suykuyR inuganoY(.).I
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .ps.von
Myosides yonagunianus
7(2) eannetnA htiw dr3 dna ht4 stnemges fo elcinuf sa gnol sa ro a elttil redaorb naht.gnol
8(9) eht naht regnol elttil a dna daorb sa gnol sa elcinuf fo tnemges d3 htiw eannetnA 4th, artyle
.lavo suyku yiRkagihsI( & etomoirI.).sI . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosides ishigakianus .ps.von
9(8) eannetnA htiw dr3 dna ht4 stnemges fo elcinuf fo eht emas epahs dna.ezis
)11(Ol awanik Os(uyku y.Reldd ie mhtttasedao r,bla vaorty lE & Kume Is.).
sp.
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides okinawanus nav.
)01(ll artylE ,demrofirypbus .eld dedihnmti htesbeda orb
12(13) ,r e9g.r3a-L7.3 mm ni ,ht ganretl yylleneve dn a y.ly dldareo odriinbmruaeomtrAsop 1~1s.
imamA( amihsO & amihsonukoT.).sI . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .ps.von
Myosides amamianus
13(12) tseda oemr hobytrlfroiret sdoepre pyaltthgia rrtesh taarrt y,lhet g6 mn.nme3il- 0,.r2ell amS
tniop ot dnuor tniojnoc .secipa arakoT 1~1s. arakaT( & amihsonakaN.).sI
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .ps.von
Myosides tokarensis
62 KMORIMOTO&C.E.LEE
)I(41 nig rda nm sihth tieimwotsipE ,etalugna level eht dniheb gniyl nfiog rraomirete tnneaewteb
,aigyretp htiw reworran erab ecaps dniheb.emotsipe
15(16) latoT ydob htgnel 6.4 ,mm htgnel fo mutonorp dna artyle 0.4 ,mm mutono ryplecracs-daorb
re naht ,gnol a r,tlyalveo-gnolbo 1.45 semit sa gnol sa ,daor bmurtsor htiw aigyretpllew
.yllaretal dednapxe otnug-ojnaD amihseM(.slsI .).I . . . . . . . . . . . . . . . . . . . . . . . . . Myosides ejimai .ps.von
)U(61 ,mm 3.3 naht ssel artyle dna mutonorp fo htgnel ,mm 0.4 naht ssel htgnel ydob latoT -onorp
.demrofiryp ro lavo artyle ,gnol naht rediw hcum nrut
17(18) l ahvcr aefaEtor nthiytliew a wor fo tnebmuced trohs ,selacs gnieb retrohs naht eht-moc
denib htgnel fo owt dnuorg ,selacs artyle ,demrofirypbus tsedaorb dniheb eht.elddim
uhsn o,HamayakaW( araN & akasO.).sferP . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosides kiiensis sp. nov.
)71(Sl f w ohoatr iawrty lfleoavret nhicaE tcerebus hdt eg r nnf,genieo sngableinhmleotoablccs
owt dnuorg.selacs
19(20) artylE ,demrofiryp murtsor htiw aigyretp llew dednapxe .yllaretal ,uhsnoH ,ukokihS.uhsuyK
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides pyrusprahS
20(19) artylE ,lavo murtsor htiw aigyretp ta tsom ylthgils dednapxe.yllaretal
21(22) tcerebuS se lr a eanc, dro srdnteeney rgallnonseamohlt 4sem ist agnyol le,scdaraaocrsb
denediw.yllatsid aeroK odujehC( .).I . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides chejuensis .ps nov.
22(21) tcerebuS ,yllats iddetal iydltcnits i,dretro hasrty lnseoela cs 2.5-3.0 . ds g a esnsomaoarilbt
. . . . . . . . . . ..tsa Era Fnaissu R,aero K,uhsuy K,ukokih S,uhsnoH Myosides seriehi.spidus sfoleoR
yeK to species by the male genitalia (ejimai sunasoznrofdna era ton)dedulcni
I(4) meullaurctis pafSgo hett 9itnsroemtlsa sa kciht msuairbu nf ao, mnreedmhegtteiatelnid
ron denettalf.yllatsid
2(3) sinePhtiw a dnuorlacipa .noitcejorp . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosidespyrus prahS
3(2) sineP ylneve . . . . . . . . . . . . . . . . . ..xep at adednuo ryldaor bdn aderepa t Myosides chejuensis .p.svon
4(l) ,nemget fo muirbunam naht rekciht hcum ,tsubor etinrets ht9 fo elartsag mulucipS ylcnitsid
detalid dna denettalf ta.xepa
j(6) sa semit eerht naht ssel turts fo esab lartnev dna xepa neewteb ecnatsid ,tsubor ,trohs sineP
gnol samumixam thickness in lateral aspect. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides marshalli) relleH(
6(5) sineP ,rednels ecnatsid neewteb xepa dna lartnev esab fo turts erom naht 3.3 semit sa taergsa
ssenkciht ni laretal.tcepsa
7(S) lanretnI cashtiw three sclerites. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides kiiensis .ps nov.
S(7) lanretnI cas tuohtiw setirelcs dniheb eht esab fo.sinep
)2I(9 sin e ehPnctenieaw wtxtseeipbda dlnaar tenseavb fo t9u.r 4ts-se3m.i3t sa gnol sa-kciht
ness.
)11(Ol sineP ,rerednels .dedis-lellarap . . . . Myosides seriehispidus sfoleoR morf( )uhsuyK
)01(ll .y l.l a ds.eaw bo .ryr l a.k,nar ee.wt rs.oihn se.P Myosides seriehispidus sf omloo)reafRe(roK
)9(2I ts as esea n2mar ea.ig trh5t e uotlsfrmaaotrbs t xdnneneepavea wetcenba t hsstiiidnweP
ssenkciht ni laretal.tcepsa
13(14) la nc rahesttinw I a sr ei.t am itpufraio et.lscos . . . . . . . . . . . . . . . . . . . . . . .. Myosides amamiurzus sp. nov.
14(13) lanretnI cas tuohtiw etinifed setirelcs ta.muitso
RJSISION OF MYOSIDES 63
15(16) sineP htiw lartnev ruotnoc ylkaew denrut-pu raen xepa ni laretal.tcepsa
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .ps.von
Myosides tokarensis
16(15) sineP htiw lartnev ruotnoc thgiarts raen xepa ni laretal.tcepsa
)M(71 -stk eac2ssmei.aairh5 tg t-etl 8suaf.ar ro4 btxtndsenenpeea avwetcenba thsstiiindweP
ness ni lateral .tcepsa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myosides okinawanus sp. nov.
18(17) eht fo semit 4.5 naht erom turts fo esab lartnev dna xepa neewteb ecnatsid ,gnol yrev sineP
.ssenkciht
19(20) no itt a aee.ch ssrt tatafc bum on. resihtditts nsi.eiwPd . . . . . . . . . .ps.von
Myosides yonagunianus
20(19) sineP tuohtiw noitacramed ta eht esab fo.sturts . . . . . . . . . . . . . . . . . . . .. Myosides ishigakianus .ps.von
Key to species by the female genitalia Cyonagunianus, ejimai dna chejuensis era ton)dedulcni
l(2) asruB xirtalupoc htiw a riap fo llams tnadneppa sebol ta eht.esab
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. (Heller)
Myosides marshalli
)02 asruB xirtalupoc tuohtiw hcus.segadneppa
3(6) rediw hcum sebol laretal htiw acehtamreps ,rotisopivo naht retrohs hcum xirtalupoc asruB
naht.gnol
4(5) ac e hhl ttaei a ttwymesalrbilpeo durplogaS tnhlasa seabbol t al,ranerineegnbtrioaallm
tsomlasa kciht as esab fo lasab.ebol . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosidespyrus Sharp
5(4) acehtamrepS htiw latsid ebol ylworran dednuor ta esab ot lasab ebol ta renni ,nigram-retal
drae oer bbloal nahf the base of basal lobe. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .ps.von
Myosides kiiensis
6(3) asruB xirtalupoc tsomla sa gnol sa.rotisopivo
700) acehtamrepS ,tsubor ylneve detalid morf eht devr utcniop fo lasab ebol ot latsi desabfo
lateral lobe.
)9(g acehtamrepS yltrohs devruc ta.xepa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosides amamianus .ps.von
)S(9 d ed al vr.calt rieecaafuhbhito ctotps.laaimdr e.pS . . . . . . . . . . . . .. Myosides okinawanus ..pvson
)7(Ol acehtamrepS htiw lasab ebol.rerednels
)41(ll acehtamrepS ,rellams ecnatsid neewteb esab dna xepa ssel naht 2.0 mm ni thgiarts.htgnel
12(13) aceht a,mtrseeplSl aemcsnatsi dneewteb esab dna xepa 41.0-31.0 mm ni thgiarts,htgnel
distal ebol ylneve .devruc . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosides tokarensis .ps.von
13(12) la t,shit dgt nh8e g1lmi. ma0nr- i t7exnds1seen.apea 0bawetcenba thastciiedwhtamrepS
e. bttohslgoimal ra.ts . . . . . . ..__........................................................ (Heller)
Myosides marshalli
14(11) acehtamrepS ,regral ecnatsid neewteb esab dna xepa erom naht 22.0 mm ni thgiarts.htgnel
15(16) .ebol lasab naht rekciht hcum ebol latsid htiw acehtamrepS Myosides formosanus.von .ps
16(15) acehtamrepS htiw latsid ebol ylecracs rekciht naht lasab.ebol
17(18) acehtamrepS htiw latsid ebol yletalugna devruc ta latsid.driht
.._.......................................... Myosides ishigukiunus sp. nov.
18(17) acehtamrepS htiw latsid ebol ylneve devruc ro tsomla.thgiarts
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Myosides seriehispidus sfoleoR
19(20) acehtamrepS htiw latsid ebol tsomla .thgiarts . . . snemicepS(.. morf ,atagi i,Nara NojihcaH).I
20(19) acehtamrepS htiw latsid ebol erom sselro .devruc
21(22) acehtamrepS htiw latsid ebol regnol naht lasab ebol dna ylneve.devruc
46 K. MORIMOTO & C. E. LEE
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . (Specimens from )uhsuyK
22(21) acehtamrepS htiw latsid ebol sa gnol sa ro a elttil retrohs naht lasab.ebol
23(24) acehtamrepS htiw latsid ebol retrohs naht lasab ,ebol tsuj lacirdnilyc ta lasab.flah
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .sne m.icepS( morf )aeroK
24(23) .yllatsid derepat ylthgils ebol latsid htiw acehtamrepS
. . . . . . . . . . . . . . . . . . . . . . .. pS( snemice morf ihcoK dna arawasagO ).s1~1
Myosides formosanas sp. nay.
-k rraemk rha tddiewlt tgonmil ayc eshrsginw ohorstbiy eersg nhe td,inww onorwt bokrrbaD
s gsnai mutonorp:swollof hlecattsiaan pbi a fhr,ette diyphnwlisieri tnn swaeoitrdiben mi fhetdinwi
neewteb dr3 airts dna ,redluohs euqilbo trohs lasabbus dnab neewteb htruof lavretni dna edis,nigram
tniojnoc depahs-V daorb dnab dniheb eht ,elddim dna netfo htiw ralugerri etinifedni kradaera
erofeb ,xepa hsiyerg dnab dniheb krad naidemrtsop dnab netfo.etinifed
4 ro 3 htiw ,seye neewteb talf daeH tcerebus murtsoR .seye evoba edis hcae no selacs S/4 semit
sa gnol sa ,daor bylk adeewworra nmorf esab ot ,eld ddiem dniesh-tlel ltaarap sedis fo;aigyretp
- rraomire tnneaew tleeb v eeernhlrooto feelb tg tnai il dydnleadn ueolr gdn na sihth tieimwotsipe
snig fo ,aigyret phtiw daorb erab ecaps dni h,eebmots inpaeidem anirac ni erab eca.pestinifedni
e tsheae n etenl ye tmirncsf lmssiwhenisogtga 1tt tanenhet,nhu logsn dAtfsliedaalmnosg2rebs
eciwt sa gnol sa ,daorb dr3 dna ht4 stnemges fo eht emas ,ezis 8/7 semit sa gnol sa ,daorb ht5 dnaht6
stn e smdagaeos rsba r on or i ee at ndlfr,aatoogh o tpnthrei oo tbhltr lmgtnplonearemsflfgaoebs
el c:i1n 2u:f31 :8 :8 :7 :7.8
seru tse2 ,ces8tht enm r.u tssluiea0ogi ead pthp-bnwxdd,t s3aoaaids7lromaa.oro0hbrtborP
,elddim eht ta tsedaorb ,daorb sa gnol sa semit 2.1 tuoba ,lavo artylE .hcae elacs tnebmucer gniraeb
ylneve dednu o,rylrorietsop dr3 lavretni tsomla sa daorb sa dna no eht emas enalp htiw ht4 lavretnita
base, tcerebus selacs tuoba sa gnol sa eerht dnuorg selacs ,denibmoc detalid.yllatsid
latoT htgnel fo ydob 1.4-4.3 ,mm htgnel fo mutonorp dna artyle 4.3-9.2.mm
:epytoloH elamef epyT( ,1772.oN uhsuyK ,).vinU ,nahsgnimgnaY raen ,iepiaT,nawiaT
,5691.v.52 .K otomiroM.gel
.gel otomiroM .K ,5691.v.72 ,.sH iepiaT ,iarU ,selamef 2 :sepytaraP & arahikaM.H ,6791.v.13
;.gel ,1ela maePf ,noI oaT nauY ,.sH ,6891.iiiv.3 .K abaB.gel
Distribution: .nawiaT
Myosides marshalli (Heller) comb. nav.
Heller, ,.tieZ .tne .neiW ,1391 89:84 .)oakaT:nawiaT(
Trachyodes marshalli
daeH dna mutonorp tsomla eht emas sa .M artylE a elttil ,reworran tsomla-thgiarts
formosanus.
yl deworr aynlroiretsop morf lacipa l/3, neht dednuor ta tniojnoc ,secipa dr3 lavretniyltcnitsid
redaorb naht gnirobhgi esnlavetni ta esab dna xnetfo ylkaew.desiar
elaM sugadea htiw sinep tsetrohs gnoma seiceps fo siht suneg ,denimaxe ecnatsid neewtebxepa
REVISION OF MYOSIDES 65
elamef ni xirtalupoc asruB .ssenkciht mumixam sa taerg sa semit 7.2 tuoba turts fo esab lartnev dna
htiw a riap fo llams tnadneppa sebol tsuj dniheb ,erop oancoeghtamreps ton devruc ta.xepa
.mm 0.3-4.2 artyle dna mutonorp fo htgnel ,mm 4.3-1.3 :htgnel ydob latoT
Specimens examined: 5 exs, ,uhihcneF iaihC ,.sH ,6691.iiv.52-42 .H ijasaS ;.gel 1 ,.xe,ihcuhC
iaih C, 6,6.9s1H.ii v..H12 ijasa S;.gel 1 ,gn i,P .-ixiaeLi-hoC a ,i,5h.6Cs9H1.vi ..3S1 oneU ; .g1el
,.xe nahsuL ,nauhcneW uotnaN ,.sH ,0891.v.72 .H arahikaM ;.gel 1 ,.xe ,)hoyraH(gnilaB nayoaT
,.sH ,8891.iiv.42 .S arumoN ;.gel 1 ,.xe ,gnilurznauK naniaT ,.sH ,5691.vi.6 .Y ekatayiM.gel
Distribution: .nawiaT
Note:eht ni eroponog erofeb sebol tnadneppa fo riap a dna elam eht ni sinep tsubor dna trohS
selamef e:rcaitsiretcara hoct siht.seiceps
Myosides yonagunianus sp. nov.
yr eeVsolc ot .M tsurbe fmfoirdf ti yb g :ne,sihrtwt enolgilaroltapooLfytdob
formosanus,
ht g3n.e4l-0. 4,mm htgnel f omutono rdpna ar t6y.l3e-5. 3;mm oitar fo htgnel yb htdiw foartyle
dteas iyalrk aneew t dfsnolaavre tgnniirobhg ine anrhet dyilwtcnit sliadvre t dnhri t3airwt y,l9e1.1
.esab sineP ,rednels ecnatsid neewteb xepa dna lartnev esab fo turts tuoba 4.5 semit sa taerg sa-kciht
,ssen htiw tcnitsid noitacramed neewteb sinep dna.sturts
:epy teolla omeHpyT( .oN ,2 7u7h 2s)u.yvK i:n sUdenpayta r2a P,s e,ilinaaumngoaSn,o.YI
,4791.iv.82 .T egakiM.gel
Distribution: napaJ inuganoY:suykuyR( ).I
Myosides ishigakianus sp. nov.
etinifedni rekrad htiw delttom gnilacs yerg-hsa esned htiw ,kcalb ot nworb tuntsehc krad mreD
,sgnikram mutonorp tuohtiw naidem rekrad ,epirts artyle htiw eugav krad sdnab ta lasab,driht
dniheb eht elddim dna no ,ytivilced tub eseht sdnab ssel etinifed naht eht rehto seiceps ronetfo
,n teyhn t ltekfheisroctwba te a dsaltemeppaku lm itea o nlohcec;o-ntsslaoaa tr bvfeptrossertinfi
no yerg-hsa netfo ,nworb krad ,seiceps gnidecerp eht naht resned dna regnol elttil a ,tcere artyle dna
ytivilced fo.artyle
da e yHdlekwaoe rwmroarnf esab ot e h,teld d ni yemydlhlettldhangraiepl txs,taeaaligyretp
, asing i yr fegdr oho mre egi td oadstnreinnd tmntpielwaennsu iytg eioiollnnewhpreaahtevteebl
erab ecap sdn iahiegby ryeltepnif ,etatcnup htiw a ka etw rnoahisdem .anirac eannet nhAtiwdn2
tnemge sf oelcinuf a e lrtettirl onhasht ,tsl d rt3nemges a el trteigln onla h,tdaor bh tt4nemgessa
gnol sa.daorb
,el dted uhitotmsbead a,odrebd nyulotrh gsi el,ds dis g as esnso5maoar8ilb.t0- 0m7u.t0onorP
roiretna dna roiretsop snigram fo eht emas ,htdiw .etacnurt artylE ,lavo tsedaorb ta eht,elddim
ylneve dednuor ecneht ,ylroiretsop dr3 lavretni tsomla sa daorb sa ro ylthgils redaorb naht dna noeht
emas enalp sa e hgtni rsolbahvgrieetnni ta ,esab tcere selacs tuoba sa gnol sa eerht dnuorgselacs
,denibmoc ,worran ylthgils detalid.yllatsid
66 K. MORIMOTO & C. E. LEE
elaM sugaedea htiw sinep tsegnol gnoma seiceps fo eht ,suneg ecnatsid neewteb xepadna
lartnev esab fo turts tuoba 6.6 semit sa taerg sa .ssenkc iahctehtamrep Sylgnorts devruc ta.xepa
latoT ydob :htgnel 0.4-3.3 ,mm htgnel fo mutonorp dna artyle 3.3-8.2.mm
,1991.iv.22-81 ,.I ikagihsI ,ekad-annaB .tM ,).vinU uhsuyK ,3772 .oN epyT( elamef :epytoloH
.K otomiroM.gel
,epytoloh htiw ytilacol emas ,sxe 4 ;epytoloh htiw atad emas ,sxe 4 :sepytartaP lS.viii.1991, .K
atagO ;.gel 39 ,sxe emas ytilacol hti w, e,p2y9t9o1l.ovhi.1 2.K amijoK ;.gel 2 ,s,xaewagarakaN
amihsariH .Y ,3691.x.5 ,.I etomoirI & .S ,6891.vi.41 ,.I etomoirI ,eripnaK ,.xe 1 ;.gel otomiroM .K
arumoN.gel
Distribution : nap a:Jsuy kiukyaRg(ihsI dna etomoirI.).sI
Note: ehT rednels sinep si citsiretcarahc ot siht.seiceps
Myosides okinawanus sp. nov.
eost oylrceV .M dna sreff itdub morf -ra hgcniwoll oe f hymtbe ht
yonagunianus ishigakianus,
e:asnr n ebeetttlcincaw Aimnourff dr3 , doetm r d osrhft3tin7lei mnogotems shttn6em gfeoseht
emas epahs dna ,ezis a elttil redaorb naht ,gnol artyle ,lavo dr3 lavretni fo eht emas htdiw dnathgieh
ht igwnir osblhagvireentni ta ,esab tcere eates no mutonorp dna artyle .trohs sineP htiwecnatsid
l ahastcaiebwhtam r.esps Se s nt e2k asm. c seai5 eilart-t shag 8ufatr xn.robtdee4tnnpeseaawvteb
ebol kciht ta eht lasab trap fo laretal.ebol
9 . :l h3 ,hamt-y2mtt ug1d. mgoatn.o3dfnT.roe3b-noemtnl7almyo.lr2ep
:ep yetl oaelmpo yH,aTn w(o.ason -Ni ,i k, 4muOi,7ahk)7gsa.2iutvn yuiu.K-nKtoUMnodeH
,.I ,1991.iii.41 .S arumoN.gel
:sepyataraP 8 ,sxe emas atad htiw ;epytoloh 91 ,sxe ,ukO ,imaginuK awanikO ,.I ,1991.iii.41.S
arumoN ;.gel 22 ,sxe .tM ,ibukureT awanikO ,.I ,5891.iii.51 .S arumoN ;.gel 11 ,.sxe,odnir-eI
awanikO ,.I ,6891.vi.22 8891.x.11 & ,1991.iii.41 .S arumoN ;.gel .tM ,ekademihsiN awanikO ,.I02
;.gel atagO .K ,1991.ii.21 ,.xe 1 ,.gel arumoN .S ,1991.iii.41 ,.sxe ,.sxe3 ,.I awanikO ,odnir-inukoO
,9891.vi.32 .K emuK ;.gel .tM ,)m004(ekadahanoY awanikO ,.I ,7891.x.02 .M iakaS ;.gel 1,.xe
,i rauwhaS n,i8k5 O9,1..Ii i.iTv. 9a1kad i,H.gel 1 ,,1.6x9e1.v ..103 ihca k;a.Ngel 1 ,.ixhesoyeuS
, 7 a8,w9 ka 1.rn,.Tai.iPkIiOv.8 onCU ,.9T891.xi .,a8.m2Iije m,uk KraamPayamu r,;a. . Ds6gxeel
onCU .gel
Distribution: nap a:Jsuyk uaywRa(nikO dna amijemuK.).sI
.ps .von
Myosides amamianus
sa emas eht tsomla noitaroloc mred dna nrettap roloc ylacS .M murtsor dna daeH
okinawanus.
kaew htiw ,elddim eht ot esab eht morf deworran ylkaew murtsor ,seiceps txen eht ot ralimis -yretp
e.aani n gheettlincw Aimnourff dr3 ot shttn6em gfeos eht em aespah sdn ay,rleetzdhiagsoirlbs
artylE .gnol naht ,demrofirypbus sineP .ylroiretsop dednuor ylneve neht ,elddim eht dniheb tsedaorb
esab lartnev dna xepa neewteb ecnatsid ,muitso ta setirelcs kaew fo riap a htiw cas lanretni ,rednels
REVISION OF MYOSIDES 76
f otur t4s.6 semit sa t aaec r erg.he. sthtssatsaemaunrrbkeocpriSht
latoT ydob :htgnel 0.4-7.3 ,mm htgnel fo mutonorp dna artyle 3.3-2.3.mm
:epytoloH elam epyT( .oN ,5772 uhsuyK ,).vinU .tM ,ekadnawuY amihsO-imamA,..I
,7891.v.5 .S arumoN .gel
:sepytaraP 61 ,..sxe 5 & ,7891.v.8 .S arumoN;.gel 1 ,.xe .tM ,ekadiuY amihsO-imamA,.I
,4891.iiiv.Ol .S arumoN ;.gel 11 ,a,em.kisahxdseaownau kk,oo.nTII 2 & ,8891.v.5 .S arumoN.gel
Distribution: napaJ :suykuy Ra(mihsO-imamA dna amihsonukoT.).sI
.ps
Myosides tohzrensis nov.
sd neatbinife drneik rha tddiewlt tgonmil ancws oyrhbt rease nhe td,ihwsikc anolwt bokrrbaD
no ,artyle tuohtiw krad naidem epirts no .mutonorp murtsoR htiw kaew ,aigyretp emotsipereworran
ht6 ot dr3 morf suorefilinom elcinuf htiw eannetnA .aigyretp fo level eht erofeb gnidurtorp llew dna
gnol sa semit 88.0-97.0 mutonorP .ht6 naht regral ylthgils tnemges ht7 ,daorb sa gnol sa ,stnemges
sa ,daorb ylthgils dednuor ta .sedis artylE 33.1-91.1 semit sa gnol sa ,daorb ,demrofirypbustsedaorb
d sa laosaravbre td,nrsi3ec itpnaioj nd on ycuolotrroiret sdoepre pyaltthgia rn te,shdtr ilhatc itpaa
,esab ta desiar ton dna slavretni gnirobhgien tcerebus selacs dnuorg eerht sa gnol sa artyle no selacs
detalid ylkaew ,denibmoc .yllacipa
-ll aamcsehtamr e.px Ser padaeennrut -ypluk ase i wrnfueoopt nloacrt nh etsviuwgae deelaaM
tse ni siht ,suneg latsid ebol ylneve.devruc
latoT ydob :htgnel 5.3-0.3 ,mm htgnel fo mutonorp dna artyle 0.3-5.2.mm
:epytoloH elam epyT( .oN ,6772 uhsuyK ,).vinU amihsonakaN ,.I arakoT ,.sI ,7891.v.2.S
arumoN.gel
:sepytaraP 81 ,sxe emas atad sa ;epytoloh 51 ,.sxe ,eki-hO amijarakaT ,.I arakoT ,.sI -02
,2991.iii.52 .S arumoN ;.gel 1 ,.xe ,ikazikarA amijarakaT ,.I ,2991.iii.22 .S arumoN ;.gel 1 ,.xe amhO
,erohsaes amijarakaT ,.I arakoT ,.sI ,2991.iii.42 .S arumoN.gel
Distribution: napaJ arakoT( :.s1~1 amihsonakaN .I dna amijarakaT.).I
Note: sihT seiceps si citsiretcarahc yb eht rellams dna demrofirypbus lareneg ,epahsdenrut-pu
sinep raen xepa dna eht tsellams.acehtamreps
.ps .von
Myosides ejimai
.ndo ai , et,h h.Ha k sh se ts ceitmilesgyasnirnaa uetlnwwewMndrlrbeoDonncidrreandbbtina
yl tx hengveinel ows.c t s meyedublyertewthos gromirRolarsnf eht esab ot ,ee hlhtdtdikiwamew
a htiw ,htooms emotsipe dniheb ecaps erab ,nigram roiretsop ta dednuor yldaorb emotsipe ,aigyretp
naid ekmaew .anirac eanne tnhnotAiitwrop ofrop )ht deih wtm(ohrtfdn eelsab fo elcinuf :)31(32
:)11(91 :)01(Ol :)01(9 :)01(8 :)11(8 .)21(Ol
m u7st9eo.m n0iso tartgPsn e o , ddlytsd ,aelaa ayeodkdeollrnatnhrndbueaatbedowvirem
,sedis ylthgis xevnoc ,yllasrod htiw esned gnorts,serutcnup
artylE ,lavo-gnolbo 4.1 semit sa gnol sa ,daorb slavretni ,xevnoc htiw enif esned ,serutcnupdr3