Records of scythridids (Lepidoptera: Scythrididae) from the southeastern Balkan

2007·1.5 MB·

by  Kari Nupponen

©EntomologicaFennica.15March2007
Records of scythridids (Lepidoptera: Scythrididae)
from the southeastern Balkan
KariNupponen,JariJunnilainen,Jari-PekkaKaitila&TimoNupponen
Nupponen,K.,Junnilainen,J.,Kaitila,J.-P.&Nupponen,T.2007:Recordsof
scythridids(Lepidoptera:Scythrididae)fromthesoutheasternBalkan.—Ento-
mol.Fennica18:1–10.
Alistof28speciesembracingabout720specimensofthefamilyScythrididae
fromthesoutheasternBalkanispresented.ThematerialwascollectedfromBul-
gariaandnorthernGreeceduring2000–2005.ScythrisbalcanicaJäckh,1978is
synonimized with S. moldavicella Caradja, 1905, syn. n. The previously un-
knownfemalesofScythrisambustellaBengtsson,1997,S.subaerariella(Stain-
ton,1867)andS.platypyga(Staudinger,1880)aredescribed.Sevenspeciesare
reported as new for Bulgaria and four for Greece. Scythris lafauryi Passerin
d’Entrèves,1986isreportedforthefirsttimefromtheBalkans.Theknowndis-
tributionrangeofeachspeciesisgiven.
Kari Nupponen, Miniatontie 1 B 9, FI-02360 Espoo, Finland; e-mail:
[email protected]
JariJunnilainen,Mahlapolku3,FI-01730Vantaa,Finland
Jari-PekkaKaitila,Kannuskuja8D37,FI-01200Vantaa,Finland
TimoNupponen,Kartanonkuja17B2,FI-02360Espoo,Finland
Received20December2005,accepted31January2006
1.Introduction as seven species new for Bulgaria and four for
Greece.Theresultsofthesestudiesandalistofall
Thescythrididfaunaofcentralandeasternparts recordedscythrididsarepresentedinthispaper.
of the Balkan region is generally rather poorly
known.Todate,45speciesofthefamilyScythri-
didaeareknownfromthatarea,ofwhich38from 2.Thecollectingsites,material
northern Greece and 27 from Bulgaria (see andmethods
Bengtsson1997).Only20speciesareincommon
forthesecountries,whichmayduetoalowcol- Theinvestigatedareaissituatedinsoutheastern
lectingactivityinthearea. part of the Balkan region: eastern and southern
During 2000–2005, Finnish lepidopterists BulgariaandnorthernGreece.Thehabitatswere
madesevenexpeditionstoBulgariaandnorthern mainly open, xerothermic localities like hill
Greece and collected a rather rich material of slopesanddifferentkindsofmeadows.
microlepidoptera, including numerous scythri- The present article is based on the material
dids.ThestudiesonScythrididaerevealedinter- collectedduring2000–2005in7differentexpedi-
estingresults,includingpreviouslyunknownfe- tions. The dates, visited areas and collectors on
malesofthreespecies,onenewsynonym,aswell eachofthetripsareasfollows:
2 Nupponenetal. (cid:127) ENTOMOL.FENNICAVol.18
Fig.1.MapofthesoutheasternBalkanregionwithcollectinglocalities.–1:BG,Asenovgrad.–2:
BG,Elesnica.–3:BG,Jakoruda.–4:BG,Kresna.–5:BG,NosSabla.–6:BG,Orelek.–7:BG,
Pirin.–8:BG,PobitiKameni.–9:BG,PopotiLivadi.–10:BG,Ropotamo.–11:BG,Sliven.–12:
BG,SvetaPetka.–13:BG,Tuzlata.–14:BG,Varvara.–15:GR,Askio.–16:GR,Grevena.–17:
GR,Kozani.–18:GR,Leptokarya1.–19:GR,Leptokarya2.–20:GR,Metamorfosi.–21:GR,
Serres.–22:GR,Vermio.–23:GR,Xirolimni.
– 1:18.–23.VI.2000;SW,C,EandNEBulga- thematerialwascollectedbysweepingandnet-
ria;J.Junnilainen. tingatdaylight.Afewspecimenswerealsocol-
– 2: 21.–27.V.2001; NW Greece; J. Junni- lectedbyartificiallightatnight.Thematerialhas
lainen. been examined by Kari Nupponen and Jari
– 3: 14.–27.VI.2001; SW, C, E and NE Bul- Junnilainen.Thecollectingsitesareshowninthe
garia;J.Junnilainen,J-P.Kaitila. map(Fig.1).
– 4:26.V.–09.VI.2002;SW,C,EandNEBul- Thecollectinglocalitiesarementionedbelow.
garia;J.Junnilainen,J-P.Kaitila. Briefvariantsoflocalitynamesaregiveninup-
– 5:04.–10.VIII.2002;SW,C,EandNEBul- percaselettersbeforeeachlocalityandusedlater
garia;J.Junnilainen. inthespecieslist.Thenumbergiventoeachof
– 6: 18.–24.V.2003; NW Greece; J. Junni- the localities is connected with that on the map
lainen,J-P.Kaitila. (Fig.1).
– 7: 03.–07.VI.2005; NW and NE Greece; T.
Nupponen. 1, BG,ASENOVGRAD:SBulgaria,42°02’N
24°52’E, 210 m, near Asenovgrad village.
Thematerial,altogetherabout720specimensof Calcareous, rocky slopes and a river valley
scythridids,isdepositedintheprivatecollections with diversified low vegetation. 30.V.2002,
oftheobservers.Theoverwhelmingmajorityof 10.VIII.2002.
ENTOMOL.FENNICAVol.18 (cid:127) ScythrididsfromthesoutheasternBalkan 3
2, BG, ELESNICA: SW Bulgaria, 41°52’N 25.VI.2001, 02.–03.VI.2002, 04.–05.VIII.
23°37’E,800m,nearElesnicavillage.Mead- 2002,10.VIII.2002.
owsandsandypastureforsheepwithdiversi- 12, BG,SVETAPETKA:SWBulgaria,42°02’N
fiedlowvegetation(Helianthemum,Thymus, 24°03’E, 800 m, near Sveta Petka village.
Genista,etc.).24.VI.2001. Grassy forest steppe slopes with diversified
3, BG, JAKORUDA: SW Bulgaria, 42°00’N low vegetation (Helianthemum, Thymus,
23°38’E,930m,nearJakorudavillage.For- Genista,etc.).31.V.2002,09.VIII.2002.
eststeppesbetweenthemountainsofRilaand 13, BG, TUZLATA: NE Bulgaria, 43°12’N
Rodopi.23.VI.2000,25.VI.2001. 27°57’E,5–30m,nearTuzlatavillage.Chalk
4, BG, KRESNA: SW Bulgaria, 41°43’N hillsclosetoBlackSeashorewithdiversified
23°09’E,200m,nearKresnavillage.Ariver low vegetation. 14.–16.VI.2001, 26.–27.VI.
valleyonfoothillsofthePirinMountains.A 2001, 26.V.2002, 28.V.2002, 05.VI.2002,
xerothermic locality with various kinds of 08.–09.VI.2002,10.VIII.2002.
dry meadows, railway banks, rocky and/or 14, BG, VARVARA: SW Bulgaria, 42°08’N
bushyslopesandruderalplacesatroadside. 24°07’E,480m,nearVarvaravillage.Ariver
21.–22.VI.2000, 21.VI.2001, 06.–07.VIII. valley withsteepslopesanddiversifiedlow
2002. vegetation.31.V.2002,06.VIII.2002.
5, BG, NOS SABLA: NE Bulgaria, 43°33’N 15, GR,ASKIO:NWGreece,prov.Makedonia,
28°35’E, 0–10 m, near Nos Sabla village. Kozani NW, 40°22’N 21°30’E, 1450 m,
Sand dunes, moist meadows and wetlands Askio Mountains. Open, grassy meadows
nearBlackSeashoreclosetoRomanianbor- androckypatches.23.V.2003.
der.27.V.2002. 16, GR, GREVENA: NW Greece, prov. Make-
6, BG, ORELEK: SW Bulgaria, 41°33’N donia,Grevena,39°49’N21°09’E,1,430m,
23°37’E, 1,800 m, southern part of Pirin Grevenatown35kmS.Ahillareawithoak
Mountains, near Orelek village. Calcareous forestsanddrymeadows.24.V.2001.
mountaintundrawithluxuriouslowvegeta- 17, GR, KOZANI: NW Greece, prov. Make-
tion.22.VI.2001,08.VIII.2002. donia, Kozani, 40°13’N 21°49’E, 480 m,
7, BG,PIRIN:SWBulgaria,41°32’N23°31’E, nearKozanitown.Open,xerothermicslopes
760 m, Pirin Mountains, near Pirin village. with diversified low vegetation (Thymus,
Dry meadows and bushy slopes. 21.– Astragalus, Helianthemum, Aster, Silene,
24.VI.2001. etc.).23.–24.V.2003.
8, BG, POBITI KAMENI: NE Bulgaria, 18, GR, LEPTOKARYA 1: NW Greece, prov.
43°12’N27°40’E,100m,PobitiKameni.A Makedonia,Pieriá,40°01’N22°29’E,750m,
relict-like area of continental sand dunes. Leptokarya village near Mount Olympos.
30.V.2002,04.VI.2002. Differentkindsofmeadowsandbushyslopes
9, BG, POPOTI LIVADI: SW Bulgaria, withinterestingvegetation(Ilex,Abies,Thy-
41°37’N 23°19’E, 900–1,300 m, Pirin mus, Scleranthus, Helianthemum, Viola,
Mountains, near Popoti Livadi village. De- etc.).21.–23.V.2001,27.V.2001.
ciduousforests(Ulmus)andbushymeadows. 19, GR, LEPTOKARYA 2: NE Greece, prov.
21.–24.VI.2001. Thraki,Evros,41°01’N26°03’E,370m,near
10, BG, ROPOTAMO: SE Bulgaria, 42°19’N Leptokarya village. Deciduous forests,
27°44’E, 10–20 m, Burgas 40 km SE, xerothermicopenslopesandmeadows.05.–
Ropotamo. Amosaicofcoastalsand dunes, 07.VI.2005.
old deciduous forests (Quercus, Ulmus, 20, GR, METAMORFOSI: NW Greece, prov.
Fraxinus), meadows, moist patches and Makedonia, Kozani, 40°22’N 21°43’E, 990
wetlands.16.–20.VI.2001,05.–07.VI.2002. m,KozanitownNW,nearMetamorfosivil-
11, BG,SLIVEN:CBulgaria,42°41’N26°22’E, lage. Rocky steppe slopes with interesting
300m,southernslopeofSlivenskaPlanina. lowvegetation(Thymus,Scleranthus,Stipa,
Rocky slopes and deciduous forests (Quer- Eryngium,Helianthemum,etc.).22.V.2003.
cus, Ulmus). 18.–19.VI.2000, 23.VI. 2000, 21, GR, SERRES: N Greece, prov. Makedonia,
4 Nupponenetal. (cid:127) ENTOMOL.FENNICAVol.18
Serres,41°16’N23°04’E,140m,Serrestown Distribution.CandSEurope,Balticcountries
41kmW,Promahonas.Roadsidemeadows, and Finland eastwards to S Siberia (Baikal re-
railwaybanks,rockyandbushyslopesclose gion).
toBulgarianborder.25.–26.V.2001.
22, GR, VERMIO: NW Greece, prov. Make- ScythrismoldavicellaCaradja,1905
donia, Imathia, 40°29’N 22°03’E, 1350 m, ScythrisbalcanicaJäckh,1978Syn.n.
KozanitownNE,VermioMountains.Bushy ScythrisottomanaJäckh,1978
meadowsandpasturegrounds.22.V.2003. BG,PobitiKameni,30.V.200225##2$$,
23, GR,XIROLIMNI:NWGreece,prov.Make- [JJ];04.VI.20021#,[JPK]
donia, Kozani, 40°18’N 21°35’E, 1,000 m, BG,Kresna,31.V.–02.VI.20027##,[JJ];9
nearXirolimnivillage.Rocky steppeslopes ##1$,[JPK]
with diversified low vegetation (Thymus, BG,Tuzlata,05.VI.2002and08.–09.VI.2002
Scleranthus, Stipa, Eryngium, Helianthe- 2##1$,[JJ]
mum, Dianthus, Artemisia, etc.). 21.–23.V. BG,Pirin,21.–24.VI.20011#,[JJ]
2003,04.–05.VI.2005. GR,Grevena,24.V.20012##1$,[JJ]
GR,Xirolimni,04.VI.20051#,[TN]
Genitaliaslide:K.Nupponenprep.no.2/15.
3.Listofscythrididspecies II.2005 ($); three genitalia preparations (##)
preservedinglycerol.
Thespeciesarepresentedinaputative,naturalse- Distribution. Bulgaria, Romania, Albania,
quence introduced in the recent review of the formerYugoslavia,Greece,Turkey.Thespecies
Scythrididae of Europe s. l. (Bengtsson 1997). isreportedalsofromPortugal(Bengtsson1991,
Theknowndistributionforeachspeciesisgiven, Fauna Europaea 2005), but according to
based on data by Bengtsson (1997) and Fauna Bengtsson(1997)therecordsfromtherearein-
Europaea(2005).Theobserverofeachrecordis correct.
mentionedinsquarebracketsasfollows:[JJ]=J. Remarks.S.balcanicaisapoorlyknownspe-
Junnilainen, [JPK] = J-P. Kaitila, [TN] = T. cies and its status has been confusing, because
Nupponen. onlyafewfemaleswereavailableandamalere-
Scythris cuspidella (Denis & Schiffermüller,
1775) Fig.2.Female
genitaliaof
BG,Sliven,18.–19.VI.2000common,[JJ]
Scythris
BG,Jakoruda,25.VI.2001common,[JJ]
moldavicella
BG,Pirin,21.–24.VI.2001common,[JJ]
Caradja,1905
BG, Ropotamo, 26.–27.VI.2001 common,
[JJ]
Distribution.CandSEurope,Russia(Chita
region,Yakutia,Amur).
Scythrisflavilaterella(Fuchs,1886)
BG,Sliven,18.–19.VI.20002##1$,[JJ]
Distribution.CandSEurope,Caucasus,Rus-
sia(SUral).
Remarks.NewtoBulgaria.
Scythrisobscurella(Scopoli,1763)
BG,PopotiLivadi,21.–24.VI.20011#1$,
[JJ]
BG,Ropotamo,26.–27.VI.20011#,[JJ]
GR,Leptokarya1,21.–23.V.20011#,[JJ]
ENTOMOL.FENNICAVol.18 (cid:127) ScythrididsfromthesoutheasternBalkan 5
mainedunknown.Furthermore,thesuperficially Remarks.S.ambustellaispreviouslyknown
similarS.moldavicellaiswidespreadinthesame by the holotype (male) only. Bengtsson (1997)
regionwherebalcanicaoccurs,butthefemaleof provisionally placed the taxon in the knochella
moldavicellahasneverbeendescribed.Thepres- species-group. There are some details in the fe-
entmaterial,includingaseriesofbothmalesand malegenitaliaofS.ambustellathatdonotexistin
females from several localities in Bulgaria, al- Europeanspeciesofthegroup,likeasclerotized
lows to study the external variation of S. mol- antrumandamedioposteriorprocessinsternum
davicella, as well as genitalia of both sexes. In VII.However,thosecharacteristicsarepresentin
threelocalities,malesandfemaleswereobserved three Central Palaearctic species (S. elenae K.
simultaneouslyandthereisnodoubtthattheybe- Nupponen,2000, S.hamardabanicaK.Nuppo-
longtothesametaxon.Themalegenitaliaequal nen,2003andS.jueriveteiK.Nupponen,2005;
tothoseofS.moldavicella(holotype)illustrated see Nupponen et. al. 2000, 2005, Nupponen
byBengtsson(1997),andthefemalegenitaliaex- 2003,2005),whichsupposedtoberelatedtoei-
actly coincide with those of the holotype of S. ther cicadella or knochella species-groups. The
balcanica(seeBengtsson1997)(Fig.2).Thus,S. latter character also exists in S. moldavicella
balcanicaisajuniorsynonymofS.moldavicella. Caradja,1905(seebelow)belongingtothecica-
S. moldavicella is externally a variable spe- della-group. Thus, S. ambustella probably be-
cies. Especially a white streak in fold of the longstotheknochella-groupasBengtsson(1997)
forewing is variable. In some specimens it is presumed, but the diagnostic characters of the
broad,distinctandfusedtoadashabovetornus, cicadella and knochella species-groups overlap
andsometimessubapicalandapicalspotsofsame (seeNupponen2003).
colour are present. On the other hand, in some ExternallythefemaleofS.ambustellaissimi-
casesthestreakisreducedtoafewpalescalesin lar to the male, but somewhat smaller in size
foldfrombasetomidwing.Theexternalappear- (wingspan 11.5 mm; 12–13 mm in male) and
anceoffemaleequalstothatofmale,excluding hindwingsareyellowishbrown(blackishbrown
theabdomen.Theabdomenoffemaleisfuscous, inmale)(Fig.3).Theabdomenoffemaleisdor-
ventrally with scattered dirty whitish scales oc-
curringmoredenselylaterallyandalmostcover-
ingventralsegmentsVI–VII.Inthefemalegeni-
talia,sternumVIIissubrectangularwithslightly
concave posterior margin and a rather membra-
nous,ovalmedioposteriorprocess.
Scythrispotentillella(Zeller,1847)
BG,Kresna,21.–22.VI.20001#,[JJ]
Distribution.Europe,AsiaMinor.
Remarks.NewtoBulgaria.
Scythrislimbella(Fabricius,1775)
BG,Pirin,21.–24.VI.20011$,[JJ]
BG,Sliven,05.VIII.20021ex.,[JJ]
Distribution.WPalaearctic,ENearctic.
ScythrisambustellaBengtsson,1997
GR, Xirolimni, 21.–23.V.2003 3 ## 1 $,
[JJ]; Genitalia slide: K. Nupponen prep. no.
2/12.IV.2004 ($); one genitalia preparation (#)
preservedinglycerol.
GR,Metamorfosi,22.V.20031#,[JPK] Fig.3.Female(up)andmale(down)ofScythris
Distribution.Greece. ambustellaBengtsson,1997.
6 Nupponenetal. (cid:127) ENTOMOL.FENNICAVol.18
Scythrispudorinella(Möschler,1866)
GR,Kozani,23.–24.V.20033##,[JJ]
Distribution. Bulgaria, Greece, Romania,
Moldova, Ukraine, Belarus, Russia (S Ural,
lowerVolgaregion,AltaiMnts),Turkey.
Scythrissubaerariella(Stainton,1867)
BG,PobitiKameni,04.VI.2002severalexx.,
[JJ];5exx.,[JPK]
BG, Tuzlata, 05.VI.2002 1 ex., 27.–28.VI.
20011ex.,[JPK]
GR, Xirolimni, 21.–23.V.2003 1 $, [JJ];
04.VI.20051#,[TN];Genitaliaslide:K.Nuppo-
nenprep.no.3/12.IV.2004($).
Distribution. Romania, Bulgaria, Greece,
Turkey,Russia(SUral).
Remarks.AlthoughS.subaerariellaiswidely
distributedandlocally rathercommoninsouth-
easternEuropeandTurkey,theexternalappear-
Fig.4.FemalegenitaliaofScythrisambustella(prep. anceandgenitaliaofthefemalehaveneverbeen
no.2/12.IV.2004KN).
described. The external appearance and size of
thefemale(wingspan13mm)issimilartothatof
themale,excludingundersideofhindwingsand
sally fuscous, more or less densely covered by the abdomen. The underside of hindwing is
rustybrownscales;ventrallyfuscousexceptseg- fuscousinbothsexes,butinfemalethebasalhalf
mentsVIandVIIthatarepaleyellowishbrown. isdenselycoveredbydarkbrownscales.Theab-
Descriptionoffemalegenitalia(Fig.4).Ste- domenoffemaleisfuscous;rusty brownscales
rigma oval, medially membranous. Antrum
sclerotized, distally wide. Sternum VII sub-
rectangular,medioposteriorlyincisedandwitha
flattenedstructure;anteriormarginmediallycon-
cave,withsmallbulgesnearcorners.Apophyses
anteriores0.55×lengthofapophysesposteriores.
Scythrisflabella(Mann,1861)
BG, Kresna, 21.–22.VI.2000 common, [JJ],
22.VI.20011#,[JPK]
Distribution. Bulgaria, former Yugoslavia,
Moldova,Ukraine,Turkey,Armenia,Iran.
ScythrisalbostriataHannemann,1961
GR,Serres,25.–26.V.20011ex,[JJ]
Distribution.Albania,Greece,Romania,for-
merYugoslavia.
Scythrisconfluens(Staudinger,1871)
GR,Xirolimni,21.–23.V.20031#2$$,[JJ]
GR,Kozani,23.–24.V.20031#2$$,[JJ]
GR,Xirolimni,04.VI.20051#2$$,[TN] Fig.5.FemalegenitaliaofScythrissubaerariella
Distribution.Greece,Makedonia. (Stainton,1867)(prep.no.3/12.IV.2004KN).
ENTOMOL.FENNICAVol.18 (cid:127) ScythrididsfromthesoutheasternBalkan 7
dorsally at segments I–IV; ventrally with scat- Scythrisanomaloptera(Staudinger,1880)
teredwhitescales,analtuftpaleyellowishwhite. BG,Jakoruda,25.VI.20011#,[JJ]
OurspecimensfromGreeceandBulgariaareless BG,Tuzlata,14.–16.VI.20012##1$,[JJ],
mottledthanspecimensfromTurkeyandSUral, 16.VI.20011#,26.V.20024##,[JPK]
aspalebrownscalesontheforewingsarescarce GR,Leptokarya2,06.VI.20051#1$,[TN]
orevenabsent. Distribution.Spain,France,Yugoslavia,Tur-
Descriptionoffemalegenitalia(Fig.5).Ste- key. Recently recorded from S Russia, lower
rigma subquadrangular, median part membra- Volga region (Volgograd distr., Olhovka village
nous; anterior margin straight and strongly 20kmS,13exx.15.V.2005,K.Nupponenleg.).
sclerotized; lateral margins concave; posterior Remarks.NewtoBulgaria,GreeceandRus-
cornersbristled;posteriormarginextended,me- sia.
diallywithamoresclerotized,medioposteriorly
incisedstructurefusedtosterigma.SternumVII ScythrishungaricellaRebel,1917
subrectangular,posteriorcornersrounded;poste- BG,Asenovgrad,20.VI.20001#,[JJ]
rior margin medially incised and with a sub- BG, Ropotamo, 16.–20.VI.2001 common,
trapezoid process. Apophyses anteriores 0.6 × [JJ]
lengthofapophysesposteriores. BG, Tuzlata, 27.–28.VI.2001 common, [JJ];
05.VI.20022##,[JPK]
Scythrisclavella(Zeller,1855) Distribution. Slovakia, Hungary, Romania,
BG,commonandwidelydistributed:Kresna, Bulgaria, Croatia, former Yugoslavia, Greece,
Ropotamo,Sliven,SvetaPetka,Tuzlata,Varvara, Syria.
[JJ]
GR,Kozani23.–24.V.2003,common,[JJ] Scythristergestinella(Zeller,1855)
Distribution.CandSEuropeeastwardstothe BG,Tuzlata,14.–16.VI.2001and05.VI.2002
southern Ural Mountains; Altai Mountains. Re- common,[JJ]
cently reported from Central Asia (Kazakhstan, GR, Leptokarya 1, 21.–23.V.2001 several
Kirgisia;K.Nupponenet.al.2005). exx.,[JJ]
Remarks.NewtoBulgariaandGreece. GR,Leptokarya2,06.VI.20051#,[TN]
Distribution.SouthernCandSEurope.
Scythrisseliniella(Zeller,1839)
BG,Varvara,31.V.20022##,[JJ] Scythrisplatypyga(Staudinger,1880)
BG,Kresna,21.–22.VI.20001$[JJ];31.V.– BG, Tuzlata, 14.–16.VI.2001 2 exx.,
02.VI.20023exx.,[JPK] 05.VI.2002 3 exx., 08.–09.VI.2002 1 ex., [JJ];
BG,Orelek,22.VI.20012exx.,[JPK] 26.V.20015##,[JPK]
BG, Tuzlata, 26.V.2002 1 ex., 05.VI.2002 3 GR, Xirolimni, 21.–23.V.2003 3 $$, [JJ];
exx.,[JPK] Genitalia slide: K. Nupponen prep. no. 1/15.II.
BG,Asenovgrad,20.VI.20001ex.,[JJ] 2005($).
GR, Leptokarya 1, 18.–21.V.2003 2 exx., GR,Kozani,23.–24.V.20031#,[JJ]
[JPK] Distribution.Bulgaria,Turkey.
GR,Vermio,18.V.20039exx.,[JPK] Remarks. New to Greece. Although S.
GR,Askio,23.V.20032exx.,[JPK] platypyga is rather widespread in Turkey and
GR,Kozani,23.–24.V.20032exx.,[JJ] southernBalkans,thefemaleremainedunknown
GR,Xirolimni,21.–23.V.20035exx.,[JPK]; to date. Externally the female is similar to the
04.VI.20052##,05.VI.20051#2$$,[TN] male,excepttheabdomen.Inventralsideofthe
GR,Leptokarya2,05.VI.20052##2$$, female abdomen, the 7th segment is pale beige
06.VI.2005 10 ## 5 $$, 07.VI.2005 14 ##, andsamecolouredscatteredscalesexistintermi-
[TN] nalpartoftheabdomen.
Distribution.CandSEurope,AsiaMinor. Descriptionoffemalegenitalia(Fig.6).Ste-
rigma large, subtriangular; anterior corners
rounded and slightly extended; anterior margin
8 Nupponenetal. (cid:127) ENTOMOL.FENNICAVol.18
italiaslide:K.Nupponenprep.no.3/08.IV.2004.
Distribution.Greece.
Remarks. S. eberhardi is previously known
bytwomalesfromsouthernGreece(seeBengts-
son 1997). However, the taxon appears to be
ratherwidelydistributedinGreeceandlocallyit
isnotrareinnorthwesternpartofthecountry.The
moth occurs in xerothermic, rocky slopes. The
mainflightperiodislateMay.
Scythristributella(Zeller,1847)
BG, Tuzlata, 05.VI.2002 common, [JJ],
16.VI.20011$,[JPK]
BG,NosSabla,27.V.20021#,[JPK],1#3
$$,[JJ]
BG,Ropotamo,05.–07.VI.2002severalexx.,
[JJ]
GR,Leptokarya1,25.VII.2001severalexx.,
[JJ]
GR,Grevena,24.V.2001severalexx.,[JJ]
GR,Xirolimni,21.–23.V.20032exx.,[JJ]
Distribution.CandSEurope,Libya,Russia
(SUral).
Remarks.NewtoBulgaria.RecordsofS.tri-
butella from Estonia (Teich 1899, Luig &
Fig.6.FemalegenitaliaofScythrisplatypyga
Kesküla1995,FaunaEuropaea2005)arebased
(Staudinger,1880)(prep.no.1/15.II.2005KN).
on misidentification. The species was recently
omitted from the Estonian list (Jürivete et. al.
convex. Sternum VIII subrectangular, membra- 2000).
nous. Dorsad segment VIII a large, sclerotized,
subtrapezoid plate. Apophyses anteriores 0.2 × ScythrislafauryiPasserind’Entrèves,1986
lengthofapophysesposteriores. BG,Kresna,21.–22.VI.20004##,1$,[JJ].
Genitalia slide: K. Nupponen prep. no. 1/12.IV.
Scythrisinertella(Zeller,1855) 2004(Figs.7–8).
BG,Tuzlata,14.–16.VI.2001common,[JJ] GR,Leptokarya2,06.VI.200512##7$$,
Distribution. S Europe, Ukraine, Belarus, 07.VI.200510##1$,[TN]
Caucasus,NAfrica,AsiaMinor,MiddleEast. Distribution.Spain,France.
Remarks. The species is previously known
Scythrispicaepennis(Haworth,1828) only from southwestern Europe. Compared to
BG,Pirin,21.–24.VI.20012exx.,[JJ] Spanishspecimens,thepatternoftheforewings
BG,Jakoruda,23.VI.2000severalexx.,[JJ] are more distinct in specimens from Balkans
GR,Serres,25.–26.V.20011ex.,[JJ] (Fig.9).However,thereisnoregionalvariation
Distribution.CandSEurope,SScandinavia, in the genitalia. The present records extend the
Estonia,Caucasus,Japan. knowndistributionrangeofthetaxonconsider-
ablytotheeast.ThehabitatsbothinBulgariaand
ScythriseberhardiBengtsson,1997 in Greece are xerothermic gravel pits. New to
GR,Serres,25.–26.V.20012##,[JJ] BulgariaandGreece.
GR,Grevena,24.V.20018##,[JJ]
GR, Xirolimni, 21.–23.V.2003 1 #, [JJ];
03.VI.20058##,04.VI.20052##,[TN];Gen-
ENTOMOL.FENNICAVol.18 (cid:127) ScythrididsfromthesoutheasternBalkan 9
Scythrisgravatella(Zeller,1847)
BG, Tuzlata, 14.–16.VI.2001 common, [JJ],
15.–16.VI.2001 10 exx., 27.–28.VI.2001 1 ex.,
26.V.20032exx.,05.V.20022exx.,[JPK]
BG,SvetaPetka,31.V.2002severalexx.,[JJ]
GR,Serres,25.–26.V.2001severalexx.,[JJ]
GR,Metamorfosi,22.V.20031#,[JPK]
GR,Xirolimni,21.–23.V.20035exx.,[JPK]
GR,Kozani,23.–24.V.20031#,[JJ]
GR, Leptokarya 2, 05.VI.2005 1 #, 06.VI.
200510##1$,07.VI.20054##,[TN]
Distribution.SEurope,Tunisia.
Fig.7.MalegenitaliaofScythrislafauryiPasserin
Scythristabidella(Herrich-Schäffer,1855)
d’Entrèves,1986(prep.no.1/12.IV.2004KN).
BG, Sliven, 18.–19.VI.2000 common, [JJ],
31.V.–02.VI.20022##,[JPK]
BG,Kresna,21.–22.VI.2000common,[JJ]
BG,Pirin,21.–24.VI.2001common,[JJ]
Distribution.CandSEurope,CAsia.
Remarks.NewtoBulgaria.
Scythrisfallacella(Schläger,1847)
BG,Jakoruda,23.VI.2000severalexx.,[JJ]
Distribution. S and C Europe, Turkey. Re-
ported also from the eastern part of European
Russia(FaunaEuropaea2005).
Fig.8.TergumVIII(left)andsternumVIII(right)of
Episcythristriangulella(Ragonot,1875)
Scythrislafauryi(prep.no.1/12.IV.2004KN).
BG,Sliven,23.VI.20001#,[JJ]
Distribution. Morocco, Portugal, Corsica,
Sardinia,Sicily,Greece,Bulgaria,Romania.
Acknowledgements. WethankKimmoSilvonen(Espoo,
Finland)andFinnishMuseumofNaturalHistory,Univer-
sity of Helsinki for their help in processing the photo-
graphs,aswellasUrmasJürivete(Tallinn,Estonia),Ole
Karsholt(Copenhagen,Denmark)andananonymousref-
ereeforusefulcommentsonthemanuscript,andFaunatica
Oyforsupportingthiswork.
Fig.9.Imago(male)ofScythrislafauryifromBulgaria. References
Bengtsson,B.Å.1991:Contributiontotheknowledgeof
the scythridid fauna of SW Europe (Lepidoptera:
ScythriscryptaHannemann,1961 Scythrididae).—SHILAPRevtaLepid.19:35–51.
BG,Elesnica,24.VI.20011#,[JJ];Genitalia Bengtsson,B.Å.1997:Scythrididae.—In:Huemer,P.,
Karsholt,O.&Lyneborg,L.(eds.),Microlepidoptera
slide:K.Nupponenprep.no.4/12.IV.2004.
ofEurope2:1–301.
Distribution.Bulgaria,CzechRepublic,Den-
Fauna Europaea 2005: Fauna Europaea. – [www docu-
mark,Greece,Italy,formerYugoslavia,Sweden, ment].URLwww.faunaeur.org.
Turkey. Jürivete,U.,Kaitila,J.,Kesküla,T.,Nupponen,K.,Viida-
10 Nupponenetal. (cid:127) ENTOMOL.FENNICAVol.18
lepp,J.&Öunap,E.2000:EestiLiblikad,Kataloog. with description of two new species (Lepidoptera:
EstonianLepidoptera,Catalogue.—Tallinn.151pp. Scythrididae).—EntomologicaFennica16(3):165–
Luig,J.&Kesküla,T.1995:CatalogusLepidopterorum 174.
Estoniae.—Tartu.130pp. Nupponen,K.,Bengtsson,B.Å.,Kaitila,J.-P.,Nupponen,
Nupponen,K.2003:Contributiontothescythrididfauna T.,Junnilainen,J.&Olschwang,V.2000:Thescythri-
ofsouthernBuryatia,withdescriptionofsevennew did fauna of the southern UralMountains, with de-
species(Lepidoptera:Scythrididae).—Entomologica scription of fourteen new species (Lepidoptera:
Fennica14(1):25–45. Scythrididae).—EntomologicaFennica11(1):5–34.
Nupponen,K.,Jürivete,U.&Pototski,A.2005:Records Teich, C. A. 1899: Vervollständigtes Verzeichnis der
ofscythrididsfromSoutheasternKazakhstan,withde- Schmetterlinge der baltischen Provinzen, soweit sie
scription of five new species (Lepidoptera: Scythri- biszumFrühjahresdesJahres1899bekanntgeworden
didae).—EntomologicaFennica16(2):65–73. sind. — Korrespondenzblatt des Naturforscher-
Nupponen, K. 2005: Notes on Palaearctic Scythrididae, VereinszuRiga42:9–76.