Plesiomenaeus poorei gen. nov., sp nov., (Crustacea: Decapoda: Pontoniinae) from Zanzibar

2009·5.7 MB·English

by  A J Bruce

MemoirsofMuseumVictoria66:25-34(2009)
ISSN 1447-2546(Print) 1447-2554(On-line)
http://museumvictoria.com.au/About/Books-and-Journals/Journals/Memoirs-of-Museum-Victoria
Pl^esiomenaeuspoorei gen. nov., sp nov., (Crustacea: Decapoda: Pontoniinae) from
X
ibar
A.J. Bruce
CrustaceaSection,QueenslandMuseum,P.O.Box3300,SouthBrisbane,Queensland,4101Australia,e-mail: abruce@
broad.net.au
Abstract Bruce,A.J.2009.Plesiomenaeuspooreigen.nov.,spnov.,(Crustacea:Decapoda:Pontoniinae)fromZanzibar.Memoirs
ofMuseumVictoria66:25-34.
Anewgenus,Plesiomenaeus,isdesignatedforanewspeciesofspongeassociatedpontoniineshrimp,P.poorei,
fromZanzibar,whichisdescribedandillustrated.ThenewgenusresemblesPericlimenaeusBorradaile,fromwhichitis
distinguished particularly by the lack of a molar process and fossa on the fingers of the major second pereiopod.
PlesiomenaeuspooreialsocloselyresemblesPericlimenaeusbouvieri(Nobili)andtherelationshipisdiscussed.
Keywords Plesiomenaeuspooreigen.nov.,sp.nov.,Crustacea:Decapoda:Pontoniinae,spongeassociate,Zanzibar.
Introduction Plesiomenaeusgen.nov.
Recent re-examination of some specimens from Zanzibar, DiagnosisRostrum greatlyreduced, compressed, uni-dentate,
provisionallyidentifiedasPericlimenaeus?sp. nov.,indicated carapace without supraorbital, epigastric, hepatic or antennal
that they were not members of the genus Periclimenaeus spines, inferior orbital angle acute, first abdominal tergite
Borradaile, 1915, sensu stricto, as the fingers of the major without anteromedian lobe, pleura rounded, telson with two
second pereiopod lacked the characteristic features of the pairs ofsmall dorsal spines, three pairs ofposteriormarginal
second pereiopod of that genus, in which the fingers are spines,scaphoceritereduced,labrumnormal,mandiblewithout
providedwithadactylarmolarprocess andopposingfossaon palp,maxillipedswithflagellaslender,withfourlong,plumose
the fixed finger. The specimens could not be referred to any terminal setae, maxilla with basal endite simple, third
otherpontoniine genusandanew genusisnowdesignatedfor maxilliped with ischiomerus and basis fused, coxa without
their reception. The new species resembles very closely the arthrobranch,fourththoracicsternitewithoutmedianprocess,
species Typton bouvieri described from Djibouti by Nobili first pereiopod chela with fingers subspatulate, cutting edges
(1904, 1906) and also reported from Suez by Balss (1927). entire, dactyl with tridentate tip, medial and lateral teeth
There have been no subsequent reports of Nobili’s species, denticulate, fixed finger distally bidentate, second pereiopods
whichwastransferredtothegenusPericlimenaeusBorradaile, well developed, unequal, similar, majordactyl fingers simple,
1915, by Holthuis (1952). Holthuis, when examining the five without molarprocess andfossa, minor fingers non-shearing,
syntypes,notedthatthemajorsecondpereiopoddactylhasthe ambulatorypereiopodsrobust,thirdpropodmostslender,fifth
cutting edge provided with “a strong hammer shaped tooth propodstoutest,dactylssimplybiunguiculate,uropodalpropod
fittingintoacavityonthefixedfinger”,clearlyindicatingthat unarmed,exopodofuropodwithdistolateraltoothandspine.
T. bouvieriiscorrectlyplacedinPericlimenaeusBorradaile.
Abbreviations used: CL, postorbital carapace length; Type species. Plesiomenaeus poorei sp. nov., by present
NMV, National Museum of Victoria, Melbourne: MNHN, selectionandmonotypy.
Museum National d’Histoire Naturelle, Paris. RMNH, Etymology. Fromplesios (Greek) near, and part ofthe name
Nationaal Natuurhistorisch Museum-Naturalis, Leiden;
OUMNH, Oxford University Museum of Natural History, Periclimenaeus, firstusedbyBorradaile (1915), asthe shrimp
Oxford; QM,QueenslandMuseum,Brisbane. was initially identified as a strange Periclimenaeus. Gender
masculine.
2
ystematicAccount Systematicposition. The genus Plesiomenaeus most closely
FAMILYPALAEMONIDAERafinesque, 1815 resembles the genus Periclimenaeus Borradailel915, type
species Periclimenaeus robustus Borradaile, 1915 (re-
SubfamilyPontoniinaeKingsley, 1878 describedbyBruce,2005).Inthisspeciesamarkedthickening
26 A.J. Bruce
ofthe posteriorthree fifths ofthe second pereiopod dactylar submedian spines more slender, finely setulose, about 0.8 of
cuttingedgeisdistinctlydemarcatedfromtheswollenbaseof intermediatespinelength.
thedactyl(Bruce,2005,fig.2C).Thisthickenedportionisless Antennule (fig. 2F) with proximal segment of peduncle
developedthan in several other species ofthe genus where it (fig. 2G)abouttwiceaslongasproximalwidth,lateralmargin
formsthetypicalposteriorlyandanteriorlydemarcatedmolar straight, non-setose, without ventromedial tooth, lateral
tooth, with a large well delineated opposing socket on the margin angular, distolateral angle rounded (fig. 6B) with 1-2
fixedfinger.InP.robustus,andotherspeciesofPericlimenaeus, short plumose setae, sometimes acute, stylocerite acute,
the occlusal surface of this thickened portion is distinctly projecting laterally, reaching to about half segment length,
flattened. In Periclimenaeus, and many other pontoniine statocystpoorlydeveloped,withoutstatolith,intermediateand
genera, the proximal occlusal end of the dactyl is normally distal segments shortandbroad,combinedlengthabout0.4of
swollen and quite distinct from the molar process which is proximal segment length, upper flagellum short, biramous,
developedontheintermediateportionofthecuttingedge(see withproximal 8 segmentsfused, shortfreeramus with single
fig, 4J, Periclimeaeus gorgonidarum (Balss), Wilson Island, long segment,with2 groups ofaesthetascs,longerfreeramus
Queensland, 6.0 m, coll. N.L. Bruce, 28 August 1980, AJB with 5 slender segments, lower flagellum short, filiform with
3106, QMW28914). Incontrast,inPlesiomenaeusthereisno 10 segments.
trace ofamolarprocess inthis position. Theproximal dactyl Antenna(fig.2H)withcarpoceritesubcylindrical,about3.0
is swollenandarticulatesinadeeplongitudinal depressionin timeslongerthanwide,basiceriterobust,withoutlateraltooth,
the proximal fixed finger but this lacks a defined anterior withlargeroundedprotuberantantennalglandprocessmedially;
margin and does not form a socket. In Plesiomenaeus the scaphocerite (fig. 21) small, subequal to carpocerite length,
second pereiopod chelae are similar but differing in size, in about 2.7 times longer than wide, distally rounded, lateral
contrastin Periclimenaeus they are ofdifferent morphology. margin straight with small acute distal tooth, at about 0.9 of
Plesiomenaeusalsolackstheanteromedianlobeonthedorsal scaphoceritelength,well shortofmarginoflamella,distal and
margin of the first abdominal tergite conspicuous in P. medialmarginswithnumerous shortplumosesetae.
robustus. Epistomeunarmed,withoutspecialfeatures.
Eye, (fig. 2J), withhemispherical cornea, well pigmented,
Plesiomenaeuspooreisp.nov. diameter about 0.13 of CL, without accessory ocellus, stalk
globular,about 1.1 timeslongerthanwide.
Figures 1-7 Mandible(fig.3A) small,withoutpalp,molarprocess (fig.
Materialexamined.1ovig.?,holotype,Chukwani,Unguja,Zanzibar, 6D) subcylindrical, tapering distally, distally obliquely
6° 13' 60"S30° 13'00"E,0.1matlowwaterspringtide,4December truncate with two small teeth and numerous rows of short
1960,coll.A.J.Bruce,#271,NMVJ59993. 1S,allotype,idem,NMV spiniformsetae,incisorprocesssmall(fig.6E)narrow,distally
J59994. Ovig. ?, dissected, idem, NMV J59995. 1 S, 1 ovig. ?, rounded with three small acute teeth laterally, two smaller
paratypes, idem, MNHN-Nal7209. 1 S, 1 ovig. 9, paratypes, idem, teethmedially.
ZQCM.200W92-81925-60.01.11<5S,,1 o1vig.ov9i,g.par9a,typpeasr,atiydpeems,,RMidNemH,DO5U31M13N.H. largMearxtihlalnulalo(wfeirg.,3lBo)wewritthabpielroinbgedwpiatlhpd(fiisgt.al6Ft)u,buerpcpleerwliotbhe
Diagnosis.Withthecharactersofthegenus.Veryshortrostrum short slender terminal seta, upper lacinia (fig. 6G) short and
generally with single dorsal tooth only, proximal segment of broad, upper margin emarginate, distal margin broadly
antennularpeduncledistolaterallyrounded,withoutacutetooth, truncatewithabout 16shortsimplespinesandscatteredsetae,
ventralborderofsecondpereiopodmerusnon-tuberculate. lower lacinia tapering distally with six terminal spines and
numerous spiniformsetae.
Description. A stoutly built shrimp of subcylindrical body Maxilla(fig.3C)withsimpletaperingpalp,withfewshort
form(fig. 1). plumose setae proximo-laterally, basal endite simple, short
Rostrum(fig.6A)veryshort,about0.06ofCL,compressed, and broad, with 10 slender sparsely setulose terminal setae,
triangular, acute, with singe acute dorsal tooth, carapace (fig. coxal endite obsolete, medial margin broadly rounded, non-
2A) smooth, without supraorbital, epigastric, hepatic or setose, scaphognathitewell developed, about2.6times longer
antennal spines, inferior orbital angle (fig. 2B) acute, than wide, anterior lobe as long as wide, medial margin
pterygostomialangle stronglyproduced,rounded. concave,posteriorlobeabout0.8ofanteriorlobelength.
Abdomen smooth, first segment without anterior median First maxilliped (fig. 3D) with palp (fig. 6H) about 2.5
dorsallobe,pleurarounded,sixthsegment(fig. 2C)depressed, timeslongerthanwide,distallyrounded,withtwopreterminal
about0.24ofCL,withsmallsubacuteposterolateraltoothand feebly setulose setae disto-medially, basal and coxal endites
muchlarger, acuteposteroventraltooth;telson(fig. 2D) about fused,distallyroundedmedialmarginstraight,withnumerous
0.35 of CL, 2.15 times longer than anterior width, lateral slender sparsely setulose marginal setae, exopod with well
marginsslightlyconvex,taperingposteriorly,posteriormargin developed caridean lobe, flagellum slender with four long
rounded (figs 2E, 6J) two pairs of small submarginal dorsal plumoseterminal setae,epipodwell developed,bilobed.
spines, about 0.08 of telson length, anterior pair at 0.65 of Second maxilliped (fig. 3E) with endopod normally
telsonlength,posteriorpairatabout0.96(seeRemarks),with developed,dactylarsegmentabout3.4timeslongerthanbroad,
threepairs ofmarginal spines,lateral spines similarto dorsal medial margin with numerous long slender coarsely setulose
spines, intermediate spines about 0.1 of telson length, or finely denticulate spines, propodal segment distomedially
Plesiomenaeuspoorei,gen. nov.,sp.nov., Pontoniinae,fromZanzibar 27
NMV
Figure1.Plesiomenaeuspooreigen.nov.,sp.,ovig.?.Holotype, J59993,Scalebarinmillimetres.
produced, with numerous long slender sparsely setulose Firstpereiopod(fig. 4A) short, robust, chela(fig. 4B) with
marginal spines, proximal endopod segments normal, basis palm 1.5timeslongerthandeep, compressed, withnumerous
with slender flagellum, coxa with small suboval epipod, short simple cleaning setae proximo-ventrally, fingers
withoutpodobranch. subspatulate, with numerous groups of short simple setae,
Third maxilliped (fig. 3F) with ischiomerus and basis fully cutting edges entire, dactyl withtridentatetip (fig. 61) medial
fused, combined segmentabout3.0 times longerthan maximal and lateral teeth posteriorly tuberculate, fixed finger distally
width, tapering distally, medial margin straight with numerous deeply bidentate (fig. 61) simple; carpus 0.59 ofchelalength,
longslendersimplesetae,carpussubcylindrical,halfischiomerus- 2.9timeslongerthandistalwidth,withtransverserowofdistal
basislength,4.0timeslongerthanwide,withnumerousspiniform marginal cleaning setaewith3 long distoventral setae; merus
setae medially, distal segment missing in dissected specimen, 1.3 times longer than chela, 5.7 times longer than wide,
exopodwith slenderflagellum, slightly exceeding distal merus, uniform, slightlybowed;ischium0.33 ofchelalength, 0.28of
withfourmajorplumoseterminal setae(brokenoffindissected meral length, basis subequal to ischial length; coxa robust,
specimen),coxalwithwelldevelopedlowroundedlateralplate, withoutventralprocess,withdorsalflange.
withoutarthrobranch.Paragnaths(fig. 6C)deeplybilobed. Second pereiopods well developed, unequal, similar,
Thoracic sternites unarmed, narrowest atfourth and fifth fingers up-curved. Major chela (fig. 4C) length subequal to
segmentlevelsandbroadeninganteriorlyandposteriorly. CL,palmsmooth,ovalinsection,about2.1 timeslongerthan
28 A.J. Bruce
Figure 2. Plesiomenaeuspoorei gen. nov., sp., ovig. 9. NMV J59994. A, carapace and rostrum. B, anterior carapace and rostrum. C, sixth
abdominal segment,lateral. D,telson. E, same,posteriorspines, dorsal spineinset. F, antennule. G, same,proximal segment. H,. antenna. I,
scaphocerite.J,eye.K,uropod.
deep, tapering distally, distal width about 0.6 of maximal edgesimilar,withoutproximaltooth,carpusrobust,about0.4
width, non-setose; fingers (fig. 4CDE) 0.25 of palm length, ofpalmlength,distallyexpanded, 1.5timeslongerthandistal
dactyl about 3.0 times longer than proximal depth, dorsal width, tapering strongly proximally, unarmed, merus about
margin convex, with strong acute tip, cutting edge without 0.5 of palm length, 3.0 times longer than wide, unarmed,
molar process, unarmed, feebly convex, entire, sharp, fixed ventrallynon-tuberculate; ischium0.8 ofmeruslength, 0.4of
finger about 1.8 times longer than proximal depth, ventral palm length, 2.4 times longer than distal width, tapering
margin convex, with acute tip distally, occlusal edge proximally, unarmed, basis and coxa robust, without special
longitudinallygroovedthroughoutlengthwithdeepdepression features. Minor second pereiopod chela (fig. 4G) similar to
proximally, dorsal margin with bluntly triangular tooth majorchela, subequaltopalmlengthofmajorchela,palm3.5
proximally, with sharp entire cutting edge, ventral cutting timeslongerthandepth,tapering slightlydistally,fingers (fig.
Plesiomenaeuspoorei,gen. nov.,sp.nov., Pontoniinae,fromZanzibar 29
Figure3. Plesiomenaeuspoorei gen. nov., sp., ovig. 9. NMVJ59994.A, mandible. B, maxillula. C, maxilla. D, firstmaxilliped. E, second
maxilliped.F,thirdmaxilliped(terminalsegmentofendopodmissing).
4H)0.25ofpalmlength,similartomajorchela,carpus0.33 of accessorytooth,about0.5ofunguislength,withseveralsimple
palm length, 2.0 times longer than distal width; proximal ventralsensorysetae;propodabout0.3 ofCL,5.5timeslonger
segmentsasformajorchelabutsmaller. than proximal width, tapering distally, distal width 0.66 of
Third ambulatory pereiopod (fig. 41) moderately slender, proximalwidth,withthree stoutspines distally,medial,lateral
reachingbeyondcarpoceritebypropodanddactyl;dactylshort, and ventral, medial spine longest, 3.7 times longerthan basal
stout, compressed, about 0.12 of propod length, unguis well width,projectingbeyonddorsalmarginofflexeddactyl,ventral
developed, curved, 2.2 times longerthan basal width, 0.33 of spine shortest, 0.8 of medial spine length, ventral margin of
corpuslength,unarmed,corpus 1.2timeslongerthanproximal propod otherwisewithout spines; carpus 0.8 ofpropodlength,
depth, with dorsal margin strongly convex, ventral margin 4.0timeslongerthandistalwidth,slightlytaperingproximally,
sinuous, distally concave, sharp, unarmed, with acute distal unarmed; merus subequal to propod length, 3.4 times longer
30 A.J. Bruce
Figure4.Plesiomenaeuspooreigen.nov.,sp.,ovig.?holotype,NMVJ59993.A,firstpereiopod.B,same,chela.C,majorsecondpereiopod,D,
same,fingers,lateral. E,same,medial.F,same,oblique. G,minorsecondpereiopod,chelaandcarpus.H,same,fingers.I,thirdpereiopod.J,
Periclimenaeusgorgonidarum(Balss),QMW28914,majorsecondpereiopoddactyl.
Plesiomenaeuspoorei,gen.nov.,sp.nov., Pontoniinae,fromZanzibar 31
NMV
Figure5.Plesiomenaeuspooreigen.nov.,sp.,ovig.9. J59994.A,thirdpereiopod,propodanddactyl.B,same,distalpropodanddactyl,.
C,fourthpereiopod,propodanddactyl. D,same, distalpropodanddactyl. E,fifthpereiopod,propodanddactyl. F, same, distalpropodand
dactyl.
32 A.J. Bruce
NMV
Figure6.Plesiomenaeuspooreigen.nov.,sp.,ovig.9. J59994.A,rostrum.B,antenna,distolateralangleofproximalsegment.C,paragnath,
D,mandible,molarprocess.E,same,incisorprocess.F,maxillula,palp.G,sameupperlacinia.H,firstmaxilliped,palp.I,firstpereiopodchela,
fingertips.J,telson,posteriormargin.K,uropod,distolateralexopod.
than wide, unarmed; ischium 0.55 ofpropod length, twice as distoventral setae, dactyl about 0.12 of propod length, with
long as distal width, tapering proximally, unarmed; basis and corpus 1.5 times longer than proximal depth, unguis 0.2 of
coxawithoutspecialfeatures.Fourthpereiopodsimilar,propod corpuslength,accessorytoothsmaller.
subequal in length but more swollen, 3.6 times longer than First pleopod (fig. 1C) male paratype (CL 3.4mm) with
proximal depth, maximal width 3.0 times distal width, with protopod about2.5times longerthanbroad, exopod subequal
shorter, more slendermedial, lateral andventral spines, dactyl to protopodlength, 5.0times longerthanwidewithnumerous
withcorpus 1.6timeslongerthanproximaldepth,unguis0.2of plumose marginal setae, endopod, 0.7 of exopod length, 5.0
corpuslength,accessorytoothsmaller. Fifthpereiopodsimilar, times longer than proximal width, tapering distally, without
propodsubequalinlengthbutpropodmoreswollenthanfourth, medial accessory lobe, with numerous, about 20, simple
3.2timeslongerthanproximaldepth,maximalwidth2.8times spiniformsetaescatteredalongmedialborder. Secondpleopod
distal width, with single small ventral spine and numerous (fig. 7D) with protopod 2.0 times longer than wide, greatest
Plesiomenaeuspoorei,gen. nov.,sp.nov., Pontoniinae,fromZanzibar 33
Figure7. Plesiomenaeuspooreigen. nov., sp. nov. A, abnormal rostrum, ovig. femaleparatype. B, rostrum, male paratype, CL3.4. C, first
pleopod.D,same,endopod.E,secondpleopod.F,same,endopod.G,same,appendices,masculinaandinterna.
width centrally, 1.2 times longerthan first pleopod protopod, convex,non-setosewithsmallacutedistaltooth(fig. 6K)with
exopod similar to first pleopod, endopod (fig. 7E) subequal small spine medially (seeRemarks)-, endopod 0.95 ofexopod
exopod length, 4.4 times longer than central width, with length,2.0timeslongerthanwide.
numerousplumosemarginalsetaedistally,appendices(fig.7F) Measurements (mms). Holotype, ovigerousfemale, postorbital
at 0.55 of medial margin length, appendix masculina
carapace length, 6.0; carapace and rostrum, 6.4; total body
subcylindrical, 2.5 times longer than wide, about 0.16 of
length(approx.) 17.0;secondpereiopod,majorchela,6.0;minor
endopod length, with 5 setulose setae of increasing length
chela,5.0;lengthofovum,0.55.
distallyalongmedialmargin,terminalsetalongest,abouttwice
corpus length’ appendixinternaslightly longerthan appendix Etymology. NamedforDrGary C.B. Poore,Principal Curator
masculina,withfewterminalcincinnuli. (Marine Biology), Museum of Victoria, in recognition of his
Uropod (fig. 2K) with protopod unarmed; exopod broad, majorcontributions to Australian and wider carcinology over
about 1.6 times longer than broad, lateral margin feebly manyyears.
34 A.J. Bruce
Host. Unidentifiedspongeencrustingroundthebaseofacoral rostraltoothinfemalesandanon-tuberculatesecondpereiopod
colony. merus as opposed to two dorsal rostral teeth and a ventrally
Colouration. Nodata. tuberculatemerusinadditiontotheabsenceofamolarprocess
and fossa on the major second pereiopod fingers. The type
Remarks.Thetelsonoftheholotypeisslightlyabnormalinthat material of Typton bouvieri is held in the collections of the
the posteriorpair ofdorsal spines are placed onthe posterior MuseoRegionaledi ScienzeNaturali,Torino.
margin of the telson which thus presents the appearance of
having four pairs of posterior marginal spines, with only a Acknowledgements
singlepairofdorsalspines. Onefemale(fig.7A)waswithouta
This study was supported by the Australian Biological
rostrum, the absence appearing congenital rather than
traumatic.Twoofthemalespecimens(fig.7B)hadtworostral Resources Study.
teeth. Some specimens had asmall acute distolateral tooth on
References
the proximal segment of the antennular peduncle, usually
asymmetrically. Balss,H. (1927)BerichtliberdieCrustaceaDecapoda(Natantiaund
The presence of slender propods on the third pereiopod Anomura).ZoologicalResultsoftheCambridgeExpeditiontothe
and stouter propods on the posterior limbs is unusual in SuezCanal, 1924.XIV.TransactionsoftheZoologicalSocietyof
pontoniine shrimps. In some species of the closely related London,26,221-230.
genusPericlimenaeusthepropodsarenotgreatlydifferentbut Bruce, A.J. 2005. A re-description of Periclimenaeus robustus
in others, such as P. crassipes Caiman and P. trispinosus Borradaile, the type species of the genus Periclimenaeus
Borradaile, 1915(Crustacea:Decapoda:Pontoniinae).Cahiersde
Bruce,thethirdpereiopodpropods areparticularly shortand
stout and the posterior propods longer and more slender, in HolthBuiiosl,ogLi.eB.Ma1r9i5n2e.,T4h6e,D3e8c9a-p39o8d,afoifgsth1e-5S.ibogaExpedition.PartXI.
contrasttotheirdevelopmentinP.pooreisp. nov. The Palaemonidae collected by the Siboga and Snellius
DespiteintensivecollectingoverthecoralreefsofZanzibar, Expeditions with remarks on other species. II. Subfamily
andsimilarreefsinTanganyikaandKenya,overseveralyears, Pontoniinae. SibogaExpeditionMonograph,39a 10, 1-252,figs,
resulting in numerous other sponge associated shrimps, no 1-110,tab. 1.
furtherspecimens ofthis specieswereevercollected. Kingsley,J.S. 1878.ListoftheNorthAmericamCrustaceabelonging
The resemblance ofP. poorei sp. nov. to Periclimenaeus tothesub-orderCaridea.BulletinoftheEssexInstitute, 10(4-6),
bouvieri (Nobili) is remarkable. Holthuis’s statement that “a 53-71.
strong hammer shapedtooth fitting into a cavity onthe fixed Nobili, G. 1904. Diagnoses preliminaires de vingt-huit especes
pfilnagceer”m,enlteavesoflittlNeobdioluib’ts conscpeercnieisng thinese fPeeartiucrleismeannadeutsh.e Rnoouuvgeel.leBsuldleetSitnomdautoMpuosdeesumetdD’ecHaipstoodierseMnaacturroeulrlee,s Pdaerilsa,Me10r,
228-238.
Re-examination ofthe type material ofNobili’s species may Nobili,G.1906.FauneCarcinologiquedelaMerRouge.Decapodeset
shedfurtherlightonthedetailedmorphologyofthefingersof Stomatopodes. Annales des Sciences naturelles, Zoologie, (9) 4
the major second pereiopod and hence its exact systematic (1-3), 1-347,figs. 1-12,pis. 1-11.
position.Plesiomenaeuspooreisp.nov.isreadilydistinguished Rafinesque,C.S. 1815.)AnalysedelanatureoutableaudeI’universet
fromPericlimenaeus bouvieriby having only a single dorsal descorpsorganises.Palermo, 1-224pp.