Proceedings of the Academy Indiana of Science VOLUME NUMBER 114 2 PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE The PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE is a journal dedicated to promoting scientific research and the diffusion ofscientific information, to encouraging communi- cation and cooperation among scientists, and to improving education in the sciences. EDITOR: James \V. Berry. Department of Biological Sciences, Butler University, Indianapolis, Indiana 46208. Telephone: (317)-940-9344; FAX: (317)-940-9519; e-mail:[email protected] EDITORIAL BOARD: Hans O. Anderson (Indiana University, Bloomington); Robert F. Dale (Purdue University, West Lafayette); Rebecca Dolan (Butler University); Kara W. Eberly (St. \lar\ "s College); Uwe J. Hansen (Indiana State University); Daryl R. Karns (Hanover College); Gene Kritsky (College ofMt. St. Joseph); StephenA. Perrill (Butler University); Paul Rothrock (Taylor University); Thomas P. Simon (U.S. Fish & Wildlife Service, Bloomington); Michael R. Tansey (Indiana University, Bloomington); Robert D. Waltz (Indiana Department ofNatural Resources, Indianapolis); J. Dan Webster (HanoverCollege); HarmonWeeks (Purdue University, West Lafayette); John O.Whitaker,Jr. (Indiana State University) THE INDIANAACADEMY OF SCIENCE PRESIDENT: Uwe J. Hansen (Indiana State University) PRESIDENT-ELECT: Clare L. Chatot (Ball State University) TREASURER: Edward L. Frazier (Indianapolis) SECRETARY: Nils I. Johansen (University ofSouthern Indiana) EXECUTIVEDIRECTOR: Nelson R. Shaffer(Indiana Geological Survey) ARCHIVIST: Holly Oster (Indiana State Library) EXCHANGE ITEMS: Items sent in exchange for the Proceedings and correspondence concerning exchangearrangementsshouldbesenttotheIndianaAcademyofScience,JohnS.WrightMemorial Library, 140 North SenateAvenue, Indianapolis, Indiana46204. Questions regardingbackissues or undelivered issues shouldbe addressedto Holly [email protected] Telephone (317)- . 232-3686. Coverphoto: The photo shows a wolfspider (Araneae: Lycosidae) from Grand Sable Dunes on the shore of Lake Superior, Pictured Rocks National Lakeshore, Alger County, Michigan. Spiders in this family are more commonly observed on the ground surface rather than in vegetation. Without examining the spider under the microscope, it is not possible to do more than guess about the spe- cific identityofthisanimal.This illustratesonereasonwhytheacquisitionofdataonthisubiquitous group ofanimals has been a slow process. See related paper on page 111. Photo by Gary Fewless (University ofWisconsin-Green Bay). Visit the IndianaAcademy ofScience website at: www.indianaacademyofscience.org Publication date: 31 January 2006 Thispapermeetsthe requirementofANSI/NISOZ39.48-1992(PermanenceofPaper). ISSN 0073-6767 2005. Proceedings of the Indiana Academy of Science 1 14(2^:83-104 PALAEOECOLOGICAL INTERPRETATION OF POLLEN, MACROFOSSILS, POLYGONAL FISSURES, AND TAPHONOMY OF THE SHAFER MASTODONT LOCALITY, WARREN COUNTY, INDIANA Anthony L. Swinehart: Department of Biology, Hillsdale College, Hillsdale, Michigan 49242 USA Ronald L. Richards: Indiana State Museum, 650 West Washington Street, Indianapolis, Indiana 46204 USA William Petty Rivers1: University of Tennessee, Center for Quaternary Studies, Knoxville, Tennessee 37996 USA Robert D. Hall and Allyson K. Anderson: Department of Geology, Indiana — University Purdue University at Indianapolis, Indianapolis, Indiana 46202 USA ABSTRACT. Discovery of thejaw of an American mastodont (Mammut americanum) and an unusual sedimentary profile in a cornfield in Warren County, Indiana, prompted a multidisciplinary study of the palaeoenvironment of the site. Wood taken from near the base of the deposit was dated at 15.540 ybp. m Stratigraphic and textural analyses of the 2.3 sedimentary profile reveal a series of inundation and desiccation events marked by polygonal fissures. Analysis of pollen from the sediment profile indicates that a boreal flora predominated during much of the time period represented by the profile. Pollen cor- relation indicates that the sedimentary record was truncated by unconformities around 10,000 ybp. Mac- rofossil analysis indicates a local environment that began as a forest dominated by white spruce and tamarack. Later inundation of the forest was indicated by the appearance of fish (Perca flavescens), meadow voles (Microtuspennsylvanicus), and submergent aquatic macrophytes (Myriopyllum exalbescens. Potamogeton pusillus, Ceratophyllum demersum, and Najasflexilis). The aquatic environment was inter- ruptedby periods ofexposure anddesiccation as indicatedbythedisappearanceofidentifiablemacrofossils and by the stratigraphy. The Shaferfossil assemblage is compared with otherlocalities, andthe taphonomy and palaeoenvironment of the mastodont are discussed. Keywords: Mastodon, polygonal fissures, paleoecology, macrofossils, pollen, Late-Pleistocene. Holo- cene, Quaternary biota On 5 July 1992, while digging a trench for mains of the mastodont and other associated drainage tile in his Warren County cornfield, fossil material. Little additional mastodont or Larry Shafer encountered teeth and jawbone other vertebrate material was recovered. How- fragments of the American mastodont {Mam- ever, the complex stratigraphy encountered at mut americanum). Upon discovery of the re- the site, including well-preserved fissure fill- mains, digging was suspended; and Shafer ings and a lower stratum rich in plant mac- contacted Purdue University archaeologists, rofossils, provided an opportunity to charac- who referred him to the Indiana State Muse- terize the paleoenvironment of the Shafer urn. A thorough excavation of the site (re- Mastodont. The discovery contributes to in- ferred to as the Shafer Mastodont Locality) laeoecological understanding of Late-glacial was then conducted by staff and volunteers of and post.glacia] environments in Endiana. The ttheembIenrdiiannaanSatattteemMprtustoeuremcofvreormad1di7t-i2o5nalSerpve-- oub,mj•'eenctt..i.tv.,hees"*foorcssti.ulh,eubip-ort.easeont,t-._t,pheapSCe1hrata,-erre*M.tao.st11xo)dd,,oonct- 'Formerly published as William H. Petty; Current Locality. 2) describe and discuss the taphon- address: Department ofBiology; St. LawrenceUni- omy of the Shafer Mastodont. 3) determine versity; Canton, New York 13617 the origin of polygonal fissure fillings at the 83 PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE 84 site, and 4) reconstruct the palaeoenvironment of the area using pollen, macrofossils, verte- brate remains, and stratigraphy. STUDY AREA The study area is in Section 7, T23N, R6W, Chatterton Quadrangle, Warren County, Indi- ana (40°27'21"N; 87°07'52"W) at an elevation of 210 m (—690 ft. asl). It occurs within shal- low, marginal wetlands associated with Otter- bein Bog, a glacial kettle depression with a maximum depth of approximately 13 m (Richards 1938). The site is situated on a por- tion of the Crawfordsville End Moraine, part of the Cartersburg Till Member of the Trafal- gar Formation, created approximately 16,000 ybp (years before present) during Wisconsin- age glaciation (Wayne & Zumberge 1965; Fullerton 1986). The peat in nearby Otterbein Bog originated from grass and sedge remains, and Richards (1938) reports that much of the recent vegetation ofthe wetland is Phragmites and Calamagrostis. The wetland surrounding Figure 1.—Map of Otterbein Bog watershed Otterbein Bog is referred to on the United (Sections 5, 6, 7, 8, 17 & 18, T23N, R6W, Warren States Geological Survey (USGS) topographic County, Indiana), showing the location ofthe study map as "Cranberry Marsh," suggesting that area. Contour interval is 10 ft.(3 m) a.s.l. the wetland may have harbored cranberry (Vaccinium cf. V. macrocarpon), a plant usu- ally restricted to peatlands with a boreal cli- encountered any additional bone. Gasoline- mate or similar microclimate. The peatland is powered pumps removed water that seeped best described as a fen or sedge-meadow be- into the excavation site, and also pumped cause it is not dominated by Sphagnum moss- clean water from adjacent Holder Ditch to es and is extremely mineral-rich (as indicated wash excavated sediments for the recovery of by the predominant flora). The soils immedi- microfauna and plant macrofossils. A site da- ately around the site are mainly poorly- tum and baseline were established for leveling drained silty loams and silty clay loams ofthe and mapping. Profiles and floor "bench" ar- Brenton, Drummer, and Williamstown-Rains- eas were created to facilitate mapping and ville series (Barnes 1990). The Shafer Mas- photography of the complex stratigraphy. todont Locality is situated on the southwest Bulk material from the trenches was variably mm mm edge of Cranberry Marsh that formed within washed through 1.2 (0.05 in) or 6.3 a kettle depression (Fig. 1). (0.25 in) mesh screens. When bulk screening METHODS from both trenches failed to yield any mas- — todont or other discernible biotic remains, the Field procedures. Only a few fragments excavation strategy changed toward develop- of mastodont bone at the Shafer Locality were ing stratigraphic profiles, down to till if pos- found in situ; the remainder was disturbed by sible, to understand the geologic context of excavating. Disturbed soils were removed by the mastodont jaw. In doing this, an organic shovel and trowel down to undisturbed soil. silt, rich in conifer cones, was encountered. Thereafter, two intersecting trenches were ex- The soils from this "cone zone" were exten- mm cavated by shovel and backhoe near the lo- sively sampled and washed through 1.2 cation of the original bone fragments in an mesh screen. Bulk soil samples were taken attempt to locate additional mastodont re- from each of the distinct strata in the profile mains. Neither the 9.5 m long east-west trench for textural analysis and recovery of macro- m nor the 10.2 long north-south cross trench fossils. Additionally, small plastic canisters SWINEHART ET AL.—SHAFER MASTODONT LOCALITY 85 — were driven into a freshly exposed profile at Macrofossil analysis. In addition to anal- 10 cm intervals from the surface ofthe profile ysis of macrofossils obtained from bulk mm to the underlying glacial diamicton. These screening in the field using 1.2 (0.05 in) mm were capped and taken to the laboratory for or 6.3 (0.25 in) mesh sieves, more careful pollen analysis. analysis of the sediment was conducted in the During the excavation of a drainage pit at laboratory to locate smaller or more delicate the south end of the north-south trench, the material. Subsamples of 300 cm3 were care- backhoe encountered a deeply-buried conifer fully broken by hand and inspected for leaf log. This prompted eight exploratory pits and impressions. To dissociate the soil, samples trenches several meters beyond the site perim- were then soaked in a 50 g per 1 solution of eter. None produced any additional vertebrate sodium phosphate for three days and rinsed material. A final widening and deepening of through a 0.4 mm sieve. Macrofossils were all the trenches and pits around the site still identified and counted with the aid of a dis- failed to yield bone of any—kind. secting microscope. Excess bulk material that Stratigraphic analysis. Analysis of soil was not used in the quantitative subsampling texture was accomplished using the Bouyou- was placed in a white enamel pan forrecovery cos Procedure (Bouyoucos 1936). Percent or- of large or infrequent macrofossils. Voucher ganic carbon in the soils was determined using specimens of macrofossils were deposited at the loss-on-ignition—method (Storer 1984). INSM. — Pollen analysis. Extraction of pollen and Radiocarbon dating. A single radiocar- spores from the sediment samples was accom- bon date was obtained from a piece of spruce plished using standard methods modified from wood located at a depth of 2.1 m ("cone Faegri & Iverson (1975). A 1 cm3 sample zone"), approximately 1.8 m below the level from each level was used in processing. Pa- from which the mastodont molars and jaw lynomorph identification was based primarily fragment were found and approximately 20 on the key by MacAndrews et al. (1973), cm above the glacial till. The wood sample, along with the aid ofthe pollen reference col- sent to Beta Analytic, Inc., was processed for lection at the Center for Quaternary Studies at a standard radiocarbon age determination the University of Tennessee, Knoxville. Tax- (Beta-62640). An attempt was made to date onomy follows Gleason & Cronquist (1963). the mastodont, but mandible fragments failed Identification ofblack spruce (Picea mariana) to produce recognizable collagen for dating and white spruce (Picea glauca) pollen was (Beta Analytic, pers. commun.). based on the morphometries developed by & RESULTS & DISCUSSION Birks Peglar (1980). Twenty spruce grains — were measured for each level and assigned to Stratigraphy. The Shafer Mastodont Lo- either black spruce, white spruce, or undiffer- cality, like most of the other lakes and bogs entiated spruce, following Hansen & Engstron that have yielded mastodonts in Indiana, is a (1985). These values were then used to assign shallow deposit of aquatic sediments that was the remaining spruce pollen to one of those truncated early in the Holocene. In this case. categories. For each level a minimum of 300 the profile represents a time period ofapprox- terrestrial pollen grains was counted. Due to imately 5000 years, beginning just after de- the existence of only a single radiocarbon glaciation nearly 16,000 ybp. Four major di- date, it was not possible to calculate pollen visions of sediments were identified in Profile influx rates for the profile. 4 and designated as A through D. oldest to Interpretation of the pollen diagram along youngest (Fig. 2). Division A has also been with assignment of chronology was augment- designated the "cone zone" (as well as Unit ed by comparison with several studies, cited XI in Fig. 2). This unit consists of wood and herein, which place the Shafer Mastodont Lo- plant remains in a matrix of silt loam. A piece cality in a regional context. The pollen tally of wood from this zone yielded a radiocarbon data are on file at the Center for Quaternary age of 15,540 ± 70 !4C ybp (Beta-62640). Studies, University of Tennessee, Knoxville, which provides a minimum age for deglacia- Tennessee 37996. Duplicate slides for each tion of the site and a maximum age for all level are held by the Indiana State Museum other sediments above. Division B consists of (INSM). interbedded silt and clav that is grav at the PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE 86 m only about 2.3 deep (Fig. 2), yet changes in the paleoenvironment are indicated by both the sediments themselves and the plant mac- rofossils and pollen they contain. The area ap- pears to have been in a relatively shallow fringe zone, or shelf, around the margin of a much deeper kettle at the south end of the Shafer site. Back-hoeing well below the cone zone exposed clay and logs. Based on the to- pography of the area, it is likely that the Sha- fer Site was on a shelf, and erosion ofthe hill- slopes (including sand and allochthonous organic matter such as wood, cones, and bones) may have transferred debris to the shelf. Waves and currents probably moved some of the debris to the deeper kettle. As the landscape became less of a direct source of sediment for the kettle, sand per- — Figure 2. Diagram of Profile 4, showing sedi- centages in the sediment declined from more ment divisions (capital letters) and units (Roman than 10% in the cone zone (Unit XI) to only numerals). Dots represent the location of pollen a few percent in the younger sediments (ex- samples taken from the profile at 10 cm intervals cept for Unit VI). The size ofthe organic mat- (see Table 1 for sedimentological details). ter also declined, and the finer sizes charac- teristic of the younger sediments were more disseminated throughout the matrix rather base (Unit X) then changes upward to blue- than being concentrated as in the cone zone. green (Unit IX), dark brown (Unit VIII), and The textures of Sediment Division B are silt brown (Unit VII) overlain by pure sand (Unit loam, silty clay loam, and silty clay, textures VI). This sand also in-fills fissures (Figs. 3, 5) common among glacio-lacustrine sediments. that may extend downward into Unit X. Be- Mean organic carbon content was relatively m sides a thickness of up to 1 of this sand as low at 15%. fissure in-fillings, there is another 5-25 cm as Delivery of organic sediments ceased tem- a blanket over younger units (Fig. 3). The unit porarily during deposition of the sand of Unit both blankets the older deposits and fills the VI. The extremely high percentage of sand in fissures within them. Sediments in Division C this unit (96%) and the fine to very fine sand are similar to those below it and consist of size suggest eolian deposition during an epi- brown silt (Unit V) that was buried, and fis- sode of aridity. A backhoe pit dug north of sures that were in-filled with brown silt loam the original excavation revealed a meter of (Unit IV) (Fig. 4). Unit IV of Division C is massive sand and may be part of a sand dune. the source of the mastodont mandible. Divi- However, it is not known whether this eolian sion D consists of a cap of brown loam (Unit sand is the same age as Unit VI. III), gray loam (Unit II), and black humus Organic deposition then resumed during the (Unit I). The oldest sediment reached in the formation of the deposits of Sediment Divi- excavation was late Wisconsinan till of the sion C. These younger sediments have about Wedron (?) Formation. This deposit of un- the same grain size, but have considerably known thickness presumably underlines the more organic matter than those of Sediment entire basin of Otterbein Bog. It was not de- Division B (Fig. 2, Table 1). Resumed depo- scribed at the Shafer Locality. Two samples sition in a quiet-water lacustrine or wetland were taken from below the cone zone, appar- environment seems likely. The youngest sed- ently from the matrix of till (Fig. 2), for lab- iments of Sediment Division D are probably oratory analyses (Table 1, Fig. 2). The texture the result oferosion in the drainage basin after of these samples is silt loam. Loss-on-ignition European settlement. Sand percentages in- averages 3.5%. crease, but organic content is very high with The sediments of the Shafer Locality are a loss-on-ignition value of 50% in Unit III. SWINEHART ET AL.—SHAFER MASTODONT LOCALITY *7 wan v* . — Figures 3-5. Photographs of fissures encountered in sediment divisions B and C. 3. Profile; 4. Cros section of sediment division C; 5. Cross section of sediment division B. — Origin of the fissure in-fillings. Three origin of polygonal fissure-fillings used in the main hypotheses for the origin of the fissure present study include Lachenbruch (1962). in-filled polygons at the Shafer Locality were Bertouille (1974), Nissen (1985). Mears considered (periglacial, desiccation, and sand- (1987), Nissen & Mears (1990). and Johnson blown mechanisms). Any reasonable hypoth- (1990). The periglacial hypothesis requires eses must include an explanation of two temperatures cold enough to maintain per- events: 1) what opened the vertically oriented mafrost. As the temperature drops, the per- fissures, and 2) how and when they were mafrost-laden soil will crack. Meltwater will filled. Moreover, there is the possibility that enter the crack and freeze, increasing the vol- the upper and lower sets of fissure in-fillings ume by 9%. Cracking may occur again and were formed by different mechanisms. again, gradually building up an ice wedge. Criteria from several sources by which the When the climate warms, the ice wedge melts. periglacial, desiccation, and sand-blow mech- and sediment begins to till the fissure. Most anisms may be compared, both to each other of the sediment is derived from the walls of and to the observations and data from the po- the crack. Some small amount ofmaterial may lygonal in-fillings at the Shafer Locality, are be washed or blown in. The most critical con- presented in Fig. 6. dition for this hypothesis to be valid is tem- Periglacial hypothesis: The primary peratures cold enough to first form permafrost. sources of information about the periglacial then contract and crack it. The time of the PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE 88 > T3 o o3 c toai3 6C0 0coo3 Z3Cofl >—c"/2 > <cSU3 o—Oc >>< o0s3 oC£u3 '?U50-3C>'£S<'3u <D <U 03 E23OUC<u/u5f'c•i3Cy. O&1DB-lhf5cf»3t* 5D0c33 ca£ur«3. "O£0a3. T^OiDh-SOXr (N H M3 vOrniOOr-CNCNinvO-^J-r^j Z Z — ITi f~, U c -J — J— J— J—. J—i — — U U a U a s a y y s j j y j C/5oOOOt/5oOOOGOOO 5^^r ZZ(N(Nv-(ONNOm^mt(ONN'ctnimr^ir^<|)-Mmin(Nna\^M(—NimnM—OOC)(N'-m-<mir)r^nih^^ihr^i^O cs < < in — r-ooaN — o^, 'oo\ooNOts '-'ronosr^r^oooom — i- o\ t < < SO Hh(s\OfimH-(H^\oa\>oin Z Z on >n 3 o Q© O >n 0o0 >0—n0 ion in + <-h <N <N m a ex 3i m o o-tii- oO o<N irn-I ooo1i oionI oini <N Q 2 -2 73 ^ ^ ^ .^ rSs Z D > > > a x X S ii < 7g3 z a ri M "O u c o £ | i. >, T5 1ND (J £ I H '5 03 u 0U0 Q O Q U U S CQ CQ S OQ CQ CQ SWINEHART ET AL.—SHAFER MASTODONT LOCALITY 89 Observationor Peri-glacial SeismicSand- Desiccation Shafer ShaferMastodont Criterion hypothesis blowHypothesis andIn-filling Mastodont Locality, UpperSet Hypothesis Locality,Lower Set 1)In-fillsoccuras Yes,activeice Usuallydike-like, Crackshould Almostallin-fills Someinfillstaperwith wedges,tapering wedgestaperwith with propagate arewedges depth;manydonot withdepth depth mushroomingif downward,thus taperingwith surfaceisreached. bewideratthe depth Usuallywiden surface withdepth 2)Map-view Yes,asshownby Notlikely,unless Yes,widely Well-defined Well-definedpolygons polygons numerousactive injectionoccurred known polygonspresent present areas alongolder mudcrackingis fissures polygonal 3)Dimensionsof Diametersof3to Nopolygons. Verylittledata Diametersof Diametersofabout75cm. polygons: a.Diamter 20m,butmostly Depthsofseveral available. about30cm, depthofabout90cm. b.Depth <1 m. Depthsof meters. Thickness depthofaboutI thicknessatlandsurface c.Thicknessat 1 to3m. of< 1 cmto>60 m,thicknessat about30cm junctionwithland Thicknessatthe cm. landsurface surface surfacemay about15cm. averageabout50 cm 4)Timingofin-filling Ifburied,ice Sedimentinjected Withfissuring Formationof Climatetoowarmtoform vs.Assuring wsleodwgley,spcearnhmaeplst swiitmhulftiasnseuoruisngly odrcyciunrgr,incgradcukrsing faitss1u2r,e0s0e0sNtiCmayted fpeorrimgleadciianlllya.stfMewa'ydehcaavdees over2000yrs. unlessfissures maydevelop BP,basedon afterditching. Sedimentswould formedpreviously veryquicklyand pollen replaceiceasit byperiglacial mayremain assemblage. melts activityor openformany desiccation. centuries. 5)Temperature -6to-8deg.C Anytemperature. Any Probablytoo Definitelytoowarmto conditions temperatureas warmtoform formperiglacially. longasdrying periglaciallyafter canoccur. ~14,000l4Cy BP 6)Compositionofin- Similarto Sand-injected Almostalways Nearlypuresand. Organicsedimentsvery filledsedimentsvs. sediments frombelow;host eoliansiltor, similartothoseadjacent adjacentsediments surroundingthe usuallyfine- morefrequently, wedges;delivered grained sand(few byslumpingfrom impurities) wallsoffissure. 7)Presenceof Canbedeposited Onlylocallyif Canbe Sandfillingthe Sedimentfillingfissures blanketinglayer oncefissuresare surfaceisreached; depositedonce fissuresalso doesnotblanket,but filled. mushrooming. fissuresare blanketsolder youngersedimentsdo. filled sediments. 8)Structureofin- Layeringparallel Usuallymassive, Probablyrather Nonenoted. Nonenoted. fillings tothewalland butmaybe massive,may horizontal vertical,stress- locallybe layeringboth producedbanding. laminated. possible,butmay alsobemassive. 9)Effectson Oftendeformed None. Crackingcan Nonenoted. Nonenoted. adjacentmaterial asicetakesup cause space. deformationin hostmaterial. — Figure 6. Observations and criteria ofthree hypotheses to explain the polygonal fissure fillings at the Shafer Mastodont Locality, Warren County, Indiana. cracking is estimated at about 13,000 l4C ybp upper in-fills are tapered and some are not based upon pollen correlation. Ice-wedge casts are usually polygonal: both When the characteristics of periglacially- upper and lower fissure fillings at the Shafer produced fissure in-fillings (i.e., ice-wedge Locality are polygonal. The dimensions ofthe casts) are compared with the observations and wedges and polygons are within the range ex- data at the Shafer Locality, there are both sim- pected for ice wedges. ilarities and differences (Fig. 6). Periglacial Other differences between ice-wedges and in-fills are wedge-shaped; at the Shafer Lo- the fissure in-fillings at the Shafer Locality in- cality, the lower fissure fills are, too, but some clude a verv different sediment for lower in- 90 PROCEEDINGS OF THE INDIANA ACADEMY OF SCIENCE fillings (pure sand) compared to the much fin- blows do match the conditions at the Shafer er surrounding materials, the lack of layering Locality. These include dimensions and lack in the in-fillings at the Shafer Locality, and of internal structure. Considering all the cri- the lack of deformation in the adjacent mate- teria for seismic sand-blows in Fig. 6, this rials. The periglacial origin for the upper set mechanism is not likely the source for either of fissure fillings at the Shafer Locality can set of in-fillings at the Shafer Locality. likely be dismissed because the late Pleisto- Desiccation and in-filling hypothesis: The cene and Holocene climate was much too third hypothesis relies on drying and cracking, warm to support permafrost. The possibility followed by in-filling with eolian sand at a of a periglacial origin for the lower sequence later date. Primary sources of information remains. Such a case is supported by the about desiccation phenomena used in the wedge-shaped in-fills, their polygonal pat- study were Conybeare & Crook (1968), Cal- terns, and range of dimensions for the in-fills, abresi & Burghignoli (1977), Haigh (1978), but these characteristics are not unique to peri- and Selley (1988). This hypothesis is not con- glacial mechanism. If the periglacial origin tradicted by any of the criteria in Fig. 6. Sim- hypothesis is considered for the lower fissure ilarities between desiccation-caused in-fillings fillings, then the permafrost would have to be and both sets of fissure fillings at the Shafer present at a time after all the silt, sand, wood Locality include the wedge shape, although fragments, bone fragments, cones and other this is more convincing in the case ofthe low- organic debris were washed into the lake; then er set. Similarly, both sets form polygons. We organic sediment accumulated under boreal have found very little data on the size of des- forest conditions for about three millennia iccation polygons, but ordinary mud cracks (i.e., by about 13,000 l4C ybp). By that time, are examples, and they seem to have dimen- it seems quite unlikely that permafrost was sions that sometimes match those at the Shafer around in Indiana. Mean annual temperatures Locality. were probably warmer than —6° to —8° C. The older set of fissures probably formed Seismic sand-blow hypothesis: Primary by 13,500 14C ybp (based on estimations de- sources of information about sand-blow phe- rived from the pollen profile) as a result of nomena used in the present study are Morris desiccation. In-filling with eolian sand (Unit 1983, Gohn et al. (1984), Obermeiere (1987), VI) must have followed quickly after Assuring Selley (1988), Munson et al. (1993), and Tut- because no other sediments are present in the tle & Barstow (1996). Sandy sediments can fissures. The upper set of fissures may have be mobilized at the time of an earthquake formed by desiccation following ditching and through a liquification process. The sand is lowering of the water tables in the early squeezed upward as a dike. The pressure will 1900's. Such fissures were filled with re- cause Assuring of older silty and clayey ma- worked organic sediment (Unit VI) before the terial as the sand is forcefully injected toward site was blanketed with sediment resulting the land surface. If it reaches the surface, the from cultivation ofthe surrounding landscape. injected materials will spread laterally for a Fissures of the upper set are still forming as short distance, forming a mushroom. witnessed by some fissures that were not filled Although the seismic sand-blow hypothesis with sediment. The fact that the upper fissures was considered as an origin for the fissure fill- do not extend deeper is probably controlled ings at the Shafer Locality, it was noted im- by the level of the lowered water table. mediately that seismic fissure in-fillings are All the observations and data at the Shafer not wedge-shaped and do not occur as poly- Locality seem to be consistent with the con- gons (Fig. 6). The seismic sand-blow origin clusion that both sets offissure fillings are the can be considered only as part of a complex product of desiccation. Neither the periglacial mechanism in which injection followed As- hypothesis nor the seismic sand-blow hypoth- suring by one of the other mechanisms. Other esis is a viable alternative to the desiccation aspects of seismic sand-blows that do not fit hypothesis. the Shafer Locality are the tendency of dikes Both desiccation events (represented by to widen with depth, the apparent lack of a Units V & VII) resulted in deep cracks (—80 sand source with depth, and lack of structure cm) in the sediment. This suggests a nearly in the fissure infillings. Some criteria for sand complete loss of the wetlands water source.