Punctapinella (Lepidoptera: Tortricidae), a new genus for three previously known and three new species from South America

9 Pages·1991·6.8 MB·English

by  John W Brown

PUNCTAPINELLA (LEPIDOPTERA: ToRTRICIDAE),
a New Genus for Three Previously Known
and Three New Species from South America
John W. Brown1
ABSTRACT. Punctapinella, new genus, is proposed for six species from northern South America: P.
conchitis(Meyrick),newcombination(typespecies);P.niphastra(Meyrick),newcombination;P.chiono-
carpa(Meyrick),newcombination;P. braziliana,newspecies;P. scleroductus,newspecies;andP. theta,
new species. Adults of Punctapinella are characterized by a unicolorous bronze-brown to dark brown
forewing with distinct roundish white spots. Putative autapomorphies for the genus include unique
forewingpattern,elongatemedial fusion ofthegnathosarms,andcompoundcornutus. Punctapinellais
morphologicallymostsimilartoSeticostaRazowski,1986,andAnopinellaPowell,1986,amongdescribed
genera. The presence ofa male foreleg hairpencil confirms the assignment ofthe genus to Euliini (Tor-
tricinae). The male of only one species, P. conchitis, is known. It possesses a row of strong spinelike
setae subbasally on the costa of the valva; this character state is interpreted as a synapomorphy for
Punctapinella and Seticosta.
(Additional key words: accessory bursa, Neotropical,Seticosta, Anopinella.)
INTRODUCTION facies. Punctapinella, new genus, is described to
accommodate Eulia chionocarpa Meyrick, Eulia
Within the recentlyredefined tortricid tribe Euliini
(Powell, 1986; Brown, 1990), Seticosta Razowski, niphastra Meyrick, Eulia conchitis Meyrick, and
1986, Anopinella Powell, 1986, and Apolychrosis three previously undescribed species. Females of
Amsel, 1962, form a well defined monophyletic Punctapinella possess a variably developed acces-
group on the basis of the following putative syn- sory bursa from the ductus bursae, unassociated
apomorphies:labialpalpielongateandporrect,male with the ductus seminalis, a character state previ-
antennaewithextremelylongcilia(greaterthan 1.5 ouslyconsideredasynapomorphyforSeticostaand
x annular segment diameter), forewing vein CuP Anopinella(BrownandPowell,unpubl.).Although
absent, valvae narrow and usually subbasally con- accessory bursae are present in Chlidanotini and
stricted with an apical or subapical region of non- Hilarographini (Chlidanotinae) and in Cochylini
deciduous bristlelike setae (analagous with cucul- (Tortricinae), these structures are considered in-
lus), and ductus seminalis from the corpus rather dependently derived among these phylogenetically
thanfrom theductus bursae. Femalesofthisgroup distant groups (Horak, 1984; K. Tuck, personal
are characterized by the presence ofelongate slen- communication). The single male ofPunctapinella
der apophyses posteriores and anteriores, a mod- possessesarowoflong,strong,spinelikesetaesub-
erately long ductus bursae, and the absence of a basally on the costa ofvalva; this characterstate is
signum. The latter character states are interpreted interpretedasasynapomorphywith Seticosta. The
as symplesiomorphies because oftheirwidespread male of Punctapinella also possesses the charac-
distribution in the outgroups (i.e., Chlidanotinae, teristic euliine male foreleg hairpencil, as do males
Olethreutinae, and several tribes of the Tortrici- ofSeticosta(Brown,1989,1990).InAnopinellaand
nae). In most Euliini the apophyses are compara- Apolychrosis this structure is apparently second-
atinvdelyunsdhiofrfte,raenntditahteeddufcrtoumsbthuerscaoeripsugsr.eatlyreduced aarmiloynglostth.esTehefouhrypgoetnheersaiziesdprcelsaednistteidcirnelFaitgiuornesh1i.p
During recent studies on Neotropical members MATERIALS AND METHODS
of the polyphyletic genus Eulia Hubner, I discov-
ered a homogeneous group of species that share Taxonomic material for this study was obtained
theabovecharactersstatesbutdifferfromtheabove through the courtesy of J.F.G. Clarke, National
genera in several morphological characters and in Museum of Natural History, Washington, D.C.
(USNM); the late F. Fernandez-Yepez, Instituto de
1. Entomology Section, Natural History Museum of Zoologia Agricola, Universidad Central de Vene-
LosAngelesCounty,900 Exposition Boulevard,LosAn- zuela, Maracay (IZAV) [currently on loan to
geles,California 90007. USNM]; M. Lodi, Naturhistorisches Museum, Vi-
Contributionsin Science,Number 423, pp. 1-9
Natural History Museum of Los Angeles County, 1991
2CCOO o ©coz O nFaWtio=nfaonrdegweinnigt;alHicWstr=uchtiurnedswifnolgl;oDwsCH=ordaiksc(a1l9c8e4l)l;.
xo>oz«. c©OQ.. cOOQ. ~oto~o SYSTEMATICS
<Q. <cz Z3 © Punctapinella new genus
CL CO
Eulia Hiibner (in part); Meyrick, 1912:681; Mey-
rick,1931:151;Meyrick,1932:257;Clarke,1958:
120; Razowski, 1964:458, 461.
TYPE SPECIES. Eulia conchitis Meyrick, 1912.
DESCRIPTION. Head: Antennal setulae (cilia)
inmale1.6-1.8 x annularsegmentdiameter.Labial
palpusporrect,elongate,broadeneddistallybyscal-
ing; segment II expanded distally to ca 1.5 x its
basal diameter, slightly curved; segment III ca 0.4
as long as II, exposed. Maxillary palpus rudimen-
tary. Frons with overhanging crown tuft. Ocelli
small. Chaetosema present. Thorax: Male foreleg
with hairpencil from proximal end of femur, ex-
tending to base of coxa. Forewing: Venation as in
APFpuitogalutyricevher1o.ssyinsHa,yppAonomotophriepnsheiilzeleasd,sPuculpanpdcoitrsattpiiicnngerltelhlaae,tibaornnasdnhciShpeitnoagcmiosptanatg.- 0F.i6gurxeF2.WLelnenggtthh;2.w3i-d2t.h4oxfDwiCdtcha;0l.e2ngxthitosflDenCgtcha;
ptoerrnreacrtealsabfioalllopwasl:pi1;)LloosnsgoafnftoenrneawlincgilvieaininCumPa;lee;lovnaglavtaee, vCeuinAs2soerpiagriantaetibngeycoand0.c6e0ll;alRon4gtolecnogsttha,oRf5DtCo;tearl-l
narrow; ductus seminalis from corpus bursae. 2) Posses- men; CuPabsent; chordapresent; M-stem reduced
sionofaccessorybursa.3)Rowofspinelikesetaeoncosta to trace. Hindwing:Venationas in Figure2. Sc+R
M
ofvalva. and Rsseparate; Rs and Mi separate; and CuAj
3
connate; CuP represented by a trace; M-stem a
trace. Abdomen: Dorsal pitsabsent. Malegenitalia
enna (NHMW); and K.R. Tuck, British Museum (based on P. conchitis only): Uncus simple, un-
(Natural History), London (BMNH). modified, slightly curved. Socii moderately short,
Dissectionmethodologyfollowedthatpresented digitate,with finesetae;not fused tognathos. Gna-
in Powell (1964). Illustrations of genitalia were thos well developed with elongate, medial process
drawn using a microprojector. Forewing measure- from fusion ofapices oflateralarms. Subscaphium
ments were made with a standard grid mounted in and hami absent. Transtilla membranous mesally;
a dissecting microscope. Terminology forwingve- well developed laterally with cuplike subbasal in-
Figure2. Wingvenation ofPunctapinella theta.
2 Contributions in Science, Number 423 Brown: Description ofPunctapinella
vagination. Valva narrow, moderately elongate; suffix “ella” refers to diminuative in size. The gen-
costa with subbasal patch of large spinelike setae; der of the new genus is interpreted as feminine.
venter weakly excavated ca 0.4-0.7 from base to
apex;sacculussimplewithoutfreeprojection,con- KEY TO SPECIES OF PUNCTAPINELLA
fined to basal 0.5. Vinculum complete. Aedeagus BASED ON FEMALES
moderately long, straight; a single long slender
compound cornutus,attached basally. Femalegen- 1. FW silver-white with bronze-gray v-shaped
italia (based on all species): Papillae anales (ovi- transverse band (Fig. 7) conchitis
positor lobes) simple. Apophyses anteriores and T. FW nearlyunicolorous bronze-brownto choc-
posteriores long, slender. Sternum of abdominal olate brown with roundish white spots 2
segment VIII weakly sclerotized. Sterigma simple; 2. FW costa with white spot ca 0.65 from base,
lamella antevaginalis poorly developed; lamella between large subbasal spot and apical spot(s)
postvaginalis a simple sclerotized transverse band, . 4
broadenedmeso-ventrally,oftenwithnarrowscob- 2'. FWcostawithoutwhitespotca0.65 from base
inate region extending toward or into ostium; os- .3
tium displaced anterad of intersegmental mem- 3. HW chocolate brown, concolorous with FW
dburcatnuesbbeutrwsaeeenususatlelrynadiVfIfIereanntidatVeIdI.I.DuCcotrupsubsurasnade 3'. (HFiWg. 3l)ight gray-brow..n...,.....much lighterntihpahnasFtrWa
narrow,elongatetomoderatelybroad.Corpusbur- (Fig. 5) hraziliana
sae narrow, weakly pear-shaped, with little or no 4. Three white spots in medial-tornal area of FW
spiculae; signa lacking. Accessory bursa variously round and equal in size scleorductus
FW
developed: Short digitate diverticulum (e.g.,brazil- 4'. Three white spots in medial-tornal area of
iana)toelongatesacfromlongslenderductus(e.g., irregularly rounded and unequal in size 5
conchitis), arising from ductus bursae (usually lost 5. Largewhitespotnearbaseofcostanearlyround
in slide-mounted preparations). Ductus seminalis (Fig. 4) .... chionocarpa
fromcorpusbursae. Entireapparatuslongandnar- 5'. Large white spot near base of costa oblong or
row. Sexual dimorphism: Based on P. conchitis, subrectangular (Fig. 6) theta
forewing shape, size, and pattern does not differ
between sexes. Males can be distinguished by the
possessionofaforeleghairpencilandlongantennal 1. Punctapinella niphastra (Meyrick)
new combination
setulae (cilia).
Punctapinella occurs from Colombia and Ven- Figures 3, 8
ezuela,southeasttoRiodeJaneiro,Brazil.Theearly
Eulia niphastra Meyrick, 1931:151; Razowski,
stages are unknown. The three previously known
1964:461.
species were described in the polyphyletic genus
Euliaandwereknownonlyfromholotypefemales. MALE. Unknown.
Members of the genus are characterized by a FEMALE. FW length 6.9 mm (n = 1). Head:
unicolorousbronze-browntodarkbrownforewing Frons smooth scaled, white-ocherous below mid-
with distinct roundish white spots. Forewing pat- eye, roughened, gray-brown above; vertex gray-
tern in the closelyrelatedAnopinellaandSeticosta brown with white-ocherous tufts medially. Labial
nearlyalways features a dark triangularpatch bor- palpus white-ocherous, gray-brown laterally. An-
deringthe costaapproximately0.33-0.66 from the tennabrown.Thorax:Smoothscaled,brown.Fore-
base and extendingabout halfway to the dorsum. wing: Chocolate brown with faint lighter brown
In Punctapinella the gnathos are well developed transverse striae; small white spot at base along
and possess an elongate medial rodlike process dorsum;largewhitespotatcostaca0.20frombase;
formed by the distal fusion of the lateral arms. By smallwhitespotatposteriorapexofDC;twoslight-
contrast, in Seticosta the gnathos are reduced; in ly larger white spots nearly equidistant from the
Anopinella each of the arms bears a laterally flat- latter,one tornad,the otherapicad;apairofwhite
tened, usually finely dentate, semicircular process costal strigulae near apex; faint ill-defined white
distally. The single male of Punctapinella has a dots in subterminal region; dorsum with several
moderately long straight aedeagus bearing a single diffuse short white transverse dashes. Fringe fer-
largecompoundcornutus.MalesofAnopinellahave ruginous-gray. Hindwing: Chocolate brown. Fringe
a comparativelyshort stoutaedeagus bearingasin- slightly lighter brown. Genitalia: As in Figure 8 (n
gle slender spinelike cornutus; males of Seticosta = 1;drawnfromRazowskislideno.4339).Lamella
almostalways lack cornuti and have a large dorso- postvaginalis a simple transverse curved band,
anterior process above the opening of the ductus broadenedmeso-ventrally.Ductusbursaelong,fairly
ejaculatoris. PutativeautapomorphiesforPunctap- broad. Corpus bursae narrow, elongate. Origin of
inellaincludetheuniqueforewingpattern,theelon- neither accessory bursa nor ductus seminalis de-
gate medial fusion ofthe arms ofthegnathos, and tectable.
the unusual compound cornutus. TYPE MATERIAL. Holotype female: BRAZIL:
ETYMOLOGY.TheGreek“puncta”meansspot State of Espiritu Santo, Guanda, December 1920,
or spotted; “pina” refers to feather or wing; the Fritz Hoffman (NHMW).
Contributions in Science,Number423 Brown: Description ofPunctapinella 3
ETYMOLOGY.TheGreekprefix“nipha”means no. 69338). Lamella postvaginalis a simple trans-
snow and “astro” or “astrum” means star. Appar- verse band with a scobinate, broadly v-shaped re-
entlythename“niphastra”referstotheroundwhite gion meso-ventrally; variable rectangular patch of
2.
spots of the forewing. scobination extending to ostium; sclerotized ridge
representing ventral lip of ostium. Ductus bursae
moderatewithshortsclerotizedantrum. Accessory
Punctapinella chionocarpa (Meyrick) bursaa short digitate diverticulum from nearjunc-
new combination tion of ductus bursae and ostium. Corpus bursae
Figures 4, 9 oblong. Ductus seminalisfromcaudal third ofcor-
pus.
Eulia chionocarpa Meyrick, 1932:257; Razowski, TYPE MATERIAL. Holotype female: BRAZIL:
1964:458.
State of Santa Catarina, Santa Catarina, XI-20-36
MALE. Unknown. [20 November 1936], Fritz HofFman (USNM).
FEMALE. FW length 9.0 mm (n = 1). Head: Fourfemaleparatypesasfollows: BRAZIL:State
Frons and vertex white. Labial palpus lacking. An- of Parana, Castro, 39$, 1898 (E. Jones, BMNH).
No
tenna light tan. Thorax: White. Forewing: Bronze- localitydata,ParaviciniCollection], 1$,B[ritish]
brown with diffuse darker reticulation; white spot M[useum] [accession no.] 1937-383 (BMNH).
ADDITIONAL MATERIAL EXAMINED.
atbasealongdorsum;largeirregularlyroundedspot
atcostaca0.25from base;foursmallerwhitespots BRAZIL: State of Rio deJaneiro, km 17, 18 km S
indistal0.5ofwing:oneatcostaca0.65frombase, Teresop4ol.is, 1180m, 1$,IV-18/19-77[18-19April
one directly below latter at posterior apex of DC, 1977], C.M. and O.S. Flint (USNM).
oneslightlylarger,betweenanddistadoflattertwo, REMARKS.ThespecimenfromtheStateofRio
one slightly oblong, at tornus; two pairs of white deJaneiro is smaller in forewing length and has a
costal strigulae near apex, forming subconfluent broader ductus bursae that is not clearly differen-
white spot; subtermen with indistinct white dots. tiated from the corpus bursae. It is possible that
Fringe light bronze-brown. Hindwing: Light gray this specimen is not conspecificwith P. braziliana.
with faint darkergray marbling. Fringegray-white. Consequently, the specimen is not designated a
zGeonwistkailisal3:i.dAesnoi.n4F3i3g2u)r.eL9am(enll=ap1;osdtvraagwinnalfirsoamsRiam-- parEatTyYpeM.OLOGY.Thename“braziliana”refersto
ple lightly sclerotized band; ostium indicated by the country of the type locality of this species.
narrowoutlineofsclerotization.Ductusbursaelong,
narrow. Corpus bursae irregularly rounded. Origin Punctapinella scleroductus
ofaccessory bursa and ductus seminails uncertain. new species
TYPEMATERIAL.Holotypefemale:COLOM- Figure 11
yBraeIsaA,r:]G,[uEDidnedpeiannr,t(mN8eH5n0Mt0Wo)ffe.Tetol[i2m7a50(?)m]],CeNntorvaelmCboerdril[lneo- FMEAMLAEL.EU.nkFnWownle.ngth 9.2 mm (n = 1). Head:
ETYMOLOGY. The Greek “chion” refers to Frons and vertex white. Labial palpus white, gray-
snow and “carpo” or “carpus” means fruit. Ap- brown laterally. Antenna light brown. Thorax:
parentlythename“chionocarpa”referstotheround White-ocherous, darker anterad. Forewing: Gray-
white forewing markings. brownwithfaintsparsered-brownoverscaling;small
whitespotatbasealongdorsum;diagonalelliptical
white spot from costa ca 0.2 from base; foursmall
Punctapinella braziliana round spotsin distal 0.5: one at costa ca0.60 from
new species base, one directly below latternearposteriorapex
of DC, two nearly equidistant from the latter, one
Figures 5, 10 tornad, the otherapicad; apexwith irregularwhite
MALE. Unknown. spotincorporatingcostal strigulae;subtermenwith
FEMALE. FW length 6.8-9.0 mm (x = 7.5; n = faint indistinct white dots. Fringe gray-brown.
5). Head: Frons and vertex white. Labial palpus Hindwing: Light gray-brown. Fringe concolorous
white,gray-brownlaterally.Antennabronze-brown. withwing. Genitalia:Asin Figure 11 (n = 1; drawn
Thorax: Yellow-ocherous to silver-white. Fore- fromUSNMslideno.69336).Lamellapostvaginalis
wing: Bronze-brown;white spotat basealongdor- asclerotizedbandwithdensepatchofscobination;
sum; large elongatediagonalwhitespot from costa edge of abdominal segment VII immediately pos-
ca 0.2 from base; white spot near posterior apex terad of ostium with broad patch of scobination
of DC; distal 0.2 of wing with row of three small and narrow sclerotized ridge; ostium round with
white spots nearly parallel to termen starting at sclerotizedperimeter.Ductusbursaebroad,strong-
apex,apicalspotformed bycoalescedcostalstrigu- ly sclerotized throughout. Corpus bursae narrow
lae; dorsum with a few scattered tiny white trans- with fine ridges ventrally. Accessory bursa from
versedashes.Fringebronze-brown.Hindwing:Light long narrow ductusarisingnearjunction ofcorpus
gray-brown. Fringe gray to dingy white. Genitalia: and ductus bursae. Ductus seminalis from caudal
As in Figure 10 (n = 3; drawn from USNM slide portion of corpus bursae.
4 Contributions in Science, Number 423 Brown: Description ofPunctapinella
Figures 3-7. Adults ofPunctapinella. 3. Holotype of P. niphastra. 4. Holotype ofP. chionocarpa. 5. Holotype of
P. braziliana. 6. Holotype ofP. theta. 7. Female ofP. conchitis. Allphotographs courtesyofVictorKrantz (USNM).
TYPEMATERIAL. Holotypefemale: BRAZIL: P. theta. Itcan bedistinguishedfromthelattertwo
State of Rio deJaneiro, Rio deJaneiro, 7/3/29 [3 by the shape and size of the three white spots of
July 1929],J.F. Zikan (USNM). thedistalhalfoftheforewing;i.e.,roundandequal
REMARKS. The adult of P. scleroductus is not in size in P. scleroductus (see key to species).
illustrated, but it is similar to P. chionocarpa and ETYMOLOGY.Thename“scleroductus”refers
Contributionsin Science, Number 423 Brown: Description ofPunctapinella 5
8
Figures 8, 9. Femalegenitalia of Punctapinella. 8.
to the strongly sclerotized ductus bursae of the white dots. Fringe red-brown. Hindwing: White-
female genitalia. ocherouswithsparsegray-brownoverscaling.Fringe
white-ocherous. Genitalia: As in Figure 12 (n = 2;
drawnfromUSNMslideno. 69334). Lamellapost-
5. Punctapinella theta new species vaginalis a sclerotized transverse band slightly
broadenedmeso-ventrad,withslenderdigitate band
Figures 2, 6, 12 ofsclerotizationextendinganteradintoostium;pe-
MALE. Unknown. rimeter of ostium unsclerotized. Ductus bursae
FEMALE. FW length 8.0-8.2 mm (x = 8.1; n = moderatelybroad.Corpusbursaeapear-shapedsac.
2). Head: Frons and vertex white. Labial palpus Ductusseminalisfromcaudal0.30ofcorpus.Weak,
white,bronze-brownlaterally.Antennalightbrown. broadductusofaccessorybursafromductusbursae
Thorax: White. Forewing: Bronze-brown; white near ostium.
spotatbasealongdorsum;largeroblongwhitespot TYPE MATERIAL. Holotype female: VENE-
from costa ca 0.2 from base; four roundish white ZUELA:StateofAragua, RanchoGrande, 1100m,
spots in distal 0.5 of wing: one on costa ca 0.65 VIII-15/21-67 [15-21 August 1967], R. W. Poole
from base; one at posterior apex of DC; one at (USNM).
dorsum near tornus, the other directly above the One female paratype same locality as holotype,
latter; apex with irregularwhite area comprised of V-15-67 [15 May 1967],J. Salcedo and C. Andara
coalesced costal strigulae; subtermen with faint (IZAV).
6 Contributions in Science,Number 423 Brown: Description of Punctapinella
11
Figures 10, 11. Femalegenitalia ofPunctapinella. 10. P. braziliana. 11. P. scleroductus.
ETYMOLOGY. The specific epithet “theta” re- laterally. Antenna brown. Thorax: Silver-white.
ferstothet-shapedsclerotizedregionofthelamella Forewing: Olive green-bronze, with two broad
postvaginalis. transverse white bands: one from costa ca 0.2-0.3
from base, extending to dorsum, posteriorly con-
tiguous with white basal spot; second band occu-
6. Punctapinella conchitis (Meyrick)
pying distal 0.3 of wing, with faint, light bronze-
new combination olive band at termen; subtermen with poorly
Figures 7, 13, 14 definedbronzestreakparalleltotermen;whitespot
atcostaca0.65 from base representingpairofcon-
Eulia conchitis Meyrick, 1912:681; Clarke, 1958:
fluentwhitecostalstrigulaeandasecondconfluent
120
. pairca 0.9 from base; a small white spot nearpos-
MALE. FW length 8.0 mm (n = 1). Head: Frons teriorapex ofDC. Fringe tan and cream-ocherous.
and vertexwhite. Labial palpus white, light brown Hindwing:Whitewithgray-brown mottling. Fringe
Contributions in Science,Number 423 Brown: Description ofPunctapinella 7
: :
12
Figures12, 13. FemalegenitaliaofPunctapinella. 12. P. theta. 13. P. conchitis. ds= ductusseminalis,ab=accessory
bursa, p = pouch.
concolorous with wing. Genitalia As in Figure 14 postvaginalis a broad scobinate band; lamella an-
(n = 1; drawn from JWB slide no. 276). Uncus tevaginaliswitha stronglysclerotized ovalpatch at
slender, rodlike, curved. Socii simple, short, digi- ostium. Ductus bursae moderately short. Corpus
tate, setose. Gnathos with strong medial process bursae simple, elongate. Accessory bursa a mem-
from apical fusion of lateral arms. Transtilla weak branous attenuate sac from long narrow weakly
mesally; basal region large, appearing as a strongly sclerotizedductusarisingoncaudalportionofcor-
sclerotized basal extension ofvalva;a deep cuplike pus bursae; unusual sclerotized distally attenuate
invagination subbasally. Valva narrow, moderately pouch attached to bursa copulatrix at junction of
elongate, attenuate distally; costa with subbasal corpus and ductus bursae. Ductus seminalis from
patch of 10-15 large spinelike setae; venter with anterior third of corpus.
rounded excavation ca 0.4-0.7 from base to apex; TYPEMATERIAL.Holotypefemale:COLOM-
sacculus confined to basal 0.5 without free projec- BIA: Department of Norte de Santander, San An-
tion. Aedeagus moderately long, slender, with a tonio. 5800 feet [1800 m], November [no year]
single large compound cornutus. (BMNH).
FEMALE. FW length 9.0 mm (n = 2). As de- ADDITIONALMATERIALEXAMINED.CO-
scribed for male. Genitalia As in Figure 13 (n = LOMBIA: Department ofNorte de Santander, San
2; drawn from USNM slide no. 68850). Lamella Antonio, 1$, “11-07” [November 1907] (USNM);
8 Contributions in Science, Number 423 Brown: Description ofPunctapinella
,
Figure 14. Malegenitalia ofPunctapinella concbitis.
DepartmentofMagdalena,Sierra Nevadade Santa Tortricinae (Lepidoptera: Tortricidae). Entomolog-
Marta,HotelSolito,2300m, Id,VII-24-73[24July icalNews 101:109-116.
1973], 7-8 PM (Oxford Expedition, BMNH). Clarke,J.F.G. 1958. Catalogue oftheTypeSpecimens
REMARKS.TheforewingpatternofP.concbitis ofMicrolepidopteraintheBritishMuseum(Natural
probablyrepresentstheplesiomorphicstateforthe History)DescribedbyEdwardMeyrick volume3.
Tortricidae, Olethreutidae, Noctuidae. Trustees of
genus becauseit ismostsimilarto thatofSeticosta the British Museum, London. 600pp.
andAnopinella. OtherspeciesofPunctapinellaex- Horak, M. 1984. Assessment of taxonomically signifi-
hibit an increase in melanic scaling, reducing the cant structures in Tortricinae (Lep., Tortricidae).
presumedwhitegroundcolortosmallroundspots. Bulletin de la Societe Entomologique Suisse 57:3-
ETYMOLOGY.TheGreekprefix“conch”means 64.
shell;thesuffix“itis”meansinflammation.Thename Meyrick,E. 1912. DescriptionsofSouthAmericanMi-
may refer to the bronze-brown, somewhat trian- cro-Lepidoptera.TransactionsoftheEntomological
gular-shaped (shell-like?) marking of the forewing. ———SocietyofLondon 1911:673-718.
1931. ExoticMicrolepidoptera4:33-192.Marl-
.
ACKNOWLEDGMENTS ———borough, Connecticut.
1932. Exotic Microlepidoptera 4:193-353.
I thank J.A. Powell, University of California, Berkeley, Ma.rlborough, Connecticut.
C.L. Hogue, Natural History Museum of Los Angeles Powell,].A. 1964. Biologicalandtaxonomicstudieson
County, and two anonymous reviewers for numerous tortricine moths, with reference to the species in
helpfulcommentsonthemanuscript.Thisworkwascom- California. UniversityofCaliforniaPublicationsin
pleted with the support of a Smithsonian Postdoctoral —Entomology32:1-317.
Fellowship. . 1986. SynopsisoftheclassificationoftheNeo-
tropicalTortricinae,withdescriptionsofnewgenera
LITERATURE CITED and species (Lepidoptera: Tortricidae). Pan-Pacific
Entomologist62:372-398.
Biown,J.W. 1989. GenericreassignmentsforNeotrop- Razowski,J. 1964. MeyricktypesofTortricoidea(Lep-
icaltortricidmoths(Lepidoptera:Tortricidae).Jour- idoptera) in the Vienna Museum. Annales Zoolo-
———naloftheLepidopterists3Society43:313-322. gicae, Polska AkademieNauk 22:451-481.
. 1990. Taxonomicdistributionandphylogenetic
significance of the male foreleg hairpencil in the Received 7February 1990; accepted 1 May 1990.
Contributionsin Science,Number423 Brown: Description ofPunctapinella 9