PLANTATION SILVICULTURE FORESTRY SCIENCES Baas P, ed: New Perspectives in Wood Anatomy. 1982. ISBN 90-247-2526-7 Prins CFL, ed: Production, Marketing and Use of Finger-Jointed Sawnwood. 1982. ISBN 90-247-2569-0 Oldeman RAA, et aI., eds: Tropical Hardwood Utilization: Practice and Prospects. 1982. ISBN 90-247-2581-X Den Ouden P and Boom BK: Manual of Cultivated Conifers: Hardy in Cold and Warm- Temperate Zone. 1982. ISBN 90-247-2148-2 Bonga JM and Durzan DJ, eds: Tissue Culture in Forestry. 1982. ISBN 90-247-2660-3 Satoo T and Magwick HAl: Forest Biomass. 1982. ISBN 90-247-2710-3 Van Nao T, ed: Forest Fire Prevention and Control. 1982. ISBN 90-247-3050-3 Douglas J: A Re-appraisal of Forestry Development in Developing Countries. 1983. ISBN 90-247-2830-4 Gordon JC and Wheeler CT, eds: Biological Nitrogen Fixation in Forest Ecosystems: Foundations and Applications. 1983. ISBN 90-247-2849-5 Nemeth MV: The Virus-Mycoplasma and Rikettsia Disease of Fruit Trees. ISBN 90-247-2868-1 Duryea ML and Landis TD, eds: Forest Nursery Manual: Production of Bareroot Seed lings. 1984. ISBN 90-247-2913-0 Hummel FC, ed: Forest Policy: A Contribution to Resource Development. 1984. ISBN 90-247-2883-5 . \ . --- Manion PD, ed: Scleroderris Canker of Conifers. 1984. ISBN 90-247-2912-2 Duryea ML and Brown GN, eds: Seedling Physiology and Reforestation Success. 1984. ISBN 90-247-2949-1 Staaf KAG and Wiksten NA: Tree Harvesting Techniques. 1984. ISBN 90-247-2994-7 Boyd JD: Biophysical Control of Microfibril Orientation in Plant Cell Walls. 1985. ISBN 90-247-3101-1 Findlay WPK, ed: Preservation of Timber in the Tropics. 1985. ISBN 90-247-3112-7 Samset I: Winch and Cable Systems. 1985. ISBN 90-247-3205-0 Leary RA: Interaction Theory in Forest Ecology and Management. 1985. ISBN 90-247-3220-4 Gessel SP: Forest Site and Productivity. 1986. ISBN 90-247-3284-0 Hennessey TC, Dougherty PM, Kossuth SV and Johnson JD, eds: Stress Physiology and Forest Productivity. 1986. ISBN 90-247-3359-6 Shepherd KR: Plantation Silviculture. 1986. ISBN 90-247-3379-0 Plantation silviculture by KENNETH R. SHEPHERD Reader in the Department of Forestry Australian National University Canberra, ACT, Australia 1986 MARTINUS NIJHOFF PUBLISHERS a member of the KLUWER ACADEMIC PUBLISHERS GROUP DORDRECHT / BOSTON I LANCASTER Distributors for the United States and Canada: Kluwer Academic Publishers, 101 Philip Drive, Assinippi Park, Norwell, MA 02061, USA for the UK and Ireland: Kluwer Academic Publishers, MTP Press Limited, Falcon House, Queen Square, Lancaster LAI 1R N, UK for all other countries: Kluwer Academic Publishers Group, Distribution Center, P.O. Box 322, 3300 AH Dordrecht, The Netherlands Library of Congress Cataloging in Publication Data Shepherd, K. R. (Kenneth Ronald) Plantation silviculture. (forestry sciences) Bibliography, p. Includes index. n. 1. Tree farms. 1. Title. Series. 58391.537 1986 634·9' 5 86-14175 IS8N-13: 978-94-01 0-8478-9 e-IS8N-13: 978-94-009-4446-6 001: 10.1007/978-94-009-4446·6 Copyright © 1986 by Martinus Nijhoff Publishers, Dordrecht. Softcover reprint of the hardcover 1 st edition 1986 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, mechanical, photocopying, recording, or otherwise, without the prior written permission of the publishers, Martinus Nijhoff Publishers, P.O. Box 163, 3300 AD Dordrecht, The Netherlands. v TABLE OF CONTENTS Preface xi Acknowledgements xiii CHAPTER 1 INTRODUCTION NATURE OF PLANTATIONS 1 Plantations Defined 3 Monoculture Tree Crops 4 plantations and Long Term Productivity 5 PLANTATIONS, ECONOMICS AND LAND USE 6 COSTS AND BENEFITS 6 Land as a Resource 8 Plantations and Development 9 SILVICULTURE AS BOTH ART AND SCIENCE 9 SILVICULTURAL SYSTEMS 10 CHAPTER 2 SEED PRODUCTION 12 THE NEED FOR GOOD QUALITY SEED 12 THE REPRODUCTIVE CYCLE IN FOREST TREES 14 Development of Gymnosperm Seeds 14 Development of Angiosperm Seeds 18 FACTORS INFLUENCING FLOWERING AND SEED PRODUCTION 21 SEED PRODUCTION FROM THE NATURAL FOREST OR FROM PLANTATION 26 SEED ORCHARDS 31 Siting of Seed Orchards 33 Orchard Establishment 34 Orchard Management 37 CHAPTER 3 SEED HANDLING 41 TRADE IN SEED 41 SEED COLLECTION 42 Pre-collection Planning 42 Field Collection Methods 43 Precautions at Harvest to Minimise Disease Infection 47 Labelling the Crop 47 SEED EXTRACTION 48 DRYING AND STORAGE OF SEED 49 Seed Quality 51 Moisture Content 52 Seed Storage 53 SEED TESTING 55 Sampling the Seedlot 56 Physical Characteristics of the Seed Sample 58 Weight 58 Moisture content 58 Genuiness and origin 58 Seed Germination Tests 58 vi SEED DORMANCY AND PRE-SOWING TREATMENT 60 Immaturity of the embryo 61 Complete inhibition of water absorption 61 A state of dormancy in the embryo itself 62 Secondary dormancy 63 CHAPTER 4 NURSERY PRACTICE 64 SEEDLINGS FOR PLANTING 64 THE NURSERY SITE 64 Selection of a Nursery site 65 Layout of the Nursery 67 BARE-ROOT SEEDLING PRODUCTION 68 The Nursery Soil 68 Texture 68 Structure 69 Nutrients 69 Organic matter 70 Soil reaction, or pH 71 Mycorrhiza 72 Soil Management 73 Tillage 73 Fertiliser use 74 Maintaining soil organic matter 75 Adjusting soil pH 76 Pest and disease control 77 Irrigation 80 Seedbed preparation, Sowing and Maintenance 83 Sowing 84 Weed control 85 Pre-emergence weed control 85 Post-emergence control 86 Interrow Cultivation 87 Root Pruning and Root Wrenching 88 Nutrient Deficiency in the Seedbed 91 Nitrogen 91 Phosphorus 91 Potassium 91 Magnesium 91 Lifting, Packing and Transport of Bare-root Stock 92 Lifting 92 Packing 93 Transport 93 PRODUCTION OF CONTAINER STOCK 93 Choice of Container 94 Cost of the container 94 Weight when filled with a growing medium 95 Durahili ty 95 Effects on the root system 95 Choice of Growing Medium 96 Fumigation and Sterilization of Potting Mixtures 96 Sowing Seed or Pricking out into Containers 97 Shading 98 Low~shade 99 High shade 99 Disease Control 100 vii Watering 100 Hardening-off 100 Handling and Transport 101 ASEXUAL OR VEGETATIVE PROPAGATION 102 Propagation by Cuttings 103 Grafting and Budding 104 Tissue Culture 105 PLANTING STOCK QUALITY 106 NURSERY RECORDS 107 CHAPTER 5 PLANTATIONS AND LAND-USE 109 PLANTATIONS AND CONSERVATION 110 SITE EVALUATION 111 CLIMATE 111 Soil and Terrain 113 Market and Economic Considerations 116 Existing and potential markets 116 Availability of infrastructure 117 Labour supply 117 Scale of operations 117 Environmental Impacts of Plantation Establishment 117 Hazards of fire, wind, erosion and landslide 118 Hazards of a biological nature 118 Conservation of native plants and animals 118 Problems of stream sedimentation and loss of water quality 118 Soil compaction and loss of fertility 118 Adverse impacts on landscape values 118 Adverse impacts on recreational values 119 SITE QUALITY 119 PLANTATION DESIGN 121 Plantations and Landscape 122 Planning the Access System 123 Stage construction 123 Classification of plantation roads 124 Roading density 125 Compartment shape 125 Non-wood Production Plantations 125 Windbreaks and shelterbelts 128 Urban forestry 130 Farm forestry and agroforestry 131 CHAPTER 6 GROWTH OF EVEN-AGED CROPS 133 TREE STANDS, AND MONOCULTURES 133 GROWTH OF SINGLE STEMS 134 Height Growth 134 Free or indeterminate growth 134 Fixed or determinate growth 134 Diameter Growth 136 Root Growth 138 Tree Form and the Influence of the Tree Crown 140 GROWTH OF THE FOREST STAND 141 Competition, Tolerance, Dominance and Crown Class 142 Competition 142 Tolerance 142 viii Dominance and crown class 143 Heigh t Growth 145 Diameter Growth 147 Root Growth 149 STAND GROWTH AND STOCKING DENSITY 151 BIOMASS PRODUCTION 153 CHAPTER 7 PLANTING, SITE PREPARATION AND MAINTENANCE 162 SILVICULTURAL DECISIONS 162 SITE PREPARATION METHODS 164 Clearing the Land 165 Broadcast felling and burning 165 Felling, windrow clearing and burning 166 Drainage and Mounding 168 Drainage 168 Mounding 168 Deep Ripping the Planting Site 170 Ploughing and Screefing 171 Weed Control on the Planting Line 172 Fertilizing at Planting 177 Benefits of adding fertilizers 177 Forms of fertilizer used at planting 181 Economics of fertilizer use at planting 184 Legumes as a source of nitrogen 184 MAINTAINING SITE PRODUCTIVITY 186 Loss of Nutrients 188 Direct loss of nutrients in harvested biomass 188 Indirect loss of nutrients from the site 189 Water Relations and Second Rotation Sites 194 Improving the water holding capacity of the soil 194 Increasing tree crop water usage 185 Soil Compaction 196 Managing the Nitrogen Supply 198 CHAPTER 8 PLANTING AND EARLY MAINTENANCE 201 CHOICE OF SPECIES 201 purpose of the Plantation 201 Plantings primarily for wood production 201 Plantations for protection 202 Plantations for fuelwood and fodder 202 Amenity and urban planting 202 Site and Climate 203 Use of Indigenous Species 204 Use of Exotic Species 204 CHOICE OF ESPACEMENT 206 Espacement, Tree and Stand Growth 206 Mortality 206 Height 207 Diameter growth 207 Stem form 208 Bral}ch size 208 Volume production 209 Wood quali ty 210 Effects of Initial Espacement on Costs 210 ix PLANTING 212 Timing of Planting Operations 212 Special Forms of Planting 213 Agro-forestry 213 Taungya, or trees with food crops 215 Line planting, or line enrichment planting 216 urban plantings 216 Planting Methods 217 PROTECTION AND EARLY MAINTENANCE 219 Protection from Vermin 221 Watering and Water Conservation 222 Minimising Competition from other Vegetation 222 Applicatiion of Fertilizer 223 CHAPTER 9 PRUNING 225 REASONS FOR PRUNING 225 NATURAL PRUNING 226 ARTIFICIAL PRUNING 227 Methods of pruning 229 Axe pruning 229 Saw pruning 230 Shear pruning 230 High pruning 231 Mechanical pruning 232 Timing and Intensity of Pruning 233 PRUNING AND WOOD QUALITY 234 CHAPTER 10 THINNING 236 THINNING AND GROWTH OF STANDS 237 OBJECTIVES OF THINNING 242 National or Organisational Objectives 243 Financial Considerations 243 Integration of Forest and Industry 243 Biological Considerations 244 THINNING METHODS 244 Commencement of Thinning 244 Type of Thinning 246 Low thinning 246 Crown thinning 247 Selection thinning or thinning from above 248 Mechanical thinning 248 Free thinning 250 Thinning Interval 251 Thinning Intensity 252 Severity of Thinning 253 THINNING, HARVESTING AND THE ENVIRONMENT 253 WIND STABILITY AND THINNING 255 THINNING AND WOOD QUALITY 258 CHAPTER 11 SILVICULTURAL MANAGEMENT 263 TACTICS AND STRATEGIES IN SILVICULTURE 263 A SILVICULTURAL REGIME 265 Shorthand Notation for a Silvicultural Regime 266 x Factors Influencing a Choice of Regime 266 Silvicultural Prescriptions 269 Use of Systems Analysis to Determine Regimes 270 Recognition 271 Definitiion of the extent of the problem and its boundaries 271 Identifying the hierarchy of goals and objectives 272 Generation of solutions 272 Modelling 272 Evaluation of potential courses of action 272 Implementation of the results 273 MODELS FOR EVALUATING SILVICULTURAL ALTERNATIVES 273 The Data Base 274 Data for building a model 274 Data for using a model 275 Yield Simulation Models 275 Economic Analysis of the Results of Simulation 277 Present net value 280 Internal rate of return 280 Benefit cost ratio 281 Seeking Optimum Management Strategies 282 SILVICULTURAL REGIMES TO SUIT DIFFERENT MANAGEMENT OBJECTIVES 283 Maximum Production Regimes 283 with a small wood market 284 with no small wood market 285 Maximum Value Regime 285 An Agro-forestry Regime 287 A Regime for Windy Climates 289 PLANTATIONS AND THE FUTURF. 290 INDEX 293 REFERENCES 297
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