Table Of Content

E iliiUJSijjfP'i !;!!! it 214 Review of New World Alfaroa and Old World Alfaropsis (Juglandaceae) 216 Bflis short (0.4—2[—3] short (0-l[-2.5] sS s*“=, ^sr'^sr1(cid:8482)' 218 Novon r„, r„, 220 Novon Volume 20, Number 2 Stone 221 2010 Alfaroa and Alfaropsis (Juglandaceae) as well as a more comprehensive overview of Alfaroa throughout Latin America, make a clear distinction between them untenable. It appears now that we are Historical background. In Manning’s 1952 treat¬ dealing with a series of disjunct populations that are ment of the Juglandaceae for the Flora of Guatemala, Alfaroa costaricensis was the only species recognized. A. guanacastensis is hereby reduced to synonymy referred to as “Engelhardtia” guatemalensis Standi, under A. manningii (Leon, 1953). are, in fact, A. guatemalensis (Standi.) L. 0. Williams & Aastganattiu.n sM ooonfl litynh aew (GWithui alltsietaemmriasll ea& ns MpAe.o ccliimonsaet,an r1sic,9 e7hn0es)i .s n Woisth e“idlse o tmhdeaeatw lithnhaget n“(hMilolloaon tdnyepin egT, iuCi.r”Ar iTTalIYbEaP),.E :1 9C oJsutlay R1i9c5a.1 ,C Ja.r taLgeoo:n P3l4at6a9- Costa Rican Alfaroa that the Guatemalan tree probably bexelpoencgtes da tt hleaat st[ thtoe thspaet cgimeneunss ][ wrehperreesaesn].t. .iat ids istoti nbcet fSitgo^n7e. 21196777 ,( hsoylno.t ynpoev, . DTUYKPEE; :’iscootsytpa eRs,i cAa,. CARp,r .F, ,D U. SE).. species” (Manning, 1952: 354). Manning was unable to Relationship of Alfaroa and Alfaropsis used to distinguish Alfaroa specimens from these two The genus Alfaropsis (Iljinskaya, 1993) is most countries. Flowering and fruiting specimens made commonly recognized as Engelhardia roxburghiana subsequently in the same general area in western Guatemala, as well as Veracruz (Narave, 1983) and Psilocarpeae Nagel emend. Leroy (Manning, 1966, 1978). It shares winged fruits with the Old World Engelhardia (Jacobs, 1960) and New World Oreo- munnea (Manning, 1959). Oreomunnea, on the other Paratypes. GUATEMALA. Quetzaltenango: slopes of hSatonnde,, i2s 00th1e) , celvoesens tt hroeulagthiv eth eo f laAttlefar rohaa s (Mwainngolse s&s, mark, ca. 1600 m, 3 Nov. 1967, D. E. Stone 2210, 2210A drupaceous fruits (Stone, 1977). There has been (DUKE), 18 May 1966, D. E. Stone 2214, 2214A, 2214B, considerable conjecture as to the validity of recog- 2215 (DUKE), 29 May 1971, D. E. Stone 3015 (DUKE). JMManEe.zX c1IaC9lpO8a1. , aDlCo.hn giEa .p raBds.r :e teodS loeJviltveoa t o&lN , BeM.g rpTai.,o .K 1eR0l laeyrko m4n9, 2f19r7o00m 0( D mRUa,K yEo1)n0; rseinlacteiv teh eyto arAel fvairrotuaa ll(yM iamnpnionsgs,i bl1e9 t4o9 ;t elSl taopnaer, t e1x9c7e2p)t, 5 km SE of Jitotol, along rd. to Bochil, Mpio. Jitotol, 1600 m, for the distinctive pistillate flowers and fruits 9 Jan. 1981, D. E. Breedlove & B. T. Keller 49370 (DUKE), (Table 2). Support for recognition of Oersted’s Or¬ 10 Sep. 1981, D. E. Breedlove & B. T. Keller 52670 (DUKE). eomunnea (1856) was provided by Hjelmqvist (1948, M13a5r0. 1m9,6 186, MM. aSr.o 1u9sa6 83,6 M19. S(DouUsKa E3)6. 05 (DUKE),^370 m, 16 b1y9 6S0t)o naen d(1 L97er2o)y e m(1p9h5a1s)i zbiansge dth eo nf rufliot raanl dm poarpttheornlosg yo,f germination, and subsequently by Manchester (1987) Synonymy within Alfaroa manningii Iljinskaya reviewed the taxonomy and phylogeny of I described Alfaroa guanacastensis D. E. Stone in the family and concluded, among other things, that E. 1977 as a segregate of A. manningii based on roxburghiana should be treated as a species of perceived differences in their respective leaf and Alfaroa, namely A. roxburghiana (Wall.) Iljinsk. nut morphology (short vs. long leaf petioles, low vs. pronounced ribs on the fruit husk, respectively) and due perhaps in part to our translation from Russian, geographic ranges (Pacific vs. Atlantic watershed in she did make the case for recognizing this taxon as a Costa Rica). Subsequent collections of these two taxa, link between Engelhardia sect. Engelhardia and the williamsii {Stone 3119) with epidermal papillae surrounding the stomate. —H. Alfaroa mexicana {Stone 2132) with epidermal bars: A-C, E-I = 5.6 pm; D = 33 pm. (N.B. Fig. 3B in Manos & Stone [2001] displays the abaxial leaf surface of Alfaroa